Cloeodes barituensis, Nieto, Carolina & Richard, Barton, 2008

Nieto, Carolina & Richard, Barton, 2008, The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species, Zootaxa 1727, pp. 1-21 : 6-8

publication ID

https://doi.org/ 10.5281/zenodo.181238

DOI

https://doi.org/10.5281/zenodo.6233767

persistent identifier

https://treatment.plazi.org/id/036D87E7-D530-FFC0-FF6C-B3BCFB2CFBAB

treatment provided by

Plazi

scientific name

Cloeodes barituensis
status

n. sp.

Cloeodes barituensis n. sp.

( Figs. 1–14 View FIGURES 1 – 4 View FIGURES 5 – 14 )

Male imago ( Fig. 1 View FIGURES 1 – 4 ). Length: body, 5.9–6.1 mm; fore wings, 6.7–6.8 mm. Head yellowish brown. Compound eyes orange brown, height of stalk half length of eye diameter. Antennae yellowish brown. Thorax yellowish brown. Legs pale yellow. Leg I: tibia 1.5 times the length of femur; tarsi as long as tibia with 4 segments decreasing in length apically. Fore wings ( Fig. 2 View FIGURES 1 – 4 ) hyaline, except costal and subcostal spaces translucent, hind wings absent. Abdomen: segments I, VII–X yellowish brown; segments II–VI pale yellow, posterior margins of terga with a dark transverse line. Genitalia ( Fig. 3 View FIGURES 1 – 4 ) with bases of forceps close together, segment III rounded. Cerci yellow.

Female imago. Length: body, 7.2–7.3 mm; fore wings, 8 mm. Similar to male imago except head yellow. Fore wings ( Fig. 4 View FIGURES 1 – 4 ) with marginal intercalary veins single. Cerci yellowish brown.

Nymph ( Fig. 5 View FIGURES 5 – 14 ). Length: body, 5.0– 6.5 mm; cerci, 1.6–1.8 mm; terminal filament, 1.5–1.6 mm. Antennae, 1.4–1.7 mm. Head yellow, longer than wide; compound eyes of the male grey brown. Antennae ( Fig. 6 View FIGURES 5 – 14 ) pale yellow, 1.5 times the head capsule, pedicel 1.5 times the length of scape. Mouthparts: labrum ( Fig. 7 View FIGURES 5 – 14 ) with one subapical seta centrally and two setae near lateral margin, anterior margin with basally bifid setae near midline and apically bifid setae near lateral margin. Left mandible (similar to Fig. 57 View FIGURES 55 – 67 ), without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible ( Fig. 8 View FIGURES 5 – 14 ) without setae between prostheca and mola. Hypopharynx ( Fig. 9 View FIGURES 5 – 14 ): lingua with a rounded projection. Maxillae ( Fig. 10 View FIGURES 5 – 14 ), palpi as long as galea-lacinia, two segmented, segment II with a constriction. Labium (similar to Fig. 61 View FIGURES 55 – 67 ), with segment III of palpi rounded.

Thorax yellowish brown, median line yellow. Fore wing pads yellow. Sterna pale yellow. Legs yellow, tarsi and tarsal claws yellowish brown. Femora ( Fig. 11 View FIGURES 5 – 14 ) dorsally with a row of short blunt spines, ventrally with spines, apically with a subquadrangular projection (similar to Fig. 48 View FIGURES 43 – 51 ). Tibia with bipectinate spines. Tarsi with short spines. Tarsal claws 0.4 times the length of tarsi. Hind wing pads absent.

Abdominal color pattern with segments I, X and anterior margin of segment II yellowish brown, segments IV, VIII and IX yellow, other segments yellowish brown with yellow spots as in Fig. 5 View FIGURES 5 – 14 . Posterior margin of terga with spines as in Fig. 12 View FIGURES 5 – 14 , except tergum I smooth. Sterna pale yellow, sterna II–VI with a small tuft of long fine setae laterally on each side. Paraprocts with spines apically, similar to Fig. 67 View FIGURES 55 – 67 . Gills ( Fig. 13 View FIGURES 5 – 14 ) translucent white, 2 times the length of each tergum, trachea pigmented. Caudal filaments yellow, 1 / 3 apically yellowish brown, with flattened setae basally sclerotized (similar to Fig. 30 View FIGURES 20 – 30 ). Cerci with long spines toward external margin every two segments, terminal filament with long spines toward midline dorsally and ventrally every two segments ( Fig. 14 View FIGURES 5 – 14 ).

Etymology. Baritú is the name of one of the rivers where this species was collected.

Diagnosis. Cloeodes barituensis n. sp. can be distinguished from the other species of the genus by the following combination of characters. In the imago, 1) female with fore wings with single marginal intercalary veins, male with fore wings with paired marginal intercalary veins; 2) hind wings absent; 3) male with genitalia ( Fig. 3 View FIGURES 1 – 4 ) with bases of forceps close together, segment III rounded. In the nymph, 1) hind wing pads absent; 2) posterior margin of abdominal tergum I smooth ( Fig. 5 View FIGURES 5 – 14 ); 3) abdominal color pattern with segments I, X and anterior margin of segment II yellowish brown, segments IV, VIII and IX yellowish, other segments yellowish brown with yellow spots as in Fig. 5 View FIGURES 5 – 14 ; 4) femora apically with a subquadrangular projection (similar to Fig. 48 View FIGURES 43 – 51 ); 5) dorsal edge of femora with a row of blunt spines (similar to Fig. 48 View FIGURES 43 – 51 ); 6) caudal filaments with flattened setae basally sclerotized (similar to Fig. 30 View FIGURES 20 – 30 ), cerci with long spines toward to external margin every two segments and terminal filament with long spines toward midline dorsally and ventrally every two segments ( Fig. 14 View FIGURES 5 – 14 ).

Material. Holotype: male nymph: ARGENTINA, Salta, Río Colorado, S 23 ° 22 '09'', W 64 ° 28 ' 11 '', 406 m, 1 / VI/ 2000, Romero, Molineri, Manzo & Nieto colls. Paratypes: 30 nymphs same data as holotype; 1 male imago (reared), 3 male and 2 female imagos, 20 male and 4 female subimagos: Depto. Santa Victoria, Baritú, Río Baritú, 1500 m, 10–17 / IX/ 1981, Domínguez col.; 5 nymphs: Aguas Blancas, Estación Jakulica, Río Pescado, Arroyo Arrazayal, 7–11 / XII/ 1984, Domínguez col. Material housed in IFML.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Ephemeroptera

Family

Baetidae

Genus

Cloeodes