Carvalhoma parvum, Namyatova & Cassis, 2016
Namyatova, Anna A. & Cassis, Gerasimos, 2016, Revision of the staphylinoid and ground-dwelling genus Carvalhoma Slater & Gross (Insecta: Heteroptera: Miridae: Cylapinae) of Australia, European Journal of Taxonomy 253, pp. 1-27: 12-15
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Carvalhoma parvum sp. nov.
Recognised by the following combination of characters: head, pronotum, hemelytron and antennal segment I shiny and dark brown to black ( Fig. 1 View Fig ); body small, 1.5–1.6 in male and 1.9–2.2 in female; head elongate, ca 3.7–4.0× as long as pronotum width in male and 4.1–4.5× in female; pronotum laterally covered with dense wrinkles and setae at sides ( Fig. 2C View Fig ); antennal segment I short, ca 1.4–1.8× as long as vertex width in male and ca 1.3–1.4× in female.
The specific epithet refers to the size of the males, which are smaller than its congeners; from the Latin “parvus”, meaning “small, minute, short”.
Known from the southeastern part of Queensland and northeastern New South Wales ( Fig. 11 View Fig ).
AUSTRALIA: New South Wales: 2 ♀♀, NE slope of Little Mountain (just W of Karuah Rd), 32.6 ° S, 151.93333 ° E, 40 m, 4 Feb. 1993 – 9 Apr. 1993, M. Gray and G. Cassis (00043346, 00043324) ( AM). — Queensland: 1 Ƌ (00043506), 1 ♀ (00043507), Belmont Hills Bushlands, site 1, 27.51305 ° S, 153.11805 ° E, 80 m, 19 Feb. 2004, QM party ( QM); 1 Ƌ, same data, 1 Mar. 2004 – 31 Mar. 2004, QM party (00043508) ( QM); 1 ♀, Chelsea Road Bushlands Reserve, 27.47634 ° S, 153.1858 ° E, 15 m, 24 Feb. 2004, QM party (00043509) ( QM); 1 Ƌ, same data, 31 Mar. 2004 – 29 Apr. 2004, QM party (AMNH_PBI 00400914) ( QM); 1 Ƌ (00043503), 1 ♀ (00043504), same locality as holotype, 30 Jan. 2004 – 1 Mar. 2004, QM party ( QM); 1 ♀, same data, 1 Mar. 2004 – 29 Mar. 2004, QM party (00043505) ( QM); 1 ♀, same data, 31 Mar. 2004 – 30 Apr. 2004, QM party (AMNH_PBI 00404482) ( QM); 1 Ƌ, Illaweena Street, Drewvale, 27.63983 ° S, 153.0578 ° E, 40 m, 17 Apr. 2003, C.J. Burwell and S. Wright (AMNH_PBI 00400916) ( QM); 1 Ƌ, same data, 26 May 2003 – 1 Jul. 2003, QM party (AMNH_PBI 00400915) ( QM); 1 ♀, same data, 31 Mar. 2004 – 29 Apr. 2004, QM party (00043510) ( QM).
BODY LENGTH. 1.5–1.6.
COLOURATION ( Fig. 1 View Fig ). Mostly brown to dark brown, sometimes with reddish tinge, clypeus pale brown, labial segment I reddish brown to reddish, segments II–IV whitish yellow to yellow, segment IV brown apically; antennal segment II yellow, often reddish apically; fore- and middle coxae whitish yellow, often darkened basally; hind coxa the same colour as fore- and middle coxae or dark brown; femora reddish or dark brown, often whitish yellow basally, tibia whitish yellow, often darkened basally, tarsi whitish yellow to yellow.
SURFACE AND VESTITURE. Head and pronotum smooth and shiny, except for shagreened collar ( Fig. 1 View Fig ); pronotum laterally wrinkled ( Fig. 1 View Fig ); hemelytron shiny, with distinct punctures ( Figs 1 View Fig , 4E View Fig ), pleura entirely covered with short setae ( Fig. 4D View Fig ). Dorsum and abdomen clothed with long simple erect pale setae.
STRUCTURE AND MEASUREMENTS. Body ca 3.7–4.0× as long as pronotum width; frons protruding anteriad of eyes at distance subequal to at least half of eye diameter ( Fig. 1 View Fig , 4B View Fig ), head ca 1.8–2.4× as wide as long, vertex ca 1.1–1.4× as wide as eye diameter; head ca 1.0–1.1× as wide as high; antennal segment I ca 1.4– 1.8× as long as vertex width; segment II ca 4.1–4.9× longer than vertex width, and ca 2.5–2.9× longer than segment I; segment III slightly longer than segment I, segment IV slightly shorter than segment III; labial segments subequal in length ( Fig. 4H View Fig ); pronotum ca 1.2–1.3× as wide as long, ca 0.8–0.9× as wide as head, with single oval depression medially; metepisternum with distinct ridge ( Fig. 4D View Fig ); tarsal segment I almost twice as long as segment II, segment II slightly shorter than segment III ( Fig. 4L View Fig ).
GENITALIA ( Fig. 9 View Fig A–E). Right paramere without dorsal ridge, its outer margin concave, apex straight, not hooked; left paramere with ridge dorsally, its apex straight, not hooked, basal swelling rounded ( Fig. 9D View Fig ); theca mostly membranous, sclerotised only apically ( Fig. 9 View Fig A–B).
BODY LENGTH. 1.9–2.2.
COLOURATION ( Fig. 1 View Fig ). Similar to male, but usually somewhat darker, colour of labial segments III–IV, coxae and tarsi varying from yellow to brown, tibiae whitish yellow to pale brown, often brown at extreme apex.
SURFACE AND VESTITURE. As in male.
STRUCTURE AND MEASUREMENTS. Body ca 4.1–4.5× longer than pronotum width; head ca 1.9–2.2 × wider than long, vertex ca 1.0–1.3× wider than eye diameter; head ca 1.1–1.3× wider than high; antennal segment I ca 1.3–1.4× longer than vertex width, segment II ca 3.1–3.7× longer than vertex width, and ca 2.2–2.5× longer than segment I, segment IV slightly longer than segment III; pronotum ca 1.2–1.3× wider than long, ca 0.8–0.9× wider than head; hind femur twice longer than head height.
GENITALIA. Dorsal labiate plate with elongate sclerite along outer margin; sclerotised ring elongate posteriorly, its dorsal portion ca 2.5–3× as long as wide ( Fig. 9F View Fig ); posterior wall of bursa copulatrix without small tubercles ( Fig. 9G View Fig ).
Carvalhoma parvum sp. nov. is similar to C. malcolmae externally and in the dark brown colouration ( Figs 1–2 View Fig View Fig , 4–5 View Fig View Fig ), but the latter species differs in the following characters: larger body ( Figs 1–2 View Fig View Fig ), 2.25 in male and 2.3–2.8 in female, smooth lateral side of the pronotum, almost without wrinkles ( Fig. 2C View Fig ), lateral sides of the pronotum and pleura not densely setose ( Fig. 2C, K View Fig ), and relatively long antennal segment I, ca 2.0× as long as vertex width in male and ca 1.8–2.2× in female.
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