identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C687DDB9282254FF38F8F0FDB9C8DA.text	03C687DDB9282254FF38F8F0FDB9C8DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goncharovtilla Williams, Bartholomay & Lopez 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Goncharovtilla Williams, Bartholomay &amp; Lopez ,  gen. nov.</p>
            <p> Type species.  Goncharovtilla oblomovi Williams, Bartholomay &amp; Lopez ,  sp. nov. , designated here. </p>
            <p> Diagnosis. FEMALE. Females of  Goncharovtilla can be separated from other South American  Dasymutillini by having the posterior surface of the head concave and separated from the dorsal surface by a sharp transversely arcuate carina that is thickened laterally. Additionally, the body is blackish with a contrasting pattern of black and whitish setae, the genal carina is distinct anteriorly, the mesosoma has a thick scutellar scale and similarly large, thickened carina antero-lateral to the scale, the T2 disc has the cuticle uniformly blackish, the body is covered with numerous brachyplumose setae, and T6 has a defined pygidial plate. MALE. Males can be differentiated from other  Dasymutillini by the following characters: clypeus with longitudinal elevated crest medially and anterior/ ventral margin strongly projected medially, mandible lacking notch or tooth ventrally, mesopleuron not swollen or projected into conspicuous tubercle in dorsal half, S1 with strong tooth-like longitudinal carina, S2 lacking seta-filled pit, metasoma lacking any reddish or orange markings. The genitalia also have useful diagnostic features: the paramere is strongly upcurved apically and densely setose ventrally, somewhat dorsoventrally depressed; the cuspis has an elongate brush of posteroventrally directed setae; and the penis valve has two simple teeth posteroventrally. </p>
            <p>Description. FEMALE. Integument. Usually reddish to dark brown, sculpture mostly concealed by dense setation, apparently densely and finely sculptured where visible. Setae predominantly brachyplumose to simple. Head. Subrounded, strongly concave posteriorly, vertex margined by transverse arcuate carina separating dorsal and posterior surfaces; finely and densely sculptured where visible. Occipital carina apparently deeply recessed, not clearly visible. Eye hemispherical, strongly protruding, surface shining, ommatidia clearly discernible. Antennal scrobe poorly developed, short, quadrate, lacking dorsal and lateral carinae. Clypeus slightly protruded basomedially, anteroventral margin virtually straight. Malar space greater than basal height of mandible. Genal carina present. Hypostomal carina strong, extending to posterior mandibular condyle. Scape simple. Pedicel almost as long as wide, shorter than F1; F1 longer than wide and longer than F2. Mandible weakly curved basally, slender, tapering, with small inconspicuous tooth subapically, no ventral basal tooth, notch, or flange. Maxillary palp six-segmented; labial palp four-segmented; basal palpomeres strongly flattened and broadened, densely covered with short erect setae. Mesosoma. Longer than wide, subrectangular, evenly and slightly narrowed posterad, lacking clear constriction on mesonotum in dorsal view, dorsum convex to flat; propodeum weakly rounded posteriorly, disk and declivity indistinct; dorsal and posterior faces of propodeum densely and coarsely sculptured; lateral face of propodeum weakly and sparsely sculptured to unsculptured. Pronotal dorsal face (excluding anterior collar) with anterior margin straight in dorsal view, reclined in lateral view; epaulet at extreme lateral margin of anterior pronotal margin, scarcely distinguished from the latter; humeral angle rounded; posterior margin indiscernible. Scutellar scale present, with conspicuous pair of transverse anterolateral carinae. Pronotal-mesopleural suture distinct, partly concealed by dense setation. Vertical mesopleural ridge present, scarcely swollen, rounded, areolate throughout. Metapleuron predominantly sculptured, almost smooth dorsally. Metapleural-propodeal suture present, inconspicuous, partly concealed by dense setae; endophragmal pit small, inconspicuous, obscured by setae, surrounding area virtually flat. Metasternal process subtriangular, slightly less than a third of metacoxal length. Legs. Tibial spur formula 1–2–2. Foreleg with tarsal comb. Meso- and metatibiae each with two rows of prominent stout spines, few spines in each row, spines smaller basad; apical spurs narrow, finely serrate laterally, outer spur shortened. Metacoxa with strong slightly arcuate carina on inner margin, ending abruptly before coxal apex. Metasoma. Surface finely and densely sculptured, sparser and coarser on sterna and sparser anterolaterally on T2. T1 shorter than wide, its width about 1/3 the width of T2, strongly constricted on anterior third and evenly expanded from constriction apicad, weakly convex in lateral view, anterior and dorsal faces distinct; anterior auricle prominent, vertically lamellate but apparently dentate from above. Lateral felt line of T2 wide. T6 with conspicuous pygidial plate, slightly convex, large, subrounded, defined laterally and apically by carinae, longitudinally and irregularly striate-rugose throughout. S2 evenly convex, with conspicuous medial longitudinal ridge on anterior third, without posterior transverse groove, lacking lateral felt line. S6 posterior margin with strong bilobate projection.</p>
            <p>MALE. Integument. Mostly blackish to brownish-black, mostly concealed by dense setation, with strong dense sculpture where visible. Setae mostly brachyplumose, with many simple setae interspersed throughout. Head. Roundly transverse, as wide as mesosoma. Occipital carina distinct dorsally. Vertex slightly swollen posteriorly. Eye broadly hemispherical, strongly protruding, surface shining, ommatidia discernible. Ocelli small. Antennal scrobe shallow, not reaching eye margin, with transverse lamellate carina on dorsal margin. Clypeus slightly depressed immediately below insertion of antennae, conspicuously convex apicomediad, ventral half with conspicuous longitudinal crest medially, almost carinate, terminating in strongly protruding acuminate anterior/ventral margin. Malar space shorter than basal height of mandible. Genal carina absent. Hypostomal carina evident, evenly high throughout, extending in a smooth curve to posterior mandibular condyle. Scape bicarinate ventrally. Pedicel approximately as wide as long, shorter than F1, which slightly longer than F2. Mandible evenly weakly curved, bidentate apically, subapical tooth minute, lacking ventral tooth or notch. Maxillary palp 6-segmented; labial palp 4-segmented; basal palpomeres strongly flattened and broadened, densely covered with short setae. Mesosoma. Surface mostly covered with dense sculpture where visible. Pronotal epaulet weakly defined, reduced to a row or tuft of short setae; humeral angle rounded in dorsal view. Tegula broadly subcircular, convex, almost entirely smooth, except anterior and inner margins sparsely sculptured. Mesoscutum lacking notaulus; parapsis present, inconspicuous, short. Mesoscutellum flat, declivous in lateral view; axilla generally flat dorsally, broadly connected to mesoscutellum, terminating in truncate/rounded lobe, with oblique longitudinal lamella connecting its ventral surface to mesoscutellum; dorsal surface of axilla smooth. Metanotum usually completely concealed by dense setae. Propodeum evenly convex in lateral view, dorsal and lateral faces indistinct. Metasternal process convergent to unidentate apex, shorter than metacoxal length. Wings. Fore wing with elongate sclerotized pterostigma; marginal cell slightly elongate, truncate apically; three closed submarginal cells, apical veins of third often inconspicuous or obliterated. Legs. Tibial spur formula 1–2–2. Mid and hind tibiae with two rows of weak spines; apical spurs finely serrated apically. Metacoxa with weak inner carina. Metasoma. Sculpture generally dense and fine; T1 longer than wide, its width about 1/3 width of T2, anterior and dorsal faces distinct, not constricted apically. T2 with lateral felt line broad and short. S1 with marked longitudinal median carina terminating posteriorly in subacute projection. S 2 in lateral view flattened weakly convex, lacking felt line; slightly elevated and projected anteromedially; with no indication of mesal seta-filled pit. Hypopygium longer than wide, surface almost flat to shallowly convex, posterior margin projected medially into sharp acute projection. Genitalia. Paramere long, free length longer than length from base of genital capsule to apex of parapenial lobe, sub-cylindrical and strongly upcurved at apical third; dorsoventrally flattened at basal 2/3; acute apically. Parapenial lobe well-developed. Cuspis short, somewhat dorsoventrally flattened, approximately 1/3 longer than digitus, paracuspis a somewhat elongated lobe. Digitus narrowly digitiform, slightly wider basad. Penis valve inner surface concave; with two sharp apical teeth on ventral margin; ventral teeth widely separated; without distinct setae.</p>
            <p>Distribution. Northeastern Brazil (Pernambuco, Ceará, and Maranhão).</p>
            <p> Etymology. This genus is named after the Russian author Ivan Goncharov (1812–1879) and the type species is named after one of his novels (see below). This name references the initial discovery of the genus in the Brazilian city of Russas (Portuguese for Russians [fem.]), Ceará, and tribal affiliation with  Gogoltilla Williams, Brothers &amp; Pitts, 2011 from Argentina (named after the Ukrainian-Russian author Nikolai Gogol). The name is feminine. </p>
            <p> Remarks. This genus was included in an unpublished molecular phylogeny of American  Dasymutillini (Williams 2012) as an undetermined  Traumatomutilla (Genbank accession numbers: JQ945822, JX002751, JQ945901). In that analysis, it was recovered as sister to  Tobantilla drosos Williams, Brothers &amp; Pitts, 2011 , a male-based species from Argentina, and this clade was recovered as the sister to the remaining New World  Dasymutillini (excluding  Lomachaeta Mickel, 1936 ). The male of this genus is structurally similar to other  Tobantilla species , particularly in the upcurved paramere and shelf-like clypeus. Females differ from  Tobantilla in many features, including body size (all  Tobantilla females are less than 6 mm in length), coloration (  Tobantilla females have a pale brown mesosoma and yellowish cuticular patches on T2), and head structure (  Goncharovtilla gen. nov. , is unique among  Dasymutillini in having the posterior margin of the vertex with a sharp transverse arcuate ridge). Furthermore, the genetic distance between  Goncharovtilla and  Tobantilla is comparable to that observed between various distinct genera within  Dasymutillini and greater than that observed within any single genus of  Dasymutillini (Williams 2012). </p>
            <p> In the best available key to Neotropical mutillid genera (Brothers 2006), females of  Goncharovtilla key out in couplet 23 as “  Dasymutilla /  Traumatomutilla ”. The unique head structure separates  Goncharovtilla from females of those genera. Males of  Goncharovtilla also key to “  Dasymutilla /  Traumatomutilla ” in couplet 65. They can be separated from those genera by the upcurved paramere and shelf-like clypeus. </p>
            <p> The female head morphology is apparently unique, with its transverse arcuate carina separating the dorsal and posterior surfaces. Some North American species in the  Dasymutilla monticola species-group approximate this head morphology; unlike  Goncharovtilla , all these species lack a genal carina and have the body color predominantly reddish-brown (Williams 2012). </p>
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	https://treatment.plazi.org/id/03C687DDB9282254FF38F8F0FDB9C8DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto. A.;Bartholomay, Pedro R.;Lopez, Vinicius M.;Brothers, Denis J.;Martins, Herbeson O. J.;Ferreira, Vinina S.;Koroiva, Ricardo;Guillermo-Ferreira, Rhainer	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.text	03C687DDB92A2252FF38F8D3FB01C982.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goncharovtilla oblomovi Williams, Bartholomay & Lopez 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Goncharovtilla oblomovi Williams, Bartholomay &amp; Lopez ,  sp. nov.</p>
            <p>(Figs 1–16, 18–20)</p>
            <p>Diagnosis. As for genus.</p>
            <p>Description. FEMALE. Body length. 6.5mm in holotype (4.0– 9.5 mm). Head. Posterior margin shallowly concave in dorsal view. Vertex width 0.8 × pronotal width. Eye height in frontal view 1.2 × distance from its ventral margin to mandibular condyle. Head surface entirely concealed by dense setation, except gena basally with sparse fine punctures. Mandible with small subapical tooth. Dorsal and lateral scrobal carinae absent. Antennal tubercle mostly asetose, shallowly and irregularly rugose. F1 1.6 × pedicel length; F2 1.3 × pedicel length. Genal carina present, ending before hypostomal carina. Occipital area concave. Dorsal and posterior surfaces of vertex separated by sharp transversely arcuate carina. Mesosoma. As long as wide. Surface completely concealed by dense setation dorsally. Anterior face of pronotum densely and conspicuously covered with longitudinal striae. Humeral carina present, narrowly separated from low inconspicuous epaulet, antero-lateral corner of pronotum rounded in dorsal view. Pronotal spiracle virtually flat against lateral margin of pronotum. Ratios of widths at humeral angles, pronotal spiracles, widest point of mesonotum, narrowest point of mesonotum and propodeum posterior to propodeal spiracles 1.00:1.05:1.15:0.91:0.94. Lateral margin of mesosoma slightly diverging anterad towards pronotal spiracle and converging slightly towards humeral angle after pronotal spiracle. Scutellar scale present, well-developed, wide, more than twice width of poorly defined anterolateral carinae, separated from the latter. Posterior and dorsal faces of propodeum undefined, propodeum almost vertical in lateral view. Propodeal spiracle virtually flat against lateral margin of mesosoma; no post-spiracular area. Metasoma. Dorsal sculpture, except pygidium and laterally on T2, completely concealed by dense setation; T2 densely and coarsely foveolate-punctate laterally. T1 0.3 × as wide as T2. T2 length 0.9 × its width. T2 maximum width anterior to midlength. S1 with conspicuous short longitudinal carina, terminating in short tooth-like projection posteriorly; S2–6 densely and coarsely foveolate-punctate. Pygidium subrounded, almost as broad as long, broadest nearer base; defined by strong arcuate lateral carina throughout its extension; surface with widely spaced longitudinal striations; interstices finely granulose. Coloration and variations. Body and appendages black to reddish-brown, except mandible, antennal tubercle, and flagellum partially yellowish-brown. Head setae silvery-white, dense, predominantly appressed and decumbent, with few sparse brownish erect setae, and dense decumbent brownish setae below occipital carina. Mesosomal dorsum with silvery-white setae laterally and black to brownish-black setae medially; mostly dense decumbent and appressed setae interspersed with sparse erect setae throughout; lateral face of pronotum, mesopleuron, metapleuron, and lateral face of propodeum silvery-white. Metasoma with interspersed appressed, decumbent, and erect silvery-white setae, except anterior half of T2 disc, posteromedial margin of T2 and medially on T3–6 with interspersed appressed, decumbent and erect brownish-black setae. Three different patterns for white setae distribution on the metasoma have been observed in this species: the holotype has a complete band of silvery-white setae on the posterior margin reaching the fringe only laterally, thus the fringe is black medially; one variant has both the silvery-white band and silvery-white fringe narrowly interrupted medially with black; a third form has both the silvery-white band and silvery-white fringe more widely interrupted medially.</p>
            <p>MALE. Body length. 9.9 in allotype (6.0–10.0) mm. Head. Transverse, subrectangular in dorsal view; posterolateral angles rounded; vertex width 0.85 × pronotal width. Eye almost circular. Ocelli small; OOD 3.7 × DLO, IOD 5.0 × DLO. Occipital carina distinct. Sculpture predominantly concealed by dense setation, coarsely and densely foveolate-punctate where visible. Gena ecarinate. Antennal scrobe concave to eye margin, with narrow transverse dorsal scrobal carina. Clypeus concave sublaterally, elevating slightly laterad and, especially, mediad; with conspicuous elevated longitudinal crest medially; ventral margin strongly acuminate, terminating medially in subacute projection; coarsely and sparsely punctate throughout, punctures slightly dense lateromedially. Scape bicarinate. F1 2.1 × pedicel length; F2 1.7 × pedicel length. Mandible slightly tapering apicad, smoothly curved medially; with low inconspicuous carina anterodorsally terminating before apical third; inconspicuously bidentate apically, subapical tooth minute; lacking ventral notch and projections. Mesosoma. Epaulets poorly defined, broadly disconnected from humeral carina, virtually flat against anterior margin of pronotum. Anterior face of pronotum mostly asetose, short, poorly and coarsely distinguished from dorsal face, indistinctly sculptured, with medial longitudinal slightly concave area. Tegula convex, mostly glabrous and impunctate except for dense punctures anteriorly and along inner margin. Dorsal sculpture of pronotum, mesoscutellum, and metanotum concealed by dense setation. Mesoscutellum flat, sloping posterad in lateral view. Axilla projected posterolaterally as truncate/ rounded lobe broadly connected to mesoscutellum; with oblique longitudinal lamella connecting its ventral surface to mesoscutellum; dorsal surface of axilla smooth. Metanotum equally long throughout. Propodeum convex, dorsal sculpture concealed by dense setation at dorsal third, densely areolate where visible; posterolateral margin smoothly rounded; dorsal face indistinguishable from posterior face. Lateral face of pronotum simply punctate. Mesopleuron mostly concealed by dense setation, densely and coarsely micropunctate where visible; smoothly elevated medially, without any conspicuous projections. Metapleuron mostly concealed by dense setation, sparsely micropunctate to smooth where visible.Lateral face of propodeum smooth to sparsely and shallowly foveolate to areolate posterodorsad. Wings. Fore wing with moderate elongate sclerotized pterostigma; length of pterostigma approximately twice its width; marginal cell elongated, its length approximately twice its width, truncate apically; three submarginal cells, apical veins of third cell vestigial. Legs. Densely setose, setae shorter and denser on tibiae and tarsi; lateral margin of spurs finely serrate. Metasoma. T1 0.1 × as wide as T2. T2 length 1.0 × its width. T1 densely and coarsely foveolate-punctate.T2 densely and finely foveolate-punctate. Sculpture of T3–7, except pygidium, concealed by dense setation. S1 densely and finely foveolate-punctate. S2 densely and coarsely foveolate-punctate, sculpture sparser posterad. S3–6 densely foveolate-punctate to punctate. S8 punctate, longer than broad, apical margin strongly acuminate. Genitalia. Ratios of free length of paramere, cuspis and digitus, 1.0:0.3:0.2. Parameres strongly convergent and broad in dorsal view, except at apical fourth where parallel and abruptly narrowed; dorsoventrally flattened, except subcylindrical apical third; in lateral view slightly curved ventrally, abruptly and strongly upcurved at apical fourth; with dense tuft of long setae on basal half, setae abruptly shorter and sparser, almost absent, on apical half. Cuspis short, stout, “clavate”, obscured by base of paramere in dorsal view, only partially visible in ventral view due to strong setation of paramere; slightly wider apicad, subparallel elsewhere; with dense long setae apicoventrally. Paracuspis short, about ¼ length of cuspis, subrounded, with few sparse setae. Digitus slightly elongate, weakly curved inward in dorsal view and slightly up-curved in lateral view, laterally compressed; apparently asetose. Penis valve concave on internal surface, with pair of acute teeth apicoventrally; without lateral pocket, asetose; apical distance between teeth 0.2 × length of valve. Coloration and variations. Body setae silvery-white, except area between posterior margin of ocellar triangle and posterior head margin, mesonotum, posterior half of T2 disc, and T5–7 with brownish-black setae. No significant variations were observed in the specimens examined.</p>
            <p>Distribution. Brazil (Pernambuco, Ceará, and Maranhão).</p>
            <p> Etymology. Named after the fictional character Ilya Ilyich Oblomov, the particularly lazy main character of the novel Oblomov by Ivan Goncharov. The first recorded specimen was collected in Russas, Ceará in 2010 and was almost immediately recognized by KAW as a new species. In a phylogenetic analysis of  Dasymutillini (Williams 2012), it was recognized to be a new genus. It took 50 pages for Oblomov to even rise from his bed, and it has taken 14 years for KAW to get around to naming and describing this spectacular wasp. </p>
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                 Material examined. [5♀ 3♂] Type material.   Holotype, ♀, BRAZIL, Pernambuco,  Petrolina , 09°19’44.2”S 42°33’30”W, 17.V.2019, coleta manual, H.O.J. Martins leg, MZSPHYM0136464 (MZSP; Fig. 18)  .  Paratypes: 1♀, same data as holotype except MZSPHYM0145123 (MZSP) ;  1♂, allotype, same data as holotype except MZSPHYM0136463 (MZSP) ;  1♂ same data as holotype except MZSPHYM0145122 (MZSP) ;  1♀, same data as holotype except Hym–002/0051, V.2018 (LESTES) ;  1♂,same data as holotype except Hym–002/0048, 24–28.III.2018 (LESTES) ;   1♀, Ceará,  
                <a title="Search Plazi for locations around (long -38.0074/lat -4.9711165)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-38.0074&amp;materialsCitation.latitude=-4.9711165">BR116, 5 km SW Russas</a>
                 , 04°58.267’S 38°00.444’W, 30 m, 22.VI.2010, C. Waichert col. (UFES; voucher JP1779)  . Other material. BRAZIL: 1♀, Maranhão, Alto Parnaíba,  Estação 8, 13.VIII.2013, L. P. Amaral-Neto (DZUP #299378; Fig. 19); 1♀, Riachão,  Fazenda Altos , 18–22.VIII.2009, Varrediura, F. Limeira-de-Oliveira leg. (INPA; Fig. 20). 
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            <p> Remarks. In both sexes, this species closely resembles  Traumatomutilla bifurca (Klug, 1821) in coloration (Bartholomay et al., 2019c) and many other mutillid species in the Brazilian Caatinga. The head structure of the female (Figs 1–3) and genitalia of the male (Figs 13–16) differentiate  Goncharovtilla oblomovi sp. nov. from any other species in the Caatinga and any known  Traumatomutilla species.</p>
            <p> The sex association is based on collection of males and females at the same locality on the same date in Pernambuco, demonstrating similar morphology (e.g., size and coloration). This association was subsequently confirmed through DNA sequence comparisons.  The Maximum Likelihood (ML) phylogeny is illustrated in Fig. 17, with posterior probability values and bootstrap values provided. The identical sequences of two females and one male corroborate the identification of a single species,  G. oblomovi sp. nov. Additionally, morphological similarities between both sexes and related genera further validate the sex association (see remarks for the genus above). </p>
            <p>The female specimens from Maranhão are not included as paratypes.They differ from the holotype and paratypes in setal coloration, having the white setal band of T2 widely interrupted by black setae (Figs 18–20). They also differ slightly from one another in the shape and thickness of the white setal bands on T2–4. Future studies, particularly if males are associated with these forms, could reveal them to belong to a separate species but, with the limited material available, they appear to form an incomplete gradient with the type series. With a nod to Goncharov’s Oblomov, we here employ the “lazy” approach of treating them as variants of a single species.</p>
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	https://treatment.plazi.org/id/03C687DDB92A2252FF38F8D3FB01C982	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto. A.;Bartholomay, Pedro R.;Lopez, Vinicius M.;Brothers, Denis J.;Martins, Herbeson O. J.;Ferreira, Vinina S.;Koroiva, Ricardo;Guillermo-Ferreira, Rhainer	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.text	03C687DDB922225BFF38FBD8FC0DC9C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyphuta Cambra, Williams & Brothers 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Dasyphuta Cambra, Williams &amp; Brothers ,  gen. nov.</p>
            <p> Type species.  Dasyphuta gorgupea Cambra, Williams &amp; Brothers ,  sp. nov. , designated here. </p>
            <p> Diagnosis. FEMALE: Females of  Dasyphuta can be separated from the other American  Dasymutillini genera by the following combination of characters: the mesosoma is more or less uniformly sculptured with deep small areolations, both dorsally and laterally, dorsally with sparse brachyplumose setae, and without a scutellar scale; the T1 shape is narrowly petiolate; and T6 is smooth and convex without a pygidial plate. MALE: Males of  Dasyphuta can be separated from other  Dasymutillini by the clypeus forming a more or less flat and densely punctate anteriorly bidentate shelf, the axilla narrowly spinose posteriorly, the pygidium smooth and convex without lateral carinae, the genitalic paramere strongly upcurved posteriorly, and the presence of erect brachyplumose setae on numerous portions of the body. </p>
            <p>Description. FEMALE. Integument. Reddish brown, sculpture clearly visible, generally confluently foveolate. Setae predominantly brachyplumose to simple. Head. Subrounded, lacking posterolateral tubercles. Eye hemispherical, strongly protruding, surface shining, ommatidia clearly discernible. Antennal scrobe poorly developed, short, quadrate, lacking dorsal and lateral carinae. Clypeus slightly protruded basomesally, anteroventral margin virtually straight. Malar space greater than basal height of mandible. Hypostomal carina strong, extending to posterior mandibular condyle. Scape simple. Pedicel as long as wide, shorter than F1; F1 longer than wide and similar in length to F2. Mandible nearly straight, slender, tapering, with small inconspicuous tooth subapically, no ventral basal tooth, notch, or flange. Maxillary palp six-segmented; labial palp four-segmented; basal palpomeres strongly flattened and broadened, densely covered with short erect setae. Mesosoma. Slightly longer than wide, pear-shaped, mesonotum bulging laterally, weakly constricted near pronotal and propodeal spiracles, dorsum convex; propodeum rounded posteriorly, disk and declivity indistinct; mesosomal dorsum and pleura more or less uniformly confluently foveolate. Pronotal dorsal face (excluding anterior collar) with anterior margin convex in dorsal view, flat in lateral view; epaulet distantly separated from humeral angle; humeral angle forming relatively sharply defined obtuse angle; posterior margin indiscernible. Scutellar scale absent. Pronotal-mesopleural suture indistinct. Mesopleuron sculptured throughout; vertical mesopleural ridge scarcely defined. Metapleuron predominantly sculptured, except narrow surface surrounding endophragmal pit. Metapleural-propodeal suture inconspicuous. Metasternal process subtriangular, slightly less than a third of metacoxal length. Legs. Tibial spur formula 1–2–2. Foreleg without tarsal comb. Meso- and metatibiae each with one row of prominent stout spines, few spines in each row, spines smaller basad; apical spurs narrow, finely serrate, outer spur shortened. Metasoma. Surface mostly confluently foveolate, sparser on sterna. T1 nearly as long as wide, its width about 1/3 width of T2, strongly constricted on anterior third and nearly parallel-sided from constriction apicad, petiolate in lateral view; anterior auricle prominent, vertically lamellate but apparently dentate from above. Lateral felt line of T2 linear. T6 convex without pygidial plate, smooth, except anterolateral portions punctate and setose. S2 evenly convex, with indistinct longitudinal ridge anteromesally, without posterior transverse groove, lacking lateral felt line. S6 posterior margin with strong bilobate projection.</p>
            <p>MALE. Integument. Mostly blackish to brownish-black, mostly concealed by dense setation, with strong dense sculpture where visible. Setae mostly brachyplumose, with many simple setae interspersed throughout. Head. Rounded, as wide as mesosoma. Occipital carina distinct. Vertex slightly swollen posteriorly. Eye broadly hemispherical, strongly protruding, surface shining, ommatidia discernible. Ocelli small. Antennal scrobe shallow, not reaching eye margin, with transverse lamellate carina on dorsal margin. Clypeus mostly flat and punctate, with slightly raised area below insertion of antennae and bidentate anterior/ventral margin. Malar space shorter than basal height of mandible. Genal carina absent. Hypostomal carina evident, evenly high throughout, extending in a smooth curve to posterior mandibular condyle. Scape bicarinate. Pedicel approximately as wide as long, slightly shorter than F1, which much shorter than F2. Mandible evenly weakly curved, bidentate apically, subapical tooth minute, lacking ventral tooth or notch. Maxillary palp 6-segmented; labial palp 4-segmented. Mesosoma. Surface mostly covered with dense sculpture. Epaulet weakly defined, reduced to a row or tuft of short setae; humeral angle rounded in dorsal view. Tegula broadly subcircular, convex, almost entirely smooth, except anterior and inner margins sparsely sculptured. Mesoscutum lacking notaulus; parapsis present, inconspicuous, short. Mesoscutellum shallowly convex with mesal longitudinal carina; axilla generally flat dorsally, narrowly connected to mesoscutellum, terminating in elongate spine; dorsal surface of axilla areolate. Metanotum concealed by dense setae. Propodeum evenly convex in lateral view, dorsal and lateral faces distinct but not separated by carina or obvious differences in sculpture. Metasternal process convergent to unidentate apex, shorter than metacoxal length. Wings. Fore wing with elongate sclerotized pterostigma; marginal cell short, truncate apically; three closed submarginal cells, apical veins of third largely inconspicuous or obliterated. Legs. Tibial spur formula 1–2–2. Mid and hind tibiae with three rows of thickened bristle-like setae, apparently without spines; apical spurs finely serrated. Metacoxa with weak inner carina. Metasoma. Sculpture generally coarse; T1 petiolate, longer than wide, its width about 1/3 width of T2, anterior and dorsal faces distinct, not constricted apically. T2 with lateral felt line linear. S1 with marked longitudinal median carina. S 2 in lateral view weakly convex, lacking felt line; with no indication of mesal seta-filled pit. Hypopygium elongate sub-triangular, surface shallowly convex, posterior margin projected mesally into sharp acute projection. Genitalia. Paramere with free length subequal to length from base of genital capsule to apex of parapenial lobe, sub-cylindrical and strongly upcurved at apical third; acute apically. Parapenial lobe well-developed. Cuspis short, somewhat dorsoventrally flattened, similar in length to digitus, paracuspis a small tubercle. Digitus narrowly digitiform, with widened apical knob. Penis valve inner surface concave; with two sharp apical teeth on ventral margin; ventral teeth widely separated; without distinct setae.</p>
            <p> Species included. Only the type species:  D. gorgupea sp. nov.</p>
            <p>Distribution. Panama.</p>
            <p> Etymology. From the Greek δασύς (dasus) “hairy”, and the genus name Ephuta Say. In reference to the tribal placement in  Dasymutillini along with the superficial resemblance to typical Ephuta in body size, coloration, and slender first metasomal segment in both sexes. The name is feminine. </p>
            <p> Remarks. This genus is similar to  Tobantilla in body size, coloration, and many structural features, but that genus is known from South America only. The undefined pygidium in both sexes, absence of scutellar scale in females, and spinose axilla of males provide strong justification to treat  Dasyphuta gen. nov. , as a separate genus. </p>
            <p> In the best available key to Neotropical mutillid genera (Brothers 2006), females of  Dasyphuta terminate in couplet 35, where their combination of characters does not match either option. Unlike  Lomachaeta , females of  Dasyphuta lack a genal carina, have densely foveolate-punctate mesosomal pleura, and have the T1 shape narrowly petiolate. Unlike the genera in couplet 36, females of  Dasyphuta lack distinctly plumose setae on the tergal fringes and the body length is less than 6 mm. Males of  Dasyphuta terminate in couplet 69, where their combination of characters does not match either option. They can be separated from the genera in couplet 70 by the bidentate mandible and uniformly black head; they can be separated from the genera in couplets 71 and 72 by the upcurved paramere, flattened indistinct epaulet, and narrowly petiolate T1 shape. </p>
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	https://treatment.plazi.org/id/03C687DDB922225BFF38FBD8FC0DC9C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto. A.;Bartholomay, Pedro R.;Lopez, Vinicius M.;Brothers, Denis J.;Martins, Herbeson O. J.;Ferreira, Vinina S.;Koroiva, Ricardo;Guillermo-Ferreira, Rhainer	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.text	03C687DDB9242243FF38FF63FD0BCC06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyphuta gorgupea Cambra, Williams & Brothers 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Dasyphuta gorgupea Cambra, Williams &amp; Brothers ,  sp. nov.</p>
            <p>(Figs 21–42)</p>
            <p>Diagnosis. As for genus.</p>
            <p>Description. FEMALE. Body length 5.0 in holotype (4.0–5.0) mm. Head. Posterior margin rounded. Vertex width subequal to pronotal width. Eye height in frontal view 1.9 × distance from its ventral margin to mandibular condyle. Head predominantly concealed by dense setae, confluently foveolate where visible. Mandible with small subapical tooth. Dorsal scrobal carina absent; lateral scrobal carina absent. Antennal tubercle asetose, faintly sculptured. Flagellomere 1 1.2 × pedicel length; flagellomere 2 1.2 × pedicel length. Genal carina apparently absent, sub-defined by irregular puncture margins. Occipital carina defined, equally wide throughout. Vertex lacking tubercles. Mesosoma. Mesosoma 1.2 × as long as wide. Mesosomal sculpture confluently foveolate. Humeral carina present, connected to short tooth-like epaulet by irregular carina, antero-lateral corner of pronotum angular in dorsal view. Pronotal spiracle slightly raised against lateral margin of pronotum. Ratios of widths at humeral angles, pronotal spiracles, widest point of mesonotum, narrowest point of mesonotum and propodeum posterior to propodeal spiracles 80:85:88:80:74. Lateral margin of mesosoma slightly emarginate anterior to propodeal spiracle, smoothly diverging anterad and converging slightly posterior to pronotal spiracle. Propodeal spiracle slightly raised against lateral margin of mesosoma; post-spiracular area not defined. Scutellar scale absent. Posterior and dorsal faces of propodeum poorly defined, posterior face apparently longer than dorsal face. Metasoma. Dorsal sculpture mostly confluently foveolate. Ratios of width of T1, width of T2 and length of T2 32:97:108. T2 maximum width anterior to midlength; its base conspicuously swollen over apex of T1. S1 with conspicuous longitudinal carina, highest anteriorly; S2 sculpture slightly sparser than T2 sculpture; S3–6 densely coarsely and irregularly foveolate-punctate to simply punctate; S2 with subapical transverse slope and anteromedial longitudinal ridge vestigial. T6 convex, without pygidial plate, sculpture smooth except for punctate and setose anterolateral portions. S6 sharply bidentate postero-mesally. Coloration and variations. Body mostly reddish brown; legs and antenna lighter orange; T2 with paired posterolateral circular yellow integumental spots. Head covered with dense pale golden setae. Mesosomal setae sparse, mostly silvery or golden except dorsally with large mesal patch (with posteromesal golden notch) and dorsolateral propodeal patches of sparse black setae. Metasoma setae sparse, mostly pale silvery except T2 disc (except lateral margins, yellow cuticular spots, and sub-circular anterolateral patch with pale golden setae) and fringes of T2 and T3 mesally black, and fringes of T2–5 brighter golden. Females vary slightly in intensity of the golden setae and some have the frons, mesosoma, and T2 disc variably darker brownish.</p>
            <p>MALE. Body length 6.0 in allotype (5.0–7.0) mm. Head. Rounded, width subequal to pronotal width. Eye almost circular. Ocelli small; OOD 4.5 × DLO, IOD 1.2 × DLO. Occipital carina distinct. Sculpture predominantly concealed by dense setation, areolate-punctate where visible. Gena ecarinate. Antennal scrobe with prominent transverse dorsal scrobal carina. Clypeus mostly flat and punctate, with slightly raised area below insertion of antennae and bidentate anterior/ventral margin. Scape bicarinate. Flagellomere 1 subequal to pedicel length; flagellomere 2 1.9 × pedicel length. Mandible narrow, bidentate; lacking ventral notch and projections. Mesosoma. Epaulets poorly defined, virtually flat against anterior margin of pronotum. Anterior face of pronotum mostly asetose, indistinctly sculptured, without mesal notch. Tegula convex, mostly glabrous and impunctate except for dense punctures anteriorly and along inner margin. Dorsal sculpture of pronotum, mesonotum, scutellum, axilla and metanotum areolate. Scutellum slightly convex, with mesal longitudinal carina. Axilla produced posterolaterally with sharp spine. Metanotum equally wide throughout. Propodeum convex, sculpture partially concealed by dense setation, areolate without enclosed area; posterolateral margins smoothly rounded; dorsal face rounded into posterior face. Lateral face of pronotum densely and confusedly areolate-punctate to micropunctate; mesopleuron densely and coarsely areolate-punctate to foveolate-punctate; mesopleuron more or less evenly convex, without any conspicuous projections. Metapleuron largely areolate with dorsal area partly smooth. Wings. Fore wing with elongate sclerotized pterostigma; marginal cell short, truncate apically; three closed submarginal cells, apical veins of third largely inconspicuous or obliterated. Legs. Mid and hind tibiae with three rows of thickened bristle-like setae, apparently without spines. Metasoma. T1 0.4 × as wide as T2. T2 length 0.9 × its width. Dorsal metasomal sculpture, except pygidium, foveolate-punctate to punctate. S1 with low longitudinal mesal carina, equally high throughout. S2 sculpture slightly sparser than that of T2, without seta-filled pit mesally. S3–6 densely punctate. S8 punctate, elongate triangular, apical margin with acute tooth-like projection mesally. Genitalia. Parapenial lobe pronounced apically, shorter than penis valve. Ratios of free length of paramere, cuspis and digitus 69:11:11. Paramere virtually straight and cylindrical throughout in dorsal view, apical third tapering towards acute apex; in lateral view conspicuously curved dorsally at apex; with sparse tuft of setae basoventrally. Cuspis short, “paddle-like”, slightly wider apicad; basically straight; with brush of elongate plumose setae emerging from posterior margin. Paracuspis a short asetose rounded tubercle. Digitus elongate, digitiform with apical knob, slightly up-curved in lateral view, apparently asetose. Penis valve concave on internal surface, with pair of acute teeth apicoventrally; apical tooth well defined, asetose; basal tooth less defined, short, without lateral pocket, asetose; apical distance between teeth 0.2 × length of valve. Coloration and variations. Body black to dark brown, except mandible, venter of flagellum and tibial bases pale yellow-brown. Setae predominantly silvery-white, except transverse black band on posterior half of T2 disc and apical tergites variably blackened. There is subtle variation in the extent of black setae on the apical tergites and various body regions have scattered erect darker setae.</p>
            <p>Distribution. Panama (Panama and Coclé).</p>
            <p> Material examined. [7 ♀, 24 ♂] Type Material:   Holotype, 1 ♀, PANAMA, PANAMA, Chorrera,  Llano Largo , 2.IV.1990, R. Cambra (CSCA)  .   Paratypes: PANAMA: Panama: Chorrera,  Playa Leona, cerca a río Perequete , 27.II.1991, yellow pan trap, R. Cambra (allotype, 1♂ CSCA)  ;   Chorrera,  Playa Leona, cerca a río Perequete : 6.II.1991, yellow pan trap, R. Cambra (1♂ CSCA)  ;  13.II.1991, yellow pan trap, R. Cambra, (2♂ MIUP) ;  18– 20.III.1991, yellow pan trap, R. Cambra (2♂ MIUP, 2♂ DJBC) ;  26.II.1991, yellow pan trap, R. Cambra (1♂ MIUP) ;  17.I.1992, R. Cambra (1♀ CSCA, 1♀ 2♂ MIUP) ,  25.I.1992, yellow pan trap, I. Henry (1♂ MIUP) ;  18.II.1992, R. Cambra (1♀ DJBC) ;  18.II.1992, A. Mena (2♀ MIUP) ;   Chorrera,  El Arado , 11.I.1993, yellow pan trap, col. R. Urriola (1♂ MIUP)  ;   oficinas Parque Nacional Soberanía , 26.II.2000, yellow pan trap, A. Santos (5♂ MIUP)  ;   Parque Metropolitano , 8.I.1993, yellow pan trap, A. Velásquez (1♂ MIUP)  ;   near barriada Don Bosco , 13.IV.2022, mangrove area, A. Santos (1♀ MIUP)  ;  Coclé: Tranquilla Norte, 12.III.1988, yellow pan trap, R. Rodríguez (3♂ MIUP) ;   Chiguirí Arriba,  Cerro La Vieja , 23.IV.1994, yellow pan trap, R. Cambra (1♂ MIUP)  . </p>
            <p> Etymology. A new word based on the combined specific epithets of the type species of  Dasymutilla Ashmead and Ephuta Say, 1836:  Mutilla (Sphaeropthalma) gorgon Blake, 1871 and  Mutilla (Ephuta) scrupea Say, 1836 . Treat as a noun in apposition. </p>
            <p> Remarks. The sexual association is based on morphological similarities of the mandible (bidentate apically, subapical tooth minute, lacking ventral tooth or notch), T1 (petiolate), similar body size, sculpture, and setation. Additionally, males and females have a sympatric distribution, captured at the same time in some places. Two other species of  Dasymutillini that are present in the same area (  Dasymutilla araneoides (Smith, 1862) and  Dasymutilla spilota Manley &amp; Pitts, 2007 ) already have associated males and females (Cambra et al. 2018). All the above is sufficient evidence to consider males and females of  Dasyphuta gorgupea as members of a single species. </p>
            <p> In both sexes, the coloration and body size are similar to many Panamanian species of Ephuta and  Pseudomethoca Ashmead, 1896 (KAW &amp; RAC, pers. obs.); females are predominantly reddish brown with a golden head and males are mostly black and covered with silvery setae. </p>
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	https://treatment.plazi.org/id/03C687DDB9242243FF38FF63FD0BCC06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto. A.;Bartholomay, Pedro R.;Lopez, Vinicius M.;Brothers, Denis J.;Martins, Herbeson O. J.;Ferreira, Vinina S.;Koroiva, Ricardo;Guillermo-Ferreira, Rhainer	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB93D2241FF38FD70FA25CBC9.text	03C687DDB93D2241FF38FD70FA25CBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasymutillini Brothers & Lelej 2017	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to American genera of  Dasymutillini</p>
            <p> The genera  Lomachaeta ,  Neomutilla , and  Protophotopsis were excluded from  Dasymutillini (Waldren et al. 2023) and are not keyed here. The key is written in a way that both sexes of velvet ants from the Americas can be checked for exclusion from  Dasymutillini or recognized as one of the 13 genera of  Dasymutillini . </p>
            <p>Females</p>
            <p>1. Eye subcircular; mandible unarmed ventrally; head unarmed ventrally; mesosoma widest at pronotum or mesonotum, in dorsal view; tergal fringes composed of simple or brachyplumose setae............................................... 2</p>
            <p>- With one or more of the following characters: eye longitudinally ovate; mandible with ventral tooth basally; gena and/or hypostoma armed with distinct ventral tooth; propodeum and pronotum both wider than mesonotum, in dorsal view; tergal fringes composed of distinctly plumose setae........................................ other tribes (not treated here)</p>
            <p>2 (1). T6 convex without defined pygidial plate.................................................................. 3</p>
            <p>- T6 with flat pygidial plate, usually with differentiated sculpture and defined by lateral carinae........................ 5</p>
            <p>3 (2). T1 shape narrowly petiolate, constricted with base of T2, T1 generally widest mesally............................... 4</p>
            <p>- T1 shape generally sessile, subsessile, or disciform, width variable, T1 generally widest at posterior margin......................................................................................... other tribes (not treated here)</p>
            <p> 4 (3). Genal carina and scutellar scale absent; mesosoma convex dorsally; T6 without longitudinal carinae; epaulet scarcely raised (Panama)..............................................  Dasyphuta gen. nov. —one species:  D. gorgupea sp. nov.</p>
            <p> - Genal carina and scutellar scale present; mesosoma flattened dorsally; T6 with two wavy longitudinal carinae mesally; epaulet usually raised onto smooth tubercle (Panama)..............................................................................................  Frigitilla Williams in Bartholomay et al. 2015 —two species, see Cambra et al. (2016) </p>
            <p>5 (2). Pygidium with lateral carinae, with acute erect tooth at posterolateral angle (mesosomal pleura uniformly covered with dense setae; tibial spurs black)................................................................................ 6</p>
            <p>- Pygidium with lateral carinae but without erect tooth at posterolateral angle...................................... 7</p>
            <p> 6 (5). Second labial palpomere mesally expanded, at least twice as wide as long; metasternal process short and truncated, with medial notch of variable depth; mesosomal dorsum with one or two narrow longitudinal stripes of dense whitish setae extending from anterior margin of pronotum to posterior margin of propodeum..............................  Suareztilla Casal, 1968</p>
            <p> - Second labial palpomere not expanded, at least as long as wide; metasternal process extended mesally into triangular plate between metathoracic coxae; mesosomal dorsum without narrow longitudinal stripes of whitish setae................................................................................................  Reedomutilla Mickel, 1964</p>
            <p>7 (5). Mesosoma broadly violin-shaped, propodeal sides divergent behind propodeal spiracle in dorsal view.................. 8</p>
            <p>- Mesosoma pear-shaped, propodeal sides convergent behind propodeal spiracle in dorsal view........................ 9</p>
            <p> 8 (7). Body uniformly pale yellow-brown; mesosoma with small scutellar scale; T2 disc with basomesal triangular patch of flattened striate setae (Argentina)................  Gogoltilla Williams et al. 2011 —One species,  G. chichikovi Williams et al. 2011</p>
            <p>- With one or more of the following characters: body color partly or entirely blackish; mesosoma without scutellar scale; T2 disc without patch of striated setae.................................................... other tribes (not treated here)</p>
            <p> 9 (7). Dorsal and posterior surfaces of vertex separated by sharp arcuate transverse carina (Brazil)............................................................................  Goncharovtilla gen. nov. —one species,  G. oblomovi sp. nov.</p>
            <p>- Posterior margin of vertex rounded or with separated posterolateral tubercles..................................... 10</p>
            <p> 10 (9). Scutellar scale bilobate; mesonotum armed with four or five dorsolateral tubercles; hypopygium apex with four spines (Bolivia, Brazil, and Paraguay)....................  Leucospilomutilla Ashmead, 1903 —three species, see Williams et al. (2020) </p>
            <p>- Scutellar scale absent or unilobate; mesonotum usually unarmed laterally, rarely with one spine; hypopygium apex lacking spines............................................................................................. 11</p>
            <p> 11 (10). Head broad and quadrate, vertex width greater than mesosomal width (T2 with large yellow or red integumental spots, paired, with marked edges; widespread in South America)....  Cephalomutilla André, 1908 —four species, see Williams et al. (2022) </p>
            <p>- Head not so broad, usually rounded posteriorly, vertex width usually less than mesosomal width..................... 12</p>
            <p>12 (11). Specimens from North and Central America or Caribbean Islands (excluding Trinidad)............................ 13</p>
            <p>- Specimens from South America (including Trinidad)........................................................ 14</p>
            <p> 13 (12). Body entirely black, except T2 with two or four yellow cuticular spots, these spots mostly bare, with sparse punctures; tibial spurs whitish (rarely encountered, only in Central America and Hispaniola)...........................................................................  Traumatomutilla André, 1901 , in part— three species, see Williams et al. (2017) </p>
            <p> - With one or more of the following characters: mesosoma reddish-brown; T2 disc without yellow cuticular spots or with spots having similar sculpture to surrounding T2 disc sculpture; tibial spurs black (widespread and common, in North and Central America and Caribbean Islands)................................  Dasymutilla Ashmead, 1899 , in part— over 100 species, see Manley &amp; Pitts (2007), Cambra et al. (2018), and Manley et al. (2020) </p>
            <p>14 (12). T2 disc cuticle uniformly blackish, lighter markings formed by setae only (genal carina absent)..................... 15</p>
            <p>- T2 disc blackish and marked with yellow or reddish cuticular patches, or T2 disc cuticle lighter reddish or yellowish brown. .................................................................................................. 17</p>
            <p>15 (14). Setal markings on T2 composed of golden circular spots (Brazilian Atlantic Forest) or T2 mostly covered with dense yellow to orange setae (countries West of Andes)................................................................. 16</p>
            <p>- Setal markings on T2 composed of small whitish setal stripes or spots (Colombia or Brazilian Caatinga and Cerrado).... 18</p>
            <p> 16 (15). Vertex smooth, without tubercles or projections of any kind; mesonotum and T2 disc with large patches of dense yellow to orange setae (Chile, Ecuador, and Peru)........................................................................................  Quwitilla Williams et al. in Bartholomay et al. 2019—three species, see Bartholomay et al. (2019b) </p>
            <p> - Vertex with large smooth tubercle posterolaterally; mesonotum with contrasting golden and black setae, T2 disc marked with large golden-orange spots composed of thickened bristle-like setae (Brazil)......................................................  Atlantilla Williams &amp; Bartholomay in Williams et al. 2020 —one species,  A. auriculata (Gerstaecker, 1874)</p>
            <p> 17 (14). Mesosoma pale reddish-brown; scutellar scale distinct and broad; vertex simple without tubercles or projections of any kind; genal carina present; lateral propodeal surface smooth and defined by irregular carina dorsally and posteriorly; body small, length 5 mm or less (widespread in South America)........  Tobantilla Casal, 1969 —nine species, see Williams et al. (2011) </p>
            <p>- With one or more of the following characters: mesosoma black; scutellar scale indistinct or absent; vertex armed with posterolateral tubercle; genal carina absent; lateral propodeal surface distinctly sculptured and not defined by carina dorsally and posteriorly; body length more than 5 mm ............................................................... 8</p>
            <p> 18 (15, 17). T2 disc with spots composed of setae only or with cuticular spots having similar sculpture to surrounding T2 disc sculpture (rarely encountered, in Colombia only, mesosoma sometimes reddish orange)..................................................  Dasymutilla Ashmead, 1899 , in part —four species, see Cambra et al. (2018), and Bartholomay et al. (2019b) </p>
            <p> - T2 disc with smooth yellow or reddish cuticular spots with sparse setae (common and widespread throughout South America; in Colombia with mesosoma blackish).......  Traumatomutilla André, 1901 —over 100 species, see Williams et al. (2017) </p>
            <p>Males</p>
            <p>1. Axilla armed with posteriorly directed tooth or lobe; eye subcircular; head unarmed ventrally; T1 shape narrowly petiolate and usually constricted with base of T2; tergal fringes composed of simple or brachyplumose setae ........................ 2</p>
            <p>- With one or more of the following characters: axilla unarmed; eye longitudinally ovate and notched; gena and/or hypostoma armed with distinct ventral tooth; T1 broadly sessile, subsessile, or disciform; tergal fringes composed of distinctly plumose setae........................................................................ other tribes (not treated here)</p>
            <p>2 (1). Mandible with ventral tooth or lobe basally................................................................ 3</p>
            <p>- Mandible unarmed ventrally............................................................................ 4</p>
            <p> 3 (2). Mandible with bilobate lamella dorsally; paramere posteriorly upcurved with apex pointing antero-dorsally (Argentina).......................................  Gogoltilla Williams et al. 2011 — One species,  G. chichikovi Williams et al. 2011</p>
            <p>- Mandible simple dorsally; paramere not upcurved, apex pointing posteriorly.............. other tribes (not treated here)</p>
            <p>4 (2). Specimens from North America, Central America, or Caribbean Islands (excluding Trinidad)......................... 5</p>
            <p>- Specimens from South America (including Trinidad)......................................................... 7</p>
            <p> 5 (4). Nearly all erect setae on body, particularly on scape and clypeus, brachyplumose; nearly entire body covered with moderately dense whitish setae; paramere posteriorly upcurved with apex pointing dorsally (Panama).................................................................................  Dasyphuta gen. nov. —one species:  D. gorgupea sp. nov.</p>
            <p>- Erect setae on body mostly simple; body setal color variable; paramere shallowly upcurved, apex pointing postero-dorsally. 6</p>
            <p> 6 (5). Body cuticle entirely black, covered only with black and silvery setae; tibial spurs whitish; free paramere length with relatively sparse setae anteroventrally (rarely encountered in Central America and Hispaniola).........................................................................................  Traumatomutilla André, 1901 , in part— three species </p>
            <p> - With one or more of the following characters: at least mesosoma or T2 with cuticle yellowish or reddish-brown; head, mesosoma, or tergites densely covered with golden to reddish setae; tibial spurs blackish; free paramere length with particularly dense brush of pale yellowish setae anteroventrally (common and widespread in North and Central America and Caribbean Islands).................................................................  Dasymutilla Ashmead, 1899 , in part— over 100 species — see Manley &amp; Pitts (2007), Cambra et al. (2018), and Manley et al. (2020) [The male of  Frigitilla panamensis Cambra et al. 2016 is not yet known, but will likely key out here with similar characters to those mentioned in couplet 9 below] </p>
            <p> 7 (4). Mandible thick and somewhat vertically oriented (with distinct dorsal carina continuous to inner apical tooth); parapenial lobe longer than penis valve (Chile, Ecuador, and Peru)................................................................  Quwitilla Williams, Bartholomay &amp; Cambra in Bartholomay et al. 2019—three species, see Bartholomay et al. (2019b) </p>
            <p>- Mandible narrowed and oblique apically (dorsal carina usually becoming obsolete before reaching inner apical tooth); parapenial lobe shorter than penis valve (widespread but rarely found West of the Andes).................................... 8</p>
            <p>8 (7). Paramere shallowly upcurved posteriorly, apex pointing postero-dorsally (brachyplumose setae generally rare and inconspicuous on body; head usually uniformly black; clypeus usually densely punctate and bidentate antero-ventrally)............... 9</p>
            <p>- Paramere sharply upcurved posteriorly, apex pointing dorsally or antero-dorsally (brachyplumose setae generally abundant and conspicuous, especially on scape and clypeus; vertex often with mesal reddish cuticular spot; clypeus usually with smooth bidentate or truncate shelf antero-ventrally, antero-ventrally unidentate, or antero-ventrally rounded).................. 14</p>
            <p> 9 (8). S2 mesally convexly swollen, with asetose mesal pit (T2 and S2 with cuticle largely orange-brown; paramere virtually straight, slightly upcurved posteriorly, with especially dense ventral setal brush on basal half; Bolivia, Brazil, and Peru)............................................  Frigitilla Williams in Bartholomay et al. 2015, in part —  F. simulatrix (Smith, 1879)</p>
            <p>- S2 flat or simply convex, if mesal pit present, then filled with setae............................................. 10</p>
            <p> 10 (9). Axillar lobe with posterior margin flat and truncate posteriorly, additionally armed with sharp down-pointed tooth along outer margin (Brazil)....................................................................................................  Atlantilla Williams &amp; Bartholomay in Williams et al., 2020 —one species,  A. auriculata (Gerstaecker, 1874)</p>
            <p>- Axillar lobe variable but without down-pointing tooth along outer margin....................................... 11</p>
            <p> 11 (10). Mesopleuron with lateral tooth-like tubercle; axilla posteriorly truncate and raised above level of depressed mesoscutellum; propodeum with conspicuous dorsolateral tubercle posterior to propodeal spiracle; penis valve with especially large pre-apical tooth, margin anterior to pre-apical tooth serrate ventrally (Bolivia, Brazil, and Paraguay)...................................................................  Leucospilomutilla Ashmead, 1903 —three species, see Williams et al. (2020) </p>
            <p>- Mesopleuron usually without tooth-like tubercle (if mesopleuron armed, then axilla dentate posteriorly or not especially raised above convex mesoscutellum); propodeum without dorsolateral tubercle posterior to propodeal spiracle; penis valve with smooth margin anterior to pre-apical ventral tooth.......................................................... 12</p>
            <p> 12 (11). S2 mesally flattened, without mesal pit; paramere dorsoventrally flattened basally (widespread in South America)....................................................  Cephalomutilla André, 1908 —four species, see Williams et al. 2022 </p>
            <p>- S2 convex (with or without mesal pit); paramere shape variable............................................... 13</p>
            <p> 13 (12). Rarely encountered, in Colombia only; with metasomal cuticle mostly reddish-orange and covered with dense red setae or metasoma predominantly black except T2 entirely pale yellow or marked with pale yellow cuticular patch.....  Dasymutilla Ashmead, 1899 , in part —four species, see Manley &amp; Pitts (2007), Cambra et al. (2018), and Bartholomay et al. (2019b) </p>
            <p> - Common and widespread throughout South America; Colombian species with metasomal cuticle uniformly black or mostly reddish-orange with sparse setae..................................  Traumatomutilla André, 1901 —over 100 species </p>
            <p>14 (8). Vertex black with reddish-orange spot mesally; clypeus, frons, and gena usually with extensive black setae (if head setae predominantly whitish, then T2 marked with round yellow cuticular spots)...................................... 15</p>
            <p>- Vertex uniformly blackish; clypeus, frons, and gena usually with silvery setae, at most interspersed with a few erect darker setae; T2 disc entirely black, reddish-black or orange (second labial palpomere not expanded, at least as long as wide).... 16</p>
            <p> 15 (14). T2 disc with cuticle uniformly black; second labial palpomere not expanded, at least as long as wide......................................................................  Reedomutilla Mickel, 1964 —three species, see Casal (1968) </p>
            <p> - T2 disc black, marked with two large circular yellow or reddish cuticular spots; second labial palpomere mesally expanded, at least twice as wide as long..................................  Suareztilla Casal, 1968 —Nine species, see Casal (1968) </p>
            <p> 16 (14). Clypeus with longitudinal elevated crest medially, with anteromedial margin strongly projected medially (Fig. 8); body cuticle entirely black (Caatinga region of Brazil).................  Goncharovtilla gen. nov. —one species,  G. oblomovi sp. nov.</p>
            <p> - Clypeus without longitudinal elevated crest medially, with anteromedial margin projected in a single raised tubercle or shallowly emarginate (see Figs 35–36 in Williams et al. 2011); metasomal cuticle largely reddish-orange, at least on T2 (widespread in South America, but absent from Caatinga)...............  Tobantilla Casal, 1969 —nine species, see Williams et al. (2011) </p>
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	https://treatment.plazi.org/id/03C687DDB93D2241FF38FD70FA25CBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Williams, Kevin A.;Cambra, Roberto. A.;Bartholomay, Pedro R.;Lopez, Vinicius M.;Brothers, Denis J.;Martins, Herbeson O. J.;Ferreira, Vinina S.;Koroiva, Ricardo;Guillermo-Ferreira, Rhainer	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
