Marma argentina (Mello-Leitão, 1941)

Salgado, Alexandre & Ruiz, Gustavo R. S., 2020, Taxonomic revision of Marma Simon, 1902 (Araneae: Salticidae: Euophryini), Zootaxa 4899 (1), pp. 287-353 : 335-338

publication ID

https://doi.org/10.11646/zootaxa.4899.1.16

publication LSID

lsid:zoobank.org:pub:27E67BBB-DFD0-4A96-8269-9E1CB6153B83

DOI

http://doi.org/10.5281/zenodo.4456895

persistent identifier

https://treatment.plazi.org/id/03403F11-FFAE-FFB3-538B-FF7308E0FAB9

treatment provided by

Plazi

scientific name

Marma argentina (Mello-Leitão, 1941)
status

 

Marma argentina (Mello-Leitão, 1941)   , revalidated

Figures 44–46 View FIGURE 44 View FIGURE 45 View FIGURE 46 , 57G View FIGURE 57 , 58I View FIGURE 58

Ocnotelus argentinus Mello-Leitão, 1941: 185   (Holotype ♀: ARGENTINA: La Rioja: Anillaco   GoogleMaps [28°48’21.7”S, 66°56’29.2”W], leg. Birabén, deposited in MLP 14980 View Materials —examined).

Thysema dorae Mello-Leitão, 1944: 390   (Holotype ♀: ARGENTINA: Tigre, Guayracá, leg. Prosen, deposited in MLP 16300 View Materials — examined). Syn. nov.

Note. Marma argentina   was previously considered one of the synonyms of M. nigritarsis   . The differences between these species were explained in the diagnosis of M. nigritarsis   above. The comparisons between the female holotypes of Ocnotelus argentinus   ( Figs 45 View FIGURE 45 A–C) and Thysema dorae   ( Figs 45 View FIGURE 45 D–F) revealed that both belong to the same species, and therefore we propose the synonymy. Both sexes were described based on well-preserved specimens deposited in the MACN. Also, the male is described for the first time.

Revised diagnosis. Among all Marma   species, M. argentina   shares more similarities with M. rosea   and M. spelunca   sp. nov. The males of these species have a thick embolic disc, with both edges curved ( Figs 44C View FIGURE 44 , 46A View FIGURE 46 , 48C View FIGURE 48 , 50A View FIGURE 50 , 53C View FIGURE 53 , 55A View FIGURE 55 , 57 View FIGURE 57 G–I), and females have very long copulatory ducts that encircle the primary spermathecae and almost touch each other ( Figs 45F View FIGURE 45 , 46D View FIGURE 46 , 49F View FIGURE 49 , 50G View FIGURE 50 , 54C View FIGURE 54 , 55G View FIGURE 55 , 58 View FIGURE 58 I–K). However, the males of M. argentina   differ by having some palpal structures elongated (cymbium, embolus and process of embolic disc; Figs 44 View FIGURE 44 C–D, 46A–B, 57G–I). Females differ by having the proximal region of the primary spermathecae laterally expanded to the external side ( Figs 58 View FIGURE 58 I–K).

Description. Male (MACN-Ar 39650). Total length: 3.86. Carapace 2.14 long, 1.42 wide, 1.02 high. Ocular quadrangle 0.90 long. Anterior eye row 1.31 wide, posterior 1.11 wide. Legs 4312. Length of legs: I 4.00 (1.30 + 1.51 + 1.19); II 3.50 (1.16 + 1.25 + 1.09); III 4.15 (1.34 + 1.43 + 1.38); IV 4.20 (1.30 + 1.39 + 1.51).

Color in alcohol ( Figs 44 View FIGURE 44 A–B): carapace with triangle of scales pointing backwards; high concentration of setae around triangle; abdomen ventrally with three dark brown stripes delimited by stripes of pale circles; legs: femur I with dark distal ring and proximal region with dark retrolateral spot, II with dark distal ring and proximal region with prolateral and retrolateral dark spot, III with dark distal ring and proximal region with incomplete dark ring (dorsally not totally colored), IV with dark distal ring and proximal region with dark proventral spot; tibia I–IV with dark proximal ring and distal region with prolateral and retrolateral dark spot; tarsus I almost black, II with black tip, III–IV with proximal dark ring.

Palp: RTA finger-shaped ( Figs 44D View FIGURE 44 , 46B View FIGURE 46 ); elongated cymbium with narrow apical region ( Figs 44C View FIGURE 44 , 46A View FIGURE 46 ); embolic disc with curved and symmetrical edges ( Figs 44C View FIGURE 44 , 46A View FIGURE 46 ); PED very long (about four times the length of the exposed portion of embolic disc; Figs 44C View FIGURE 44 , 46A View FIGURE 46 ); PED emerging from the middle-distal part of embolic disc ( Figs 44C View FIGURE 44 , 46A View FIGURE 46 ); embolus tip ends slightly beyond tip of PED ( Figs 44C View FIGURE 44 , 46A View FIGURE 46 ).

Female (MACN-Ar 39648). Total length: 4.63. Carapace 1.81 long, 1.35 wide, 0.94 high. Ocular quadrangle 0.91 long. Anterior eye row 1.24 wide, posterior 1.06 wide. Legs 4321. Length of legs: I 2.82 (0.95 + 1.03 + 0.84); II 2.86 (0.96 + 1.05 + 0.85); III 3.83 (1.33 + 1.29 + 1.21); IV 3.92 (1.30 + 1.25 + 1.37).

Color in alcohol ( Figs 45 View FIGURE 45 G–H): carapace as in male; abdomen ventrally pale with dark lateral borders; leg: femur I with dark distal ring and proximal region with dark retrolateral spot, II with dark distal ring and proximal region with dark retroventral spot, III with dark distal ring and proximal region with incomplete dark ring (retrolaterally not totally colored), IV with dark distal ring and proximal region with dark proventral spot; tibia I–II with dark proximal ring and distal region with prolateral and retrolateral dark spot; III–IV with proximal and distal dark ring; tarsus I–II without dark marks, III–IV with dark proximal ring.

Epigyne ( Figs 45C, F, I View FIGURE 45 , 46D View FIGURE 46 ): copulatory openings very close to each other and placed more anteriorly than primary spermathecae; copulatory duct long, encircling primary spermathecae; proximal copulatory duct shorter than distal section (half the length); primary spermathecae arched and with distal portion slighty dilated; secondary spermathecae well developed.

Note. Due to the lack of time to examine all specimens of Marma   available at MACN, the macrosetae pattern of Marma argentina   has not been documented.

Other material examined. ARGENTINA: Salta: Rosario de La Frontera [25°47’56.9”S, 64°58’08.0”W], leg. Donadio, IX.1986, 1♁ (MACN-Ar 39632); GoogleMaps   Cafayate [26°04’22.9”S, 65°58’34.7”W], leg. M.E. Galiano, XI.1960, 1♁ 1♀ (MACN-Ar 39571). GoogleMaps   Córdoba: Cerro Colorado [30°05’42.5”S, 63°55’55.8”W], leg. M.E. Galiano, 21.XI.1983, 2♁ 2♀ (MACN-Ar 39584); GoogleMaps   Bialet Massé [31°17’57.4”S, 64°28’03.6”W], leg. A. Gonzalez, XI.1985, 1♁ (MACN-Ar 39634); GoogleMaps   Argüello [31°20’55.8”S, 64°15’00.1”W], leg. J. A. De Carlo, XII.1943 – II.1946, 2♀ (MACN-Ar 39622), 1♀ (MACN-Ar 39629); GoogleMaps   San Roque, Parque Siquimán, Villa Playa [31°21’52.2”S, 64°28’43.7”W], leg. M. Miranda, I.1982, 1♀ (MACN-Ar 39544), leg. M.E. Galiano 20.XI.1983, 3♁ 2♀ (MACN-Ar 39631); GoogleMaps   Road Between R. Encantado and Panaholma [31°22’39.9”S, 64°09’39.5”W], leg. M.E. Galiano, 15.XI.1987, 2♁ (MACN-Ar 39494); GoogleMaps   Cuesta Blanca, Carlos Paz [31°28’56.4”S, 64°34’10.8”W], leg. Peretti, IX.1990, 2♀ (MACN-Ar 39621, 39624); GoogleMaps   Alta Gracia, Los Paredones [31°39’17.9”S, 64°26’32.6”W], leg. M.E. Galiano, 16.XI.1984, 3♁ 1♀ (MACN-Ar 39623); GoogleMaps   Cura Brochero [31°40’00.4”S, 65°01’05.5”W], leg. M.E. Galiano, 17.XI.1983 – XII.1987, 3♁ 4♀ (MACN-Ar 39526), 1♀ (MACN-Ar 39529), 3♁ 3♀ (MACN-Ar 39545), 1♁ (MACN-Ar 39619), 1♀ (MACN-Ar 39648), 6♁ 2♀ (MACN-Ar 39649), 1♁ (MACN-Ar 39650); GoogleMaps   Pampa de Achala [31°40’51.6”S, 64°50’14.1”W], leg. M.E. Galiano, 15.XI.1984, 2♁ 2♀ (MACN-Ar 39620); GoogleMaps   Alta Grácia, Vila Paisanita [31°43’08.9”S, 64°28’47.6”W], leg. A. Oliva, 7–13.XII.1984, 1♁ (MACN-Ar 39573). GoogleMaps   San Luis: Merlo, Pasos Malos [32°19’49.3”S, 64°59’55.0”W], leg. M.E. Galiano & Miranda, XI.1985, 2♁ (MACN-Ar 39628); GoogleMaps   Merlo [32°20’48.3”S, 65°00’56.8”W], leg. M.E. Galiano, 15.XI.1983 – 14.XI.1984, 2♁ 3♀ (MACN-Ar 39625), 2♁ 3♀ (MACN-Ar 39626), 2♁ (MACN-Ar 39627) GoogleMaps   .

Distribution. Known from the provinces of Salta, Córdoba and San Luis, Argentina ( Fig. 59A View FIGURE 59 ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Salticidae

Genus

Marma

Loc

Marma argentina (Mello-Leitão, 1941)

Salgado, Alexandre & Ruiz, Gustavo R. S. 2020
2020
Loc

Thysema dorae Mello-Leitão, 1944: 390

Mello-Leitao 1944: 390
1944
Loc

Thysema dorae

Mello-Leitao 1944
1944
Loc

Ocnotelus argentinus Mello-Leitão, 1941: 185

Mello-Leitao 1941: 185
1941
Loc

Ocnotelus argentinus

Mello-Leitao 1941
1941