Polyplectropus Ulmer

Chamorro, Maria Lourdes & Holzenthal, Ralph W., 2010, 2582, Zootaxa 2582, pp. 1-252 : 44-45

Genus Polyplectropus Ulmer

Polyplectropus Ulmer, 1905a: 103 [Type species: Polyplectropus flavicornis Ulmer, 1905a , monotypic].— Flint, 1964: 476 [larva described as Genus C].— Flint, 1968: 23 [larva].— Flint, 1968: 20 [systematics of the genus].—Bueno- Soria, 1990: 357 [revision of Mexican and Central American species].

Ecnomodes Ulmer, 1911:17 [Type species: Ecnomodes buchwaldi Ulmer, 1911 , monotypic, preoccupied by Turner, 1903 in Lepidoptera ].— Flint 1968: 21 [synonym].

Cordillopsyche Banks, 1913: 238 [Type species: Cordillopsyche costalis Banks, 1913 , monotypic].— Flint, 1967: 6 [synonym].

Ecnomodellina Ulmer, 1962: 5 [Type species: Ecnomodes buchwaldi Ulmer, 1911 , replacement name for Ecnomodes ].— Flint, 1968: 21 [synonym].

Eodipseudopsis Marlier, 1959: 117 [Type species: Eodipseudopsis tomensis Marlier, 1959 , monotypic]. — Oláh & Johanson, 2010: 22 [synonym].

Adult. Length of forewing 2.5–7.8 mm. Antennae stout, approximately 2/3rds length of forewing; scape no more than 1/3rd length of head. Maxillary palps 5-segmented in both sexes; 1st and 2nd segments as long as wide; 2nd with apical, mesal setose cushion; 3rd inserted preapically into 2nd and length equal to 1st and 2nd together; 4th 1.5 x length of 1st; 5th annulated, twice length of 3rd ( Fig. 2B). Labial palps 3-segmented; 1st and 2nd equal in length; 3rd 1.5 x longer than 2nd. Head with frontal, antennal, and occipital setal warts well developed; preocellar, ocellar, and postocellar setal warts less defined, but clearly present ( Fig. 2A); occipital warts 2/3rds length of head; ocellar warts rectangular, not fused. Prothorax with elevated rectangular median pronotal setal warts; lateral pronotal setal warts less pronounced or elevated; cervical setal warts bulbous ( Fig. 2A). Paired tegula heart-shaped with lateral lobe slightly more pronounced then mesal lobe ( Fig. 2A). Mesoscutum with distinct oval scutal setal warts located parallel to mid-line and not at angle. Mesoscutellum with single median oblong scutellar setal wart, with mid-line devoid of setae ( Fig. 2A). Tibial spur formula 3,4,4; foretibiae flattened, particularly in females. Color of body ranging from yellow to dark brown; setae on body brown with grey clusters on apex of head to entirely brown with golden reflections; setae on tegula long, golden brown and grey to dark brown; forewings ranging from yellowish-brown to dark brown or almost black, lighter wings may have brown to golden brown mottling. Forewing venation ( Figs. 2C, D): forks 1–5 present, rarely fork 1 absent; fork 1 petiolate, fork 2 sessile, fork 3 petiolate [with respect to median (m) crossvein], fork 4 petiolate, fork 5 petiolate [with respect to median-cubital (m-cu) crossvein]; discoidal, medial, and thyridial cells closed; Cu 2 vein apically bent beyond arculus, recurving proximally and joining combined anal veins 2+3 (A 2+3) at arculus; anal vein 1 (A 1) lightly pigmented, positioned anterad towards Cu 2 vein and distally meeting cu-a 1 and a 1 - a 2 crossveins. Abdominal segment V with short, occasionally elongate, external, paired, laterally digitate processes with associated internal bulbous glands.

Male genitalia. Sternum IX in lateral view rounded to deltoid, posterior margin shape ranging from more or less entire to possessing acute median projection. Tergum X membranous, oblong, bearing dorsal microsetae; lower lip of tergum X either produced and apically bilobed or entire or not readily apparent. Intermediate appendage of varying lengths, shapes, and pubescence, yet always distinct and not fused mesally to tergum X. Preanal appendage bipartite or tripartite; if tripartite dorsolateral process originating from dorsum of mesolateral process and either bulbous with dorsal microsetae or produced into sclerotized, sometimes short or elongate recurved, lanceolate process; mesolateral process setose, usually oblong to quadrate; mesoventral process setose, apex usually produced ventrad, dorsally produced into short or long digitate lobe, apically processes separated or completely fused, with or without peg-like setae. Inferior appendage bipartite with anterior basal plate either extending anterad beyond sternum IX (when observed in lateral view) or contained entirely within sternum IX, anteriorly bilobed or entire; dorsal branch usually highly setose and variously shaped, peg-like setae present or absent; ventral branch setose and variously shaped, but never larger than dorsal branch. Phallus long or short; dorsal phallic sclerite distinct, apex variously shaped; endothecal membrane with or without embedded spines of varying numbers, lengths, shapes, and robustness.

Female genitalia. Ventral plates of segment VIII setose and variously shaped; external part of gonopod of segment VIII in ventral view with posterior margin ranging from truncate to medially produced; internal part of gonopod VIII sclerotized, variously shaped and textured; several species with highly sclerotized paired structure, here termed internal rods, associated with internal part of gonopod VIII and genital chamber. Tergum IX devoid of setae, lightly sclerotized. Segment X setose, apically bearing pair of pigmented cerci surrounded by 2 pairs of membranous, digitate processes. Genital chamber opening to processus spermathecae through sclerotized circular opening of ductus spermathecae ( Nielsen 1980).