Syleter papua Darlington, 1962

Balkenohl, Michael, 2021, Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini), Revue suisse de Zoologie 128 (2), pp. 357-374 : 363-365

Syleter papua Darlington, 1962

Figs 10 , 17 , 24

Syleter papua Darlington, 1962: 357 . – Moore et al., 1987: 78. – Lorenz, 1998: 138. – Lorenz, 2005: 148. – Balkenohl, 2001: 30. – Baehr, 2008: 13, figs 2, 16, 49, 62, 63, 177, 212.

Type material examined: Paratypes: BMNH; 1 ♂, TNHMUK 013460395 , white, black printed “Dobodura, Papua, N.G., Mar-July, 1944, Darlington ”, red, printed “Paratype Syleter papua Darl. ”, white, printed “Brit. Mus. 1963-26.”, circle, yellow framed, black printed “Paratype”. – 1 ♀, TNHMUK 013460393, with labels and data white, black printed: “Muffin Bay Dutch N.G. Aug 1944 Darlington ”, red: “ Paratype Syleter papua Darl. ”, white: “ Brit. Mus. 1963-26.”, circle, yellow framed, black printed “ Paratype ”. Damaged specimen with right mesotibia, left tarsomeres of the mesotibia, and right antennomeres three to eleven missing . – 1 ♀, BMNH; same data but barcode with No 13460398. Specimen with missing antennomeres three to eleven bilaterally.

Extended diagnosis: Head anterior eyes conspicuously elongated, with clypeal furrow slightly indicated by indistinct lateral rugae, smooth at middle, with an indistinct longitudinal pore on frons; clypeus and frons with distinct isodiametric reticulation, supraorbital plate almost smooth; neck constriction almost incomplete, developed as row of punctures with a more or less wide gap at middle; labrum seven-setose; mandible moderately elongated. Pronotum with reflexed lateral margin complete, diverging but slightly convex in anterior half; anterior margin excised, anterior angle projecting; lateral channel of pronotum of same width over entire length; basal constriction half as wide as flange, flange convex; disk with reticulation and small group of few punctures; with posterior angle marked as obtuse tooth. Elytron with all intervals longitudinally reticulated, with small tubercle at declivity of third interval; striae moderately wide, finely granulated, with punctures indistinct, umbilical setigerous punctures of equal distance, lateral margin at apex about 40% widened. Male genitalia ( Fig. 17 ) with aedeagus regularly curved, at apex conspicuously acute in ventral view, in lateral view developed apically as asymmetric distorted spatula curved ventrally, rounded at tip of apex, parameres asetose. Female genitalia ( Fig. 24 ) with coxostylus gently curved at apex, with large conelike seta, ventromedially with eighteen nematiform setae, five of them elongated, laterotergite with three long and two moderately long nematiform setae. Measurements: Body length 3.32-3.6 mm; for other measurements see Baehr (2008).

Remark: The species is well described by Darlington (1962) and additional general characters are provided by Baehr (2008) including measurements based on ten specimens, and description of the male and female genitalia. The ink sketches of the male genitalia and the shape of the female coxostyli in general reflect more or less the reality. However, the two females investigated show a different setae pattern as figured in Baehr (2008), and the subapical setose organ figured could not be observed (500 fold, Polyvar compound microscope with optimized condenser). The female specimens investigated by Baehr (2008) were from Dobodura whereas the females investigated in this contribution were found in the Indonesian part of the island (Vogelkop). Therefore it is possible the two populations belong to different forms.

Distribution: New Guinea: Papua-New Guinea: Dobodura; Indonesia: Muffin Bay ( Darlington, 1962). Northeast Australia (Queensland): Cape York ( Moore, 1987; Baehr, 2008).

Type locality: Dobodura in Papua-New Guinea.

Figures

Figs 7-13. Syleter spp., habitus. (7) S. paradoxus (Putzeys, 1868), specimen from Bangkok, female. (8) S. acutipalpis (Putzeys, 1877), lectotype. (9) S. fulvaster (Andrewes, 1927), paratype, male. (10) S. papua Darlington, 1962, paratype, female. (11) S. sinepunctatus sinepunctatus sp. nov., holotype. (12) S. sinepunctatus kluangensis ssp. nov., holotype. (13) S. gradus sp. nov. holotype.

Figs 14-20. Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (14) Syleter paradoxus (Putzeys, 1868), specimen from Tale Noi, Thailand. (15) S. acutipalpis (Putzeys, 1877), lectotype. (16) S. fulvaster (Andrewes, 1927), paratype. (17) S. papua Darlington, 1962, paratype.(18) S. sinepunctatus sinepunctatus sp. nov., holotype. (19) S. sinepunctatus kluangensis ssp. nov., holotype. (20) S. gradus sp. nov. holotype.

Figs 21-27. Female coxostyli with laterotergite. (21) Syleter paradoxus (Putzeys, 1868), specimen from Tale Noi, Thailand, with latero- and mediotergite. (22) S. acutipalpis (Putzeys, 1877), specimen from Lilong, China. (23) S. fulvaster (Andrewes, 1927), holotype. (24) S. papua Darlington, 1962, paratype. (25) S. sinepunctatus sinepunctatus sp. nov., paratype. (26) S. sinepunctatus kluangensis sp. nov., paratype. (27) S. gradus sp. nov. paratype.