taxonID	type	description	language	source
03C687DDB9282254FF38F8F0FDB9C8DA.taxon	type_taxon	Type species. Goncharovtilla oblomovi Williams, Bartholomay & Lopez, sp. nov., designated here.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9282254FF38F8F0FDB9C8DA.taxon	diagnosis	Diagnosis. FEMALE. Females of Goncharovtilla can be separated from other South American Dasymutillini by having the posterior surface of the head concave and separated from the dorsal surface by a sharp transversely arcuate carina that is thickened laterally. Additionally, the body is blackish with a contrasting pattern of black and whitish setae, the genal carina is distinct anteriorly, the mesosoma has a thick scutellar scale and similarly large, thickened carina antero-lateral to the scale, the T 2 disc has the cuticle uniformly blackish, the body is covered with numerous brachyplumose setae, and T 6 has a defined pygidial plate. MALE. Males can be differentiated from other Dasymutillini by the following characters: clypeus with longitudinal elevated crest medially and anterior / ventral margin strongly projected medially, mandible lacking notch or tooth ventrally, mesopleuron not swollen or projected into conspicuous tubercle in dorsal half, S 1 with strong tooth-like longitudinal carina, S 2 lacking seta-filled pit, metasoma lacking any reddish or orange markings. The genitalia also have useful diagnostic features: the paramere is strongly upcurved apically and densely setose ventrally, somewhat dorsoventrally depressed; the cuspis has an elongate brush of posteroventrally directed setae; and the penis valve has two simple teeth posteroventrally.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9282254FF38F8F0FDB9C8DA.taxon	description	Description. FEMALE. Integument. Usually reddish to dark brown, sculpture mostly concealed by dense setation, apparently densely and finely sculptured where visible. Setae predominantly brachyplumose to simple. Head. Subrounded, strongly concave posteriorly, vertex margined by transverse arcuate carina separating dorsal and posterior surfaces; finely and densely sculptured where visible. Occipital carina apparently deeply recessed, not clearly visible. Eye hemispherical, strongly protruding, surface shining, ommatidia clearly discernible. Antennal scrobe poorly developed, short, quadrate, lacking dorsal and lateral carinae. Clypeus slightly protruded basomedially, anteroventral margin virtually straight. Malar space greater than basal height of mandible. Genal carina present. Hypostomal carina strong, extending to posterior mandibular condyle. Scape simple. Pedicel almost as long as wide, shorter than F 1; F 1 longer than wide and longer than F 2. Mandible weakly curved basally, slender, tapering, with small inconspicuous tooth subapically, no ventral basal tooth, notch, or flange. Maxillary palp six-segmented; labial palp four-segmented; basal palpomeres strongly flattened and broadened, densely covered with short erect setae. Mesosoma. Longer than wide, subrectangular, evenly and slightly narrowed posterad, lacking clear constriction on mesonotum in dorsal view, dorsum convex to flat; propodeum weakly rounded posteriorly, disk and declivity indistinct; dorsal and posterior faces of propodeum densely and coarsely sculptured; lateral face of propodeum weakly and sparsely sculptured to unsculptured. Pronotal dorsal face (excluding anterior collar) with anterior margin straight in dorsal view, reclined in lateral view; epaulet at extreme lateral margin of anterior pronotal margin, scarcely distinguished from the latter; humeral angle rounded; posterior margin indiscernible. Scutellar scale present, with conspicuous pair of transverse anterolateral carinae. Pronotal-mesopleural suture distinct, partly concealed by dense setation. Vertical mesopleural ridge present, scarcely swollen, rounded, areolate throughout. Metapleuron predominantly sculptured, almost smooth dorsally. Metapleural-propodeal suture present, inconspicuous, partly concealed by dense setae; endophragmal pit small, inconspicuous, obscured by setae, surrounding area virtually flat. Metasternal process subtriangular, slightly less than a third of metacoxal length. Legs. Tibial spur formula 1 – 2 – 2. Foreleg with tarsal comb. Meso- and metatibiae each with two rows of prominent stout spines, few spines in each row, spines smaller basad; apical spurs narrow, finely serrate laterally, outer spur shortened. Metacoxa with strong slightly arcuate carina on inner margin, ending abruptly before coxal apex. Metasoma. Surface finely and densely sculptured, sparser and coarser on sterna and sparser anterolaterally on T 2. T 1 shorter than wide, its width about 1 / 3 the width of T 2, strongly constricted on anterior third and evenly expanded from constriction apicad, weakly convex in lateral view, anterior and dorsal faces distinct; anterior auricle prominent, vertically lamellate but apparently dentate from above. Lateral felt line of T 2 wide. T 6 with conspicuous pygidial plate, slightly convex, large, subrounded, defined laterally and apically by carinae, longitudinally and irregularly striate-rugose throughout. S 2 evenly convex, with conspicuous medial longitudinal ridge on anterior third, without posterior transverse groove, lacking lateral felt line. S 6 posterior margin with strong bilobate projection. MALE. Integument. Mostly blackish to brownish-black, mostly concealed by dense setation, with strong dense sculpture where visible. Setae mostly brachyplumose, with many simple setae interspersed throughout. Head. Roundly transverse, as wide as mesosoma. Occipital carina distinct dorsally. Vertex slightly swollen posteriorly. Eye broadly hemispherical, strongly protruding, surface shining, ommatidia discernible. Ocelli small. Antennal scrobe shallow, not reaching eye margin, with transverse lamellate carina on dorsal margin. Clypeus slightly depressed immediately below insertion of antennae, conspicuously convex apicomediad, ventral half with conspicuous longitudinal crest medially, almost carinate, terminating in strongly protruding acuminate anterior / ventral margin. Malar space shorter than basal height of mandible. Genal carina absent. Hypostomal carina evident, evenly high throughout, extending in a smooth curve to posterior mandibular condyle. Scape bicarinate ventrally. Pedicel approximately as wide as long, shorter than F 1, which slightly longer than F 2. Mandible evenly weakly curved, bidentate apically, subapical tooth minute, lacking ventral tooth or notch. Maxillary palp 6 - segmented; labial palp 4 - segmented; basal palpomeres strongly flattened and broadened, densely covered with short setae. Mesosoma. Surface mostly covered with dense sculpture where visible. Pronotal epaulet weakly defined, reduced to a row or tuft of short setae; humeral angle rounded in dorsal view. Tegula broadly subcircular, convex, almost entirely smooth, except anterior and inner margins sparsely sculptured. Mesoscutum lacking notaulus; parapsis present, inconspicuous, short. Mesoscutellum flat, declivous in lateral view; axilla generally flat dorsally, broadly connected to mesoscutellum, terminating in truncate / rounded lobe, with oblique longitudinal lamella connecting its ventral surface to mesoscutellum; dorsal surface of axilla smooth. Metanotum usually completely concealed by dense setae. Propodeum evenly convex in lateral view, dorsal and lateral faces indistinct. Metasternal process convergent to unidentate apex, shorter than metacoxal length. Wings. Fore wing with elongate sclerotized pterostigma; marginal cell slightly elongate, truncate apically; three closed submarginal cells, apical veins of third often inconspicuous or obliterated. Legs. Tibial spur formula 1 – 2 – 2. Mid and hind tibiae with two rows of weak spines; apical spurs finely serrated apically. Metacoxa with weak inner carina. Metasoma. Sculpture generally dense and fine; T 1 longer than wide, its width about 1 / 3 width of T 2, anterior and dorsal faces distinct, not constricted apically. T 2 with lateral felt line broad and short. S 1 with marked longitudinal median carina terminating posteriorly in subacute projection. S 2 in lateral view flattened weakly convex, lacking felt line; slightly elevated and projected anteromedially; with no indication of mesal seta-filled pit. Hypopygium longer than wide, surface almost flat to shallowly convex, posterior margin projected medially into sharp acute projection. Genitalia. Paramere long, free length longer than length from base of genital capsule to apex of parapenial lobe, sub-cylindrical and strongly upcurved at apical third; dorsoventrally flattened at basal 2 / 3; acute apically. Parapenial lobe well-developed. Cuspis short, somewhat dorsoventrally flattened, approximately 1 / 3 longer than digitus, paracuspis a somewhat elongated lobe. Digitus narrowly digitiform, slightly wider basad. Penis valve inner surface concave; with two sharp apical teeth on ventral margin; ventral teeth widely separated; without distinct setae.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9282254FF38F8F0FDB9C8DA.taxon	distribution	Distribution. Northeastern Brazil (Pernambuco, Ceará, and Maranhão).	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9282254FF38F8F0FDB9C8DA.taxon	etymology	Etymology. This genus is named after the Russian author Ivan Goncharov (1812 – 1879) and the type species is named after one of his novels (see below). This name references the initial discovery of the genus in the Brazilian city of Russas (Portuguese for Russians [fem.]), Ceará, and tribal affiliation with Gogoltilla Williams, Brothers & Pitts, 2011 from Argentina (named after the Ukrainian-Russian author Nikolai Gogol). The name is feminine.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9282254FF38F8F0FDB9C8DA.taxon	discussion	Remarks. This genus was included in an unpublished molecular phylogeny of American Dasymutillini (Williams 2012) as an undetermined Traumatomutilla (Genbank accession numbers: JQ 945822, JX 002751, JQ 945901). In that analysis, it was recovered as sister to Tobantilla drosos Williams, Brothers & Pitts, 2011, a male-based species from Argentina, and this clade was recovered as the sister to the remaining New World Dasymutillini (excluding Lomachaeta Mickel, 1936). The male of this genus is structurally similar to other Tobantilla species, particularly in the upcurved paramere and shelf-like clypeus. Females differ from Tobantilla in many features, including body size (all Tobantilla females are less than 6 mm in length), coloration (Tobantilla females have a pale brown mesosoma and yellowish cuticular patches on T 2), and head structure (Goncharovtilla gen. nov., is unique among Dasymutillini in having the posterior margin of the vertex with a sharp transverse arcuate ridge). Furthermore, the genetic distance between Goncharovtilla and Tobantilla is comparable to that observed between various distinct genera within Dasymutillini and greater than that observed within any single genus of Dasymutillini (Williams 2012). In the best available key to Neotropical mutillid genera (Brothers 2006), females of Goncharovtilla key out in couplet 23 as “ Dasymutilla / Traumatomutilla ”. The unique head structure separates Goncharovtilla from females of those genera. Males of Goncharovtilla also key to “ Dasymutilla / Traumatomutilla ” in couplet 65. They can be separated from those genera by the upcurved paramere and shelf-like clypeus. The female head morphology is apparently unique, with its transverse arcuate carina separating the dorsal and posterior surfaces. Some North American species in the Dasymutilla monticola species-group approximate this head morphology; unlike Goncharovtilla, all these species lack a genal carina and have the body color predominantly reddish-brown (Williams 2012).	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	description	(Figs 1 – 16, 18 – 20)	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	diagnosis	Diagnosis. As for genus.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	description	Description. FEMALE. Body length. 6.5 mm in holotype (4.0 – 9.5 mm). Head. Posterior margin shallowly concave in dorsal view. Vertex width 0.8 × pronotal width. Eye height in frontal view 1.2 × distance from its ventral margin to mandibular condyle. Head surface entirely concealed by dense setation, except gena basally with sparse fine punctures. Mandible with small subapical tooth. Dorsal and lateral scrobal carinae absent. Antennal tubercle mostly asetose, shallowly and irregularly rugose. F 1 1.6 × pedicel length; F 2 1.3 × pedicel length. Genal carina present, ending before hypostomal carina. Occipital area concave. Dorsal and posterior surfaces of vertex separated by sharp transversely arcuate carina. Mesosoma. As long as wide. Surface completely concealed by dense setation dorsally. Anterior face of pronotum densely and conspicuously covered with longitudinal striae. Humeral carina present, narrowly separated from low inconspicuous epaulet, antero-lateral corner of pronotum rounded in dorsal view. Pronotal spiracle virtually flat against lateral margin of pronotum. Ratios of widths at humeral angles, pronotal spiracles, widest point of mesonotum, narrowest point of mesonotum and propodeum posterior to propodeal spiracles 1.00: 1.05: 1.15: 0.91: 0.94. Lateral margin of mesosoma slightly diverging anterad towards pronotal spiracle and converging slightly towards humeral angle after pronotal spiracle. Scutellar scale present, well-developed, wide, more than twice width of poorly defined anterolateral carinae, separated from the latter. Posterior and dorsal faces of propodeum undefined, propodeum almost vertical in lateral view. Propodeal spiracle virtually flat against lateral margin of mesosoma; no post-spiracular area. Metasoma. Dorsal sculpture, except pygidium and laterally on T 2, completely concealed by dense setation; T 2 densely and coarsely foveolate-punctate laterally. T 1 0.3 × as wide as T 2. T 2 length 0.9 × its width. T 2 maximum width anterior to midlength. S 1 with conspicuous short longitudinal carina, terminating in short tooth-like projection posteriorly; S 2 – 6 densely and coarsely foveolate-punctate. Pygidium subrounded, almost as broad as long, broadest nearer base; defined by strong arcuate lateral carina throughout its extension; surface with widely spaced longitudinal striations; interstices finely granulose. Coloration and variations. Body and appendages black to reddish-brown, except mandible, antennal tubercle, and flagellum partially yellowish-brown. Head setae silvery-white, dense, predominantly appressed and decumbent, with few sparse brownish erect setae, and dense decumbent brownish setae below occipital carina. Mesosomal dorsum with silvery-white setae laterally and black to brownish-black setae medially; mostly dense decumbent and appressed setae interspersed with sparse erect setae throughout; lateral face of pronotum, mesopleuron, metapleuron, and lateral face of propodeum silvery-white. Metasoma with interspersed appressed, decumbent, and erect silvery-white setae, except anterior half of T 2 disc, posteromedial margin of T 2 and medially on T 3 – 6 with interspersed appressed, decumbent and erect brownish-black setae. Three different patterns for white setae distribution on the metasoma have been observed in this species: the holotype has a complete band of silvery-white setae on the posterior margin reaching the fringe only laterally, thus the fringe is black medially; one variant has both the silvery-white band and silvery-white fringe narrowly interrupted medially with black; a third form has both the silvery-white band and silvery-white fringe more widely interrupted medially. MALE. Body length. 9.9 in allotype (6.0 – 10.0) mm. Head. Transverse, subrectangular in dorsal view; posterolateral angles rounded; vertex width 0.85 × pronotal width. Eye almost circular. Ocelli small; OOD 3.7 × DLO, IOD 5.0 × DLO. Occipital carina distinct. Sculpture predominantly concealed by dense setation, coarsely and densely foveolate-punctate where visible. Gena ecarinate. Antennal scrobe concave to eye margin, with narrow transverse dorsal scrobal carina. Clypeus concave sublaterally, elevating slightly laterad and, especially, mediad; with conspicuous elevated longitudinal crest medially; ventral margin strongly acuminate, terminating medially in subacute projection; coarsely and sparsely punctate throughout, punctures slightly dense lateromedially. Scape bicarinate. F 1 2.1 × pedicel length; F 2 1.7 × pedicel length. Mandible slightly tapering apicad, smoothly curved medially; with low inconspicuous carina anterodorsally terminating before apical third; inconspicuously bidentate apically, subapical tooth minute; lacking ventral notch and projections. Mesosoma. Epaulets poorly defined, broadly disconnected from humeral carina, virtually flat against anterior margin of pronotum. Anterior face of pronotum mostly asetose, short, poorly and coarsely distinguished from dorsal face, indistinctly sculptured, with medial longitudinal slightly concave area. Tegula convex, mostly glabrous and impunctate except for dense punctures anteriorly and along inner margin. Dorsal sculpture of pronotum, mesoscutellum, and metanotum concealed by dense setation. Mesoscutellum flat, sloping posterad in lateral view. Axilla projected posterolaterally as truncate / rounded lobe broadly connected to mesoscutellum; with oblique longitudinal lamella connecting its ventral surface to mesoscutellum; dorsal surface of axilla smooth. Metanotum equally long throughout. Propodeum convex, dorsal sculpture concealed by dense setation at dorsal third, densely areolate where visible; posterolateral margin smoothly rounded; dorsal face indistinguishable from posterior face. Lateral face of pronotum simply punctate. Mesopleuron mostly concealed by dense setation, densely and coarsely micropunctate where visible; smoothly elevated medially, without any conspicuous projections. Metapleuron mostly concealed by dense setation, sparsely micropunctate to smooth where visible. Lateral face of propodeum smooth to sparsely and shallowly foveolate to areolate posterodorsad. Wings. Fore wing with moderate elongate sclerotized pterostigma; length of pterostigma approximately twice its width; marginal cell elongated, its length approximately twice its width, truncate apically; three submarginal cells, apical veins of third cell vestigial. Legs. Densely setose, setae shorter and denser on tibiae and tarsi; lateral margin of spurs finely serrate. Metasoma. T 1 0.1 × as wide as T 2. T 2 length 1.0 × its width. T 1 densely and coarsely foveolate-punctate. T 2 densely and finely foveolate-punctate. Sculpture of T 3 – 7, except pygidium, concealed by dense setation. S 1 densely and finely foveolate-punctate. S 2 densely and coarsely foveolate-punctate, sculpture sparser posterad. S 3 – 6 densely foveolate-punctate to punctate. S 8 punctate, longer than broad, apical margin strongly acuminate. Genitalia. Ratios of free length of paramere, cuspis and digitus, 1.0: 0.3: 0.2. Parameres strongly convergent and broad in dorsal view, except at apical fourth where parallel and abruptly narrowed; dorsoventrally flattened, except subcylindrical apical third; in lateral view slightly curved ventrally, abruptly and strongly upcurved at apical fourth; with dense tuft of long setae on basal half, setae abruptly shorter and sparser, almost absent, on apical half. Cuspis short, stout, “ clavate ”, obscured by base of paramere in dorsal view, only partially visible in ventral view due to strong setation of paramere; slightly wider apicad, subparallel elsewhere; with dense long setae apicoventrally. Paracuspis short, about ¼ length of cuspis, subrounded, with few sparse setae. Digitus slightly elongate, weakly curved inward in dorsal view and slightly up-curved in lateral view, laterally compressed; apparently asetose. Penis valve concave on internal surface, with pair of acute teeth apicoventrally; without lateral pocket, asetose; apical distance between teeth 0.2 × length of valve. Coloration and variations. Body setae silvery-white, except area between posterior margin of ocellar triangle and posterior head margin, mesonotum, posterior half of T 2 disc, and T 5 – 7 with brownish-black setae. No significant variations were observed in the specimens examined.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	distribution	Distribution. Brazil (Pernambuco, Ceará, and Maranhão).	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	etymology	Etymology. Named after the fictional character Ilya Ilyich Oblomov, the particularly lazy main character of the novel Oblomov by Ivan Goncharov. The first recorded specimen was collected in Russas, Ceará in 2010 and was almost immediately recognized by KAW as a new species. In a phylogenetic analysis of Dasymutillini (Williams 2012), it was recognized to be a new genus. It took 50 pages for Oblomov to even rise from his bed, and it has taken 14 years for KAW to get around to naming and describing this spectacular wasp.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	materials_examined	Material examined. [5 ♀ 3 ♂] Type material. Holotype, ♀, BRAZIL, Pernambuco, Petrolina, 09 ° 19 ’ 44.2 ” S 42 ° 33 ’ 30 ” W, 17. V. 2019, coleta manual, H. O. J. Martins leg, MZSPHYM 0136464 (MZSP; Fig. 18). Paratypes: 1 ♀, same data as holotype except MZSPHYM 0145123 (MZSP); 1 ♂, allotype, same data as holotype except MZSPHYM 0136463 (MZSP); 1 ♂ same data as holotype except MZSPHYM 0145122 (MZSP); 1 ♀, same data as holotype except Hym – 002 / 0051, V. 2018 (LESTES); 1 ♂, same data as holotype except Hym – 002 / 0048, 24 – 28. III. 2018 (LESTES); 1 ♀, Ceará, BR 116, 5 km SW Russas, 04 ° 58.267 ’ S 38 ° 00.444 ’ W, 30 m, 22. VI. 2010, C. Waichert col. (UFES; voucher JP 1779). Other material. BRAZIL: 1 ♀, Maranhão, Alto Parnaíba, Estação 8, 13. VIII. 2013, L. P. Amaral-Neto (DZUP # 299378; Fig. 19); 1 ♀, Riachão, Fazenda Altos, 18 – 22. VIII. 2009, Varrediura, F. Limeira-de-Oliveira leg. (INPA; Fig. 20).	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB92A2252FF38F8D3FB01C982.taxon	discussion	Remarks. In both sexes, this species closely resembles Traumatomutilla bifurca (Klug, 1821) in coloration (Bartholomay et al., 2019 c) and many other mutillid species in the Brazilian Caatinga. The head structure of the female (Figs 1 – 3) and genitalia of the male (Figs 13 – 16) differentiate Goncharovtilla oblomovi sp. nov. from any other species in the Caatinga and any known Traumatomutilla species. The sex association is based on collection of males and females at the same locality on the same date in Pernambuco, demonstrating similar morphology (e. g., size and coloration). This association was subsequently confirmed through DNA sequence comparisons. The Maximum Likelihood (ML) phylogeny is illustrated in Fig. 17, with posterior probability values and bootstrap values provided. The identical sequences of two females and one male corroborate the identification of a single species, G. oblomovi sp. nov. Additionally, morphological similarities between both sexes and related genera further validate the sex association (see remarks for the genus above). The female specimens from Maranhão are not included as paratypes. They differ from the holotype and paratypes in setal coloration, having the white setal band of T 2 widely interrupted by black setae (Figs 18 – 20). They also differ slightly from one another in the shape and thickness of the white setal bands on T 2 – 4. Future studies, particularly if males are associated with these forms, could reveal them to belong to a separate species but, with the limited material available, they appear to form an incomplete gradient with the type series. With a nod to Goncharov’s Oblomov, we here employ the “ lazy ” approach of treating them as variants of a single species.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.taxon	type_taxon	Type species. Dasyphuta gorgupea Cambra, Williams & Brothers, sp. nov., designated here.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.taxon	diagnosis	Diagnosis. FEMALE: Females of Dasyphuta can be separated from the other American Dasymutillini genera by the following combination of characters: the mesosoma is more or less uniformly sculptured with deep small areolations, both dorsally and laterally, dorsally with sparse brachyplumose setae, and without a scutellar scale; the T 1 shape is narrowly petiolate; and T 6 is smooth and convex without a pygidial plate. MALE: Males of Dasyphuta can be separated from other Dasymutillini by the clypeus forming a more or less flat and densely punctate anteriorly bidentate shelf, the axilla narrowly spinose posteriorly, the pygidium smooth and convex without lateral carinae, the genitalic paramere strongly upcurved posteriorly, and the presence of erect brachyplumose setae on numerous portions of the body.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.taxon	description	Description. FEMALE. Integument. Reddish brown, sculpture clearly visible, generally confluently foveolate. Setae predominantly brachyplumose to simple. Head. Subrounded, lacking posterolateral tubercles. Eye hemispherical, strongly protruding, surface shining, ommatidia clearly discernible. Antennal scrobe poorly developed, short, quadrate, lacking dorsal and lateral carinae. Clypeus slightly protruded basomesally, anteroventral margin virtually straight. Malar space greater than basal height of mandible. Hypostomal carina strong, extending to posterior mandibular condyle. Scape simple. Pedicel as long as wide, shorter than F 1; F 1 longer than wide and similar in length to F 2. Mandible nearly straight, slender, tapering, with small inconspicuous tooth subapically, no ventral basal tooth, notch, or flange. Maxillary palp six-segmented; labial palp four-segmented; basal palpomeres strongly flattened and broadened, densely covered with short erect setae. Mesosoma. Slightly longer than wide, pear-shaped, mesonotum bulging laterally, weakly constricted near pronotal and propodeal spiracles, dorsum convex; propodeum rounded posteriorly, disk and declivity indistinct; mesosomal dorsum and pleura more or less uniformly confluently foveolate. Pronotal dorsal face (excluding anterior collar) with anterior margin convex in dorsal view, flat in lateral view; epaulet distantly separated from humeral angle; humeral angle forming relatively sharply defined obtuse angle; posterior margin indiscernible. Scutellar scale absent. Pronotal-mesopleural suture indistinct. Mesopleuron sculptured throughout; vertical mesopleural ridge scarcely defined. Metapleuron predominantly sculptured, except narrow surface surrounding endophragmal pit. Metapleural-propodeal suture inconspicuous. Metasternal process subtriangular, slightly less than a third of metacoxal length. Legs. Tibial spur formula 1 – 2 – 2. Foreleg without tarsal comb. Meso- and metatibiae each with one row of prominent stout spines, few spines in each row, spines smaller basad; apical spurs narrow, finely serrate, outer spur shortened. Metasoma. Surface mostly confluently foveolate, sparser on sterna. T 1 nearly as long as wide, its width about 1 / 3 width of T 2, strongly constricted on anterior third and nearly parallel-sided from constriction apicad, petiolate in lateral view; anterior auricle prominent, vertically lamellate but apparently dentate from above. Lateral felt line of T 2 linear. T 6 convex without pygidial plate, smooth, except anterolateral portions punctate and setose. S 2 evenly convex, with indistinct longitudinal ridge anteromesally, without posterior transverse groove, lacking lateral felt line. S 6 posterior margin with strong bilobate projection. MALE. Integument. Mostly blackish to brownish-black, mostly concealed by dense setation, with strong dense sculpture where visible. Setae mostly brachyplumose, with many simple setae interspersed throughout. Head. Rounded, as wide as mesosoma. Occipital carina distinct. Vertex slightly swollen posteriorly. Eye broadly hemispherical, strongly protruding, surface shining, ommatidia discernible. Ocelli small. Antennal scrobe shallow, not reaching eye margin, with transverse lamellate carina on dorsal margin. Clypeus mostly flat and punctate, with slightly raised area below insertion of antennae and bidentate anterior / ventral margin. Malar space shorter than basal height of mandible. Genal carina absent. Hypostomal carina evident, evenly high throughout, extending in a smooth curve to posterior mandibular condyle. Scape bicarinate. Pedicel approximately as wide as long, slightly shorter than F 1, which much shorter than F 2. Mandible evenly weakly curved, bidentate apically, subapical tooth minute, lacking ventral tooth or notch. Maxillary palp 6 - segmented; labial palp 4 - segmented. Mesosoma. Surface mostly covered with dense sculpture. Epaulet weakly defined, reduced to a row or tuft of short setae; humeral angle rounded in dorsal view. Tegula broadly subcircular, convex, almost entirely smooth, except anterior and inner margins sparsely sculptured. Mesoscutum lacking notaulus; parapsis present, inconspicuous, short. Mesoscutellum shallowly convex with mesal longitudinal carina; axilla generally flat dorsally, narrowly connected to mesoscutellum, terminating in elongate spine; dorsal surface of axilla areolate. Metanotum concealed by dense setae. Propodeum evenly convex in lateral view, dorsal and lateral faces distinct but not separated by carina or obvious differences in sculpture. Metasternal process convergent to unidentate apex, shorter than metacoxal length. Wings. Fore wing with elongate sclerotized pterostigma; marginal cell short, truncate apically; three closed submarginal cells, apical veins of third largely inconspicuous or obliterated. Legs. Tibial spur formula 1 – 2 – 2. Mid and hind tibiae with three rows of thickened bristle-like setae, apparently without spines; apical spurs finely serrated. Metacoxa with weak inner carina. Metasoma. Sculpture generally coarse; T 1 petiolate, longer than wide, its width about 1 / 3 width of T 2, anterior and dorsal faces distinct, not constricted apically. T 2 with lateral felt line linear. S 1 with marked longitudinal median carina. S 2 in lateral view weakly convex, lacking felt line; with no indication of mesal seta-filled pit. Hypopygium elongate sub-triangular, surface shallowly convex, posterior margin projected mesally into sharp acute projection. Genitalia. Paramere with free length subequal to length from base of genital capsule to apex of parapenial lobe, sub-cylindrical and strongly upcurved at apical third; acute apically. Parapenial lobe well-developed. Cuspis short, somewhat dorsoventrally flattened, similar in length to digitus, paracuspis a small tubercle. Digitus narrowly digitiform, with widened apical knob. Penis valve inner surface concave; with two sharp apical teeth on ventral margin; ventral teeth widely separated; without distinct setae.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.taxon	distribution	Distribution. Panama.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.taxon	etymology	Etymology. From the Greek δασύς (dasus) “ hairy ”, and the genus name Ephuta Say. In reference to the tribal placement in Dasymutillini along with the superficial resemblance to typical Ephuta in body size, coloration, and slender first metasomal segment in both sexes. The name is feminine.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB922225BFF38FBD8FC0DC9C8.taxon	discussion	Remarks. This genus is similar to Tobantilla in body size, coloration, and many structural features, but that genus is known from South America only. The undefined pygidium in both sexes, absence of scutellar scale in females, and spinose axilla of males provide strong justification to treat Dasyphuta gen. nov., as a separate genus. In the best available key to Neotropical mutillid genera (Brothers 2006), females of Dasyphuta terminate in couplet 35, where their combination of characters does not match either option. Unlike Lomachaeta, females of Dasyphuta lack a genal carina, have densely foveolate-punctate mesosomal pleura, and have the T 1 shape narrowly petiolate. Unlike the genera in couplet 36, females of Dasyphuta lack distinctly plumose setae on the tergal fringes and the body length is less than 6 mm. Males of Dasyphuta terminate in couplet 69, where their combination of characters does not match either option. They can be separated from the genera in couplet 70 by the bidentate mandible and uniformly black head; they can be separated from the genera in couplets 71 and 72 by the upcurved paramere, flattened indistinct epaulet, and narrowly petiolate T 1 shape.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	description	(Figs 21 – 42)	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	diagnosis	Diagnosis. As for genus.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	description	Description. FEMALE. Body length 5.0 in holotype (4.0 – 5.0) mm. Head. Posterior margin rounded. Vertex width subequal to pronotal width. Eye height in frontal view 1.9 × distance from its ventral margin to mandibular condyle. Head predominantly concealed by dense setae, confluently foveolate where visible. Mandible with small subapical tooth. Dorsal scrobal carina absent; lateral scrobal carina absent. Antennal tubercle asetose, faintly sculptured. Flagellomere 1 1.2 × pedicel length; flagellomere 2 1.2 × pedicel length. Genal carina apparently absent, sub-defined by irregular puncture margins. Occipital carina defined, equally wide throughout. Vertex lacking tubercles. Mesosoma. Mesosoma 1.2 × as long as wide. Mesosomal sculpture confluently foveolate. Humeral carina present, connected to short tooth-like epaulet by irregular carina, antero-lateral corner of pronotum angular in dorsal view. Pronotal spiracle slightly raised against lateral margin of pronotum. Ratios of widths at humeral angles, pronotal spiracles, widest point of mesonotum, narrowest point of mesonotum and propodeum posterior to propodeal spiracles 80: 85: 88: 80: 74. Lateral margin of mesosoma slightly emarginate anterior to propodeal spiracle, smoothly diverging anterad and converging slightly posterior to pronotal spiracle. Propodeal spiracle slightly raised against lateral margin of mesosoma; post-spiracular area not defined. Scutellar scale absent. Posterior and dorsal faces of propodeum poorly defined, posterior face apparently longer than dorsal face. Metasoma. Dorsal sculpture mostly confluently foveolate. Ratios of width of T 1, width of T 2 and length of T 2 32: 97: 108. T 2 maximum width anterior to midlength; its base conspicuously swollen over apex of T 1. S 1 with conspicuous longitudinal carina, highest anteriorly; S 2 sculpture slightly sparser than T 2 sculpture; S 3 – 6 densely coarsely and irregularly foveolate-punctate to simply punctate; S 2 with subapical transverse slope and anteromedial longitudinal ridge vestigial. T 6 convex, without pygidial plate, sculpture smooth except for punctate and setose anterolateral portions. S 6 sharply bidentate postero-mesally. Coloration and variations. Body mostly reddish brown; legs and antenna lighter orange; T 2 with paired posterolateral circular yellow integumental spots. Head covered with dense pale golden setae. Mesosomal setae sparse, mostly silvery or golden except dorsally with large mesal patch (with posteromesal golden notch) and dorsolateral propodeal patches of sparse black setae. Metasoma setae sparse, mostly pale silvery except T 2 disc (except lateral margins, yellow cuticular spots, and sub-circular anterolateral patch with pale golden setae) and fringes of T 2 and T 3 mesally black, and fringes of T 2 – 5 brighter golden. Females vary slightly in intensity of the golden setae and some have the frons, mesosoma, and T 2 disc variably darker brownish. MALE. Body length 6.0 in allotype (5.0 – 7.0) mm. Head. Rounded, width subequal to pronotal width. Eye almost circular. Ocelli small; OOD 4.5 × DLO, IOD 1.2 × DLO. Occipital carina distinct. Sculpture predominantly concealed by dense setation, areolate-punctate where visible. Gena ecarinate. Antennal scrobe with prominent transverse dorsal scrobal carina. Clypeus mostly flat and punctate, with slightly raised area below insertion of antennae and bidentate anterior / ventral margin. Scape bicarinate. Flagellomere 1 subequal to pedicel length; flagellomere 2 1.9 × pedicel length. Mandible narrow, bidentate; lacking ventral notch and projections. Mesosoma. Epaulets poorly defined, virtually flat against anterior margin of pronotum. Anterior face of pronotum mostly asetose, indistinctly sculptured, without mesal notch. Tegula convex, mostly glabrous and impunctate except for dense punctures anteriorly and along inner margin. Dorsal sculpture of pronotum, mesonotum, scutellum, axilla and metanotum areolate. Scutellum slightly convex, with mesal longitudinal carina. Axilla produced posterolaterally with sharp spine. Metanotum equally wide throughout. Propodeum convex, sculpture partially concealed by dense setation, areolate without enclosed area; posterolateral margins smoothly rounded; dorsal face rounded into posterior face. Lateral face of pronotum densely and confusedly areolate-punctate to micropunctate; mesopleuron densely and coarsely areolate-punctate to foveolate-punctate; mesopleuron more or less evenly convex, without any conspicuous projections. Metapleuron largely areolate with dorsal area partly smooth. Wings. Fore wing with elongate sclerotized pterostigma; marginal cell short, truncate apically; three closed submarginal cells, apical veins of third largely inconspicuous or obliterated. Legs. Mid and hind tibiae with three rows of thickened bristle-like setae, apparently without spines. Metasoma. T 1 0.4 × as wide as T 2. T 2 length 0.9 × its width. Dorsal metasomal sculpture, except pygidium, foveolate-punctate to punctate. S 1 with low longitudinal mesal carina, equally high throughout. S 2 sculpture slightly sparser than that of T 2, without seta-filled pit mesally. S 3 – 6 densely punctate. S 8 punctate, elongate triangular, apical margin with acute tooth-like projection mesally. Genitalia. Parapenial lobe pronounced apically, shorter than penis valve. Ratios of free length of paramere, cuspis and digitus 69: 11: 11. Paramere virtually straight and cylindrical throughout in dorsal view, apical third tapering towards acute apex; in lateral view conspicuously curved dorsally at apex; with sparse tuft of setae basoventrally. Cuspis short, “ paddle-like ”, slightly wider apicad; basically straight; with brush of elongate plumose setae emerging from posterior margin. Paracuspis a short asetose rounded tubercle. Digitus elongate, digitiform with apical knob, slightly up-curved in lateral view, apparently asetose. Penis valve concave on internal surface, with pair of acute teeth apicoventrally; apical tooth well defined, asetose; basal tooth less defined, short, without lateral pocket, asetose; apical distance between teeth 0.2 × length of valve. Coloration and variations. Body black to dark brown, except mandible, venter of flagellum and tibial bases pale yellow-brown. Setae predominantly silvery-white, except transverse black band on posterior half of T 2 disc and apical tergites variably blackened. There is subtle variation in the extent of black setae on the apical tergites and various body regions have scattered erect darker setae.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	distribution	Distribution. Panama (Panama and Coclé).	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	materials_examined	Material examined. [7 ♀, 24 ♂] Type Material: Holotype, 1 ♀, PANAMA, PANAMA, Chorrera, Llano Largo, 2. IV. 1990, R. Cambra (CSCA). Paratypes: PANAMA: Panama: Chorrera, Playa Leona, cerca a río Perequete, 27. II. 1991, yellow pan trap, R. Cambra (allotype, 1 ♂ CSCA); Chorrera, Playa Leona, cerca a río Perequete: 6. II. 1991, yellow pan trap, R. Cambra (1 ♂ CSCA); 13. II. 1991, yellow pan trap, R. Cambra, (2 ♂ MIUP); 18 – 20. III. 1991, yellow pan trap, R. Cambra (2 ♂ MIUP, 2 ♂ DJBC); 26. II. 1991, yellow pan trap, R. Cambra (1 ♂ MIUP); 17. I. 1992, R. Cambra (1 ♀ CSCA, 1 ♀ 2 ♂ MIUP), 25. I. 1992, yellow pan trap, I. Henry (1 ♂ MIUP); 18. II. 1992, R. Cambra (1 ♀ DJBC); 18. II. 1992, A. Mena (2 ♀ MIUP); Chorrera, El Arado, 11. I. 1993, yellow pan trap, col. R. Urriola (1 ♂ MIUP); oficinas Parque Nacional Soberanía, 26. II. 2000, yellow pan trap, A. Santos (5 ♂ MIUP); Parque Metropolitano, 8. I. 1993, yellow pan trap, A. Velásquez (1 ♂ MIUP); near barriada Don Bosco, 13. IV. 2022, mangrove area, A. Santos (1 ♀ MIUP); Coclé: Tranquilla Norte, 12. III. 1988, yellow pan trap, R. Rodríguez (3 ♂ MIUP); Chiguirí Arriba, Cerro La Vieja, 23. IV. 1994, yellow pan trap, R. Cambra (1 ♂ MIUP).	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	etymology	Etymology. A new word based on the combined specific epithets of the type species of Dasymutilla Ashmead and Ephuta Say, 1836: Mutilla (Sphaeropthalma) gorgon Blake, 1871 and Mutilla (Ephuta) scrupea Say, 1836. Treat as a noun in apposition.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB9242243FF38FF63FD0BCC06.taxon	discussion	Remarks. The sexual association is based on morphological similarities of the mandible (bidentate apically, subapical tooth minute, lacking ventral tooth or notch), T 1 (petiolate), similar body size, sculpture, and setation. Additionally, males and females have a sympatric distribution, captured at the same time in some places. Two other species of Dasymutillini that are present in the same area (Dasymutilla araneoides (Smith, 1862) and Dasymutilla spilota Manley & Pitts, 2007) already have associated males and females (Cambra et al. 2018). All the above is sufficient evidence to consider males and females of Dasyphuta gorgupea as members of a single species. In both sexes, the coloration and body size are similar to many Panamanian species of Ephuta and Pseudomethoca Ashmead, 1896 (KAW & RAC, pers. obs.); females are predominantly reddish brown with a golden head and males are mostly black and covered with silvery setae.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
03C687DDB93D2241FF38FD70FA25CBC9.taxon	discussion	The genera Lomachaeta, Neomutilla, and Protophotopsis were excluded from Dasymutillini (Waldren et al. 2023) and are not keyed here. The key is written in a way that both sexes of velvet ants from the Americas can be checked for exclusion from Dasymutillini or recognized as one of the 13 genera of Dasymutillini.	en	Williams, Kevin A., Cambra, Roberto. A., Bartholomay, Pedro R., Lopez, Vinicius M., Brothers, Denis J., Martins, Herbeson O. J., Ferreira, Vinina S., Koroiva, Ricardo, Guillermo-Ferreira, Rhainer (2024): Two new genera of Neotropical Dasymutillini (Hymenoptera, Mutillidae, Sphaeropthalminae): Goncharovtilla gen. nov. from Brazil and Dasyphuta gen. nov. from Panama. Zootaxa 5538 (2): 127-152, DOI: 10.11646/zootaxa.5538.2.2, URL: https://doi.org/10.11646/zootaxa.5538.2.2
