Parasyleter, Balkenohl, 2021

Balkenohl, Michael, 2021, Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini), Revue suisse de Zoologie 128 (2), pp. 357-374 : 369

Genus Parasyleter gen. nov.

Figs 2 , 5 , 28, 29, 30

Type species: Dyschirius doriae Putzeys, 1873: 14 .

Genus diagnosis: Size 1.8 to 2.3 mm. Head ( Fig. 2 ) with clypeus and frons distinctly elongated and flattened anteriorly; clypeus and clypeal wing fused, delimitation between clypeal wing and supraantennal plate observable as indistinct obtuse notch in two of the species; clypeus straight or slightly convex anteriorly, as far as or slightly more projecting than clypeal wing; mandible elongated, narrow in apical third, laterally nearly straight or convex, hooked at apical tip; labrum three- or five-setose; maxillary palpomere distinctly narrowed and slender towards apex, nearly straight, two apical segments securiform; antenna with scapus and pedicellus eccentrically attached; eye without or with distinct reduction, gena small or extended posteriorly. Mentum not elongated, with lateral lobe convex, median tooth as wide as lateral lobe, acute, distinctly more projecting than lateral lobe, surface with slight transverse rugae. Pronotum square or indistinctly trapezoid, with reflexed lateral margin incomplete, running from anterior angle up to posterior setigerous puncture, posterior angle rounded off without tooth, lateral channel of equal width, median line narrow, finely or just not joining basal constriction; proepisternum visible basally in dorsal view; flange at base convex, as narrow as or up to two times as wide as basal constriction. Elytron with interval eight convex, lateral channel distinctly flared apically, with interval eight distinctly emarginated apically and row of umbilical setigerous punctures distinctly following the emargination, interval eight with distinct emarginated carina at apex; third interval without setigerous punctures. Hind wings fully developed or reduced. Legs with front and intermediate tarsomeres slightly widened, more distinct in males; mesotibia with four setae laterally, preapical seta either without any protuberance or arising from minute tubercle ( Fig. 5 ). Male genitalia ( Fig. 29 ) with aedeagus distinctly curved, apical part elongated, apex more or less spatulate or securiform. Female genitalia ( Fig. 30 ) with coxostylus one and two completely fused, with evident cone-like seta in apical third; laterotergite with isolated setae.

Differential diagnosis: See key to the genera.

Etymology: The name is a combination of the Greek adjective para- (besides) and the original genus name.

Species: Three described species belong into the genus: Dyschirius doriae Putzeys, 1873 known from Indonesia ( Figs 2 , 5 , 28, 29, 30 ), Psilus malayicus Andrewes, 1927 distributed in Malaysia, and Psilus porphyreus Andrewes, 1923 known from Sri Lanka. The genus includes also species not described yet and is currently in revision by the author.

Figures

Figs 1-3. Head, dorsal view. (1) Syleter paradoxus (Putzeys, 1868).(2) Parasyleter doriae (Putzeys, 1873) comb.nov.(3) Clypeuspinus validus (Andrewes, 1936) comb. nov.

Figs 4-6. Right mesotibia. (4) Syleter paradoxus (Putzeys, 1868) showing the small preapical protuberance and its seta inserted apically (arrow). (5) Parasyleter doriae (Putzeys, 1873) comb. nov. with no tubercles at all. (6) Clypeuspinus validus (Andrewes, 1936) comb. nov. showing the long protuberance and its seta inserted laterally (arrow).

Figs 28-30. Parasyleter doriae (Putzeys, 1873) comb. nov., specimen from Sarawak collected by G. Doria. (28) Habitus, male. (29) Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (30) Female coxostyli and laterotergite.