taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745577/files/figure.png	https://doi.org/10.5281/zenodo.16745577	Fig. 1. Clinobiantes paradoxus Roewer, 1927, lectotype, ♂ (SMF 9900064), habitus photos. A. Dorsal view. B. Ventral view. C. Ventral view with detail of coxae I–III. D. Ventral view with detail of coxa IV and free sternites. E. Lateral view. F. Posterior view. Scale bars: A–B = 1 mm; C–D, F = 500 µm; E = 2 mm.	Fig. 1. Clinobiantes paradoxus Roewer, 1927, lectotype, ♂ (SMF 9900064), habitus photos. A. Dorsal view. B. Ventral view. C. Ventral view with detail of coxae I–III. D. Ventral view with detail of coxa IV and free sternites. E. Lateral view. F. Posterior view. Scale bars: A–B = 1 mm; C–D, F = 500 µm; E = 2 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745581/files/figure.png	https://doi.org/10.5281/zenodo.16745581	Fig. 3. Clinobiantes paradoxus Roewer, 1927, lectotype, ♂ (SMF 9900064), drawings of habitus, pedipalp, and chelicera. A. Habitus, dorsal view. B–C. Right pedipalp. B. Ectal view. C. Mesal view, D–E. Left chelicera. D. Ectal view. E. Frontal view. Scale bars = 500 µm.	Fig. 3. Clinobiantes paradoxus Roewer, 1927, lectotype, ♂ (SMF 9900064), drawings of habitus, pedipalp, and chelicera. A. Habitus, dorsal view. B–C. Right pedipalp. B. Ectal view. C. Mesal view, D–E. Left chelicera. D. Ectal view. E. Frontal view. Scale bars = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745583/files/figure.png	https://doi.org/10.5281/zenodo.16745583	Fig. 4. Clinobiantes paradoxus Roewer, 1927, lectotype, ♂ (SMF 9900064), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 4. Clinobiantes paradoxus Roewer, 1927, lectotype, ♂ (SMF 9900064), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745586/files/figure.png	https://doi.org/10.5281/zenodo.16745586	Fig. 5. Clinobiantes paradoxus Roewer, 1927, paralectotype, ♀ (SMF 9900064). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A = 1 mm; B = 2 mm; C = 500 µm; D–F = 100 µm.	Fig. 5. Clinobiantes paradoxus Roewer, 1927, paralectotype, ♀ (SMF 9900064). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A = 1 mm; B = 2 mm; C = 500 µm; D–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745588/files/figure.png	https://doi.org/10.5281/zenodo.16745588	Fig. 6. Clinobiantes paradoxus Roewer, 1927, sexual dimorphism, right legs II photos. A, C, E. Paralectotype, ♀ (SMF 9900064). B, D, F. Lectotype, ♂ (SMF 9900064). A–B. Femur, retrolateral view. C–D. Tibia, retrolateral view. E–F. Metatarsus, retrolateral view. Arrow indicates a pseudoarticular ring. Scale bars = 1 mm.	Fig. 6. Clinobiantes paradoxus Roewer, 1927, sexual dimorphism, right legs II photos. A, C, E. Paralectotype, ♀ (SMF 9900064). B, D, F. Lectotype, ♂ (SMF 9900064). A–B. Femur, retrolateral view. C–D. Tibia, retrolateral view. E–F. Metatarsus, retrolateral view. Arrow indicates a pseudoarticular ring. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745579/files/figure.png	https://doi.org/10.5281/zenodo.16745579	Fig. 2. Label accompanying the types Clinobiantes paradoxus Roewer, 1927 (SMF 9900064). Label text: Arachn. Coll. Roewer – Lfd. No.64, Opiliones: Phalangodid, Biantinae No.1, Clinobiantes paradoxus Rwr. 1♂, 2♀, Kamerun: Bibundi, Typus, Roewer det. 1927.	Fig. 2. Label accompanying the types Clinobiantes paradoxus Roewer, 1927 (SMF 9900064). Label text: Arachn. Coll. Roewer – Lfd. No.64, Opiliones: Phalangodid, Biantinae No.1, Clinobiantes paradoxus Rwr. 1♂, 2♀, Kamerun: Bibundi, Typus, Roewer det. 1927.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745681/files/figure.png	https://doi.org/10.5281/zenodo.16745681	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745590/files/figure.png	https://doi.org/10.5281/zenodo.16745590	Fig. 7. Metabiantes elongatus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.219865), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 7. Metabiantes elongatus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.219865), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745592/files/figure.png	https://doi.org/10.5281/zenodo.16745592	Fig. 8. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45430), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 200 µm.	Fig. 8. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45430), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 200 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745594/files/figure.png	https://doi.org/10.5281/zenodo.16745594	Fig. 9. Metabiantes elongatus sp. nov., scanning electron micrographs of pedipalp and chelicera. A–F. Paratype, ♂ (MACN-Ar 45425), left pedipalp. A. Mesal view. B. Femur, ventral view. C. Tibia and tarsus, ventral view. D. Detail of femur, ventral view. E. Detail of proximal spines on tibia. F. Detail of microtrichia. G–H. Paratype, ♂ (MACN-Ar 45430), left chelicera. G. Mesal view. H. Frontal view. Scale bars: A = 200 µm; B–C, G–H = 100 µm; D–E = 20 µm; F = 1 µm.	Fig. 9. Metabiantes elongatus sp. nov., scanning electron micrographs of pedipalp and chelicera. A–F. Paratype, ♂ (MACN-Ar 45425), left pedipalp. A. Mesal view. B. Femur, ventral view. C. Tibia and tarsus, ventral view. D. Detail of femur, ventral view. E. Detail of proximal spines on tibia. F. Detail of microtrichia. G–H. Paratype, ♂ (MACN-Ar 45430), left chelicera. G. Mesal view. H. Frontal view. Scale bars: A = 200 µm; B–C, G–H = 100 µm; D–E = 20 µm; F = 1 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745598/files/figure.png	https://doi.org/10.5281/zenodo.16745598	Fig. 10. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45425), scanning electron micrographs of left leg II. A. Femur, prolateral view. B. Detail of femur surface. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Distal metatarsus with division between astragalus and calcaneus visible. G. Detail of astragalus surface. H. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A, C, E = 200 µm; B, G = 20 µm; D, H = 10 µm; F = 30 µm.	Fig. 10. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45425), scanning electron micrographs of left leg II. A. Femur, prolateral view. B. Detail of femur surface. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Distal metatarsus with division between astragalus and calcaneus visible. G. Detail of astragalus surface. H. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A, C, E = 200 µm; B, G = 20 µm; D, H = 10 µm; F = 30 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745600/files/figure.png	https://doi.org/10.5281/zenodo.16745600	Fig. 11. Metabiantes elongatus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.219865), penis drawings. A–B. Dorsal view (detail of conductors within the box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 11. Metabiantes elongatus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.219865), penis drawings. A–B. Dorsal view (detail of conductors within the box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745604/files/figure.png	https://doi.org/10.5281/zenodo.16745604	Fig. 12. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45430), scanning electron micrographs of penis. A. Dorsal view. B. Lateral view. C. Ventral view. D. Detail of conductors, dorsal view. E. Detail of conductors, ventral view. F. Anterior view. G. Details of conductors, anterior view. Conductors in red, titillators in magenta, microsetae in yellow. Abbreviations: DF = dorsal fold; VF = ventral fold. Scale bars: A = 30 µm; B = 40 µm; C = 50 µm; D = 5 µm; E, G = 10 µm; F = 20 µm.	Fig. 12. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45430), scanning electron micrographs of penis. A. Dorsal view. B. Lateral view. C. Ventral view. D. Detail of conductors, dorsal view. E. Detail of conductors, ventral view. F. Anterior view. G. Details of conductors, anterior view. Conductors in red, titillators in magenta, microsetae in yellow. Abbreviations: DF = dorsal fold; VF = ventral fold. Scale bars: A = 30 µm; B = 40 µm; C = 50 µm; D = 5 µm; E, G = 10 µm; F = 20 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745606/files/figure.png	https://doi.org/10.5281/zenodo.16745606	Fig. 13. Metabiantes elongatus sp. nov., paratype, ♀ (MACN-Ar 45440). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A–C = 500 µm; D–F = 100 µm.	Fig. 13. Metabiantes elongatus sp. nov., paratype, ♀ (MACN-Ar 45440). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A–C = 500 µm; D–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745608/files/figure.png	https://doi.org/10.5281/zenodo.16745608	Fig. 14. Metabiantes elongatus sp. nov., sexual dimorphism, left legs II photos. A, C, E, G. Paratype, ♀ (MACN-Ar 45440). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.219865). A–B. Leg II, retrolateral view. C–D. Femur, retrolateral view. E–F. Tibia, retrolateral view. G–H. Metatarsus, retrolateral view. Scale bars: A–B = 1 mm; C–H = 500 µm.	Fig. 14. Metabiantes elongatus sp. nov., sexual dimorphism, left legs II photos. A, C, E, G. Paratype, ♀ (MACN-Ar 45440). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.219865). A–B. Leg II, retrolateral view. C–D. Femur, retrolateral view. E–F. Tibia, retrolateral view. G–H. Metatarsus, retrolateral view. Scale bars: A–B = 1 mm; C–H = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745664/files/figure.png	https://doi.org/10.5281/zenodo.16745664	Fig. 33. Metabiantes serratus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.223705), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 33. Metabiantes serratus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.223705), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745666/files/figure.png	https://doi.org/10.5281/zenodo.16745666	Fig. 34. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 200 µm.	Fig. 34. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 200 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745673/files/figure.png	https://doi.org/10.5281/zenodo.16745673	Fig. 36. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Detail of calcaneus. G. Detail of astragalus. H. Detail of calcaneus surface. Black triangles indicate trichomes; white triangle indicates a sensillum chaeticum. Scale bars: A–C, E = 400 µm; D, F–G = 40 µm; H = 20 µm.	Fig. 36. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Detail of calcaneus. G. Detail of astragalus. H. Detail of calcaneus surface. Black triangles indicate trichomes; white triangle indicates a sensillum chaeticum. Scale bars: A–C, E = 400 µm; D, F–G = 40 µm; H = 20 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745679/files/figure.png	https://doi.org/10.5281/zenodo.16745679	Fig. 39. Metabiantes serratus sp. nov., sexual dimorphism, photos of left legs II. A, C, E, G. Paratype, ♀ (MACN-Ar 45474). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.223705). A–B. Leg II, retrolateral view. C–D. Detail of femur and patella, retrolateral view. E–F. Detail of tibia, retrolateral view. G–H. Detail of metatarsus, retrolateral view. Scale bars: A–B, D = 500 µm; C, E–H = 200 µm.	Fig. 39. Metabiantes serratus sp. nov., sexual dimorphism, photos of left legs II. A, C, E, G. Paratype, ♀ (MACN-Ar 45474). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.223705). A–B. Leg II, retrolateral view. C–D. Detail of femur and patella, retrolateral view. E–F. Detail of tibia, retrolateral view. G–H. Detail of metatarsus, retrolateral view. Scale bars: A–B, D = 500 µm; C, E–H = 200 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFE4FFA64421A41460D1F3D9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745681/files/figure.png	https://doi.org/10.5281/zenodo.16745681	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745610/files/figure.png	https://doi.org/10.5281/zenodo.16745610	Fig. 15. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), habitus photos. A. Dorsal view. B. Ventral view. C. Ventral view with detail of coxae I–IV. D. Ventral view with detail of coxa IV and free sternites. E. Lateral view. F. Posterior view. Scale bars: A–B, E = 1 mm; C = 200 µm; D, F = 500 µm.	Fig. 15. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), habitus photos. A. Dorsal view. B. Ventral view. C. Ventral view with detail of coxae I–IV. D. Ventral view with detail of coxa IV and free sternites. E. Lateral view. F. Posterior view. Scale bars: A–B, E = 1 mm; C = 200 µm; D, F = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745612/files/figure.png	https://doi.org/10.5281/zenodo.16745612	Fig. 16. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473). A–C, F–H. Drawings of habitus, pedipalp, and chelicera. D–E. Photos of pedipalp femur. A. Habitus, dorsal view. B–F. Left pedipalp. B. Ectal view. C. Mesal view. D. Femur, mesal view. E. Detail of femur surface. F. Femur, ventral view. G–H. Left chelicera. G. Ectal view. H. Frontal view. White triangle indicates a granule. Scale bars: A–C, F–H = 500 µm; D = 250 µm; E = 200 µm.	Fig. 16. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473). A–C, F–H. Drawings of habitus, pedipalp, and chelicera. D–E. Photos of pedipalp femur. A. Habitus, dorsal view. B–F. Left pedipalp. B. Ectal view. C. Mesal view. D. Femur, mesal view. E. Detail of femur surface. F. Femur, ventral view. G–H. Left chelicera. G. Ectal view. H. Frontal view. White triangle indicates a granule. Scale bars: A–C, F–H = 500 µm; D = 250 µm; E = 200 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745614/files/figure.png	https://doi.org/10.5281/zenodo.16745614	Fig. 17. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), scanning electron micrographs of right leg II. A. Leg II, retrolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, ventroprolateral view. F. Detail of astragalus and calcaneus surfaces. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum; black arrow indicates a glandular opening. Scale bars: A = 300 µm; B = 200 µm; C, E = 100 µm; D = 40 µm; F = 10 µm.	Fig. 17. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), scanning electron micrographs of right leg II. A. Leg II, retrolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, ventroprolateral view. F. Detail of astragalus and calcaneus surfaces. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum; black arrow indicates a glandular opening. Scale bars: A = 300 µm; B = 200 µm; C, E = 100 µm; D = 40 µm; F = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745616/files/figure.png	https://doi.org/10.5281/zenodo.16745616	Fig. 18. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 18. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745620/files/figure.png	https://doi.org/10.5281/zenodo.16745620	Fig. 19. Metabiantes herculeus sp. nov., paratype, ♀ (BE_RMCA_ARA.Opi.247663). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A, C = 500 µm; B = 1 mm; D–F = 100 µm.	Fig. 19. Metabiantes herculeus sp. nov., paratype, ♀ (BE_RMCA_ARA.Opi.247663). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A, C = 500 µm; B = 1 mm; D–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745622/files/figure.png	https://doi.org/10.5281/zenodo.16745622	Fig. 20. Metabiantes herculeus sp. nov., sexual dimorphism, left legs II photos. A, C, E, G. Paratype, ♀ (BE_RMCA_ARA.Opi.247663). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.173473). A–B. Leg II, retrolateral view. C–D. Femur, retrolateral view. E–F. Tibia, retrolateral view. G–H. Metatarsus, retrolateral view. Arrow indicates a pseudoarticular ring. Scale bars: A–B = 1 mm; C–H = 500 µm.	Fig. 20. Metabiantes herculeus sp. nov., sexual dimorphism, left legs II photos. A, C, E, G. Paratype, ♀ (BE_RMCA_ARA.Opi.247663). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.173473). A–B. Leg II, retrolateral view. C–D. Femur, retrolateral view. E–F. Tibia, retrolateral view. G–H. Metatarsus, retrolateral view. Arrow indicates a pseudoarticular ring. Scale bars: A–B = 1 mm; C–H = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745635/files/figure.png	https://doi.org/10.5281/zenodo.16745635	Fig. 23. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), photographs and scanning electron micrographs of left leg II. A. Femur, retrolateral view. B. Femur, ventral view. C–D. Tibia, retrolateral view. E–F. Metatarsus, retrolateral view. G. Metatarsus, ventral view. H. Detail of astragalus and calcaneus, retrolateral view. I. Detail of astragalus and calcaneus surface, ventral view. J. Detail of calcaneus surface, retrolateral view. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum.Abbreviations: As = astragalus; Ca = calcaneus. Scale bars: A, C, E = 500 µm; B, F = 200 µm; D, G = 100 µm; H = 40 µm; I = 30 µm; J = 5 µm.	Fig. 23. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), photographs and scanning electron micrographs of left leg II. A. Femur, retrolateral view. B. Femur, ventral view. C–D. Tibia, retrolateral view. E–F. Metatarsus, retrolateral view. G. Metatarsus, ventral view. H. Detail of astragalus and calcaneus, retrolateral view. I. Detail of astragalus and calcaneus surface, ventral view. J. Detail of calcaneus surface, retrolateral view. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum.Abbreviations: As = astragalus; Ca = calcaneus. Scale bars: A, C, E = 500 µm; B, F = 200 µm; D, G = 100 µm; H = 40 µm; I = 30 µm; J = 5 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745648/files/figure.png	https://doi.org/10.5281/zenodo.16745648	Fig. 28. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Detail of astragalus and calcaneus, prolateral view. F. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A = 400 µm; B = 200 µm; C = 100 µm; D = 20 µm; E = 40 µm; F = 10 µm.	Fig. 28. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Detail of astragalus and calcaneus, prolateral view. F. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A = 400 µm; B = 200 µm; C = 100 µm; D = 20 µm; E = 40 µm; F = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745637/files/figure.png	https://doi.org/10.5281/zenodo.16745637	Fig. 24. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 24. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745653/files/figure.png	https://doi.org/10.5281/zenodo.16745653	Fig. 29. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 29. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745639/files/figure.png	https://doi.org/10.5281/zenodo.16745639	Fig. 25. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 25. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745659/files/figure.png	https://doi.org/10.5281/zenodo.16745659	Fig. 31. Metabiantes kivuensis sp. nov., sexual dimorphism. A–F. Paratype, ♀ (BE_RMCA_ARA. Opi.247664). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. G–H. Paratype, minor ♂ (BE_RMCA_ARA. Opi.247664), photo habitus. G. Dorsal view. H. Lateral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr= truncus. Scale bars: A–B, G–H = 1 mm; C–F = 100 µm.	Fig. 31. Metabiantes kivuensis sp. nov., sexual dimorphism. A–F. Paratype, ♀ (BE_RMCA_ARA. Opi.247664). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. G–H. Paratype, minor ♂ (BE_RMCA_ARA. Opi.247664), photo habitus. G. Dorsal view. H. Lateral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr= truncus. Scale bars: A–B, G–H = 1 mm; C–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFFAFFA94421A47460D1F7B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745681/files/figure.png	https://doi.org/10.5281/zenodo.16745681	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745626/files/figure.png	https://doi.org/10.5281/zenodo.16745626	Fig. 21. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), habitus photos. A. Dorsal view. B. Ventral view. C. Ventral view with detail of coxae I–IV. D. Ventral view with detail of coxa IV and free sternites. E. Lateral view. F. Posterior view. Scale bars: A–B, E = 1 mm; C = 200 µm; D, F = 500 µm.	Fig. 21. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), habitus photos. A. Dorsal view. B. Ventral view. C. Ventral view with detail of coxae I–IV. D. Ventral view with detail of coxa IV and free sternites. E. Lateral view. F. Posterior view. Scale bars: A–B, E = 1 mm; C = 200 µm; D, F = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745632/files/figure.png	https://doi.org/10.5281/zenodo.16745632	Fig. 22. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), drawings of habitus, pedipalp, and chelicera. A. Habitus, dorsal view. B–D. Left pedipalp. B. Mesal view. C. Ectal view. D. Femur, ventral view. E–F. Left chelicera. E. Ectal view. F. Frontal view. Scale bars: A = 1 mm; B–D = 500 µm; E–F = 250 µm.	Fig. 22. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), drawings of habitus, pedipalp, and chelicera. A. Habitus, dorsal view. B–D. Left pedipalp. B. Mesal view. C. Ectal view. D. Femur, ventral view. E–F. Left chelicera. E. Ectal view. F. Frontal view. Scale bars: A = 1 mm; B–D = 500 µm; E–F = 250 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745635/files/figure.png	https://doi.org/10.5281/zenodo.16745635	Fig. 23. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), photographs and scanning electron micrographs of left leg II. A. Femur, retrolateral view. B. Femur, ventral view. C–D. Tibia, retrolateral view. E–F. Metatarsus, retrolateral view. G. Metatarsus, ventral view. H. Detail of astragalus and calcaneus, retrolateral view. I. Detail of astragalus and calcaneus surface, ventral view. J. Detail of calcaneus surface, retrolateral view. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum.Abbreviations: As = astragalus; Ca = calcaneus. Scale bars: A, C, E = 500 µm; B, F = 200 µm; D, G = 100 µm; H = 40 µm; I = 30 µm; J = 5 µm.	Fig. 23. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), photographs and scanning electron micrographs of left leg II. A. Femur, retrolateral view. B. Femur, ventral view. C–D. Tibia, retrolateral view. E–F. Metatarsus, retrolateral view. G. Metatarsus, ventral view. H. Detail of astragalus and calcaneus, retrolateral view. I. Detail of astragalus and calcaneus surface, ventral view. J. Detail of calcaneus surface, retrolateral view. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum.Abbreviations: As = astragalus; Ca = calcaneus. Scale bars: A, C, E = 500 µm; B, F = 200 µm; D, G = 100 µm; H = 40 µm; I = 30 µm; J = 5 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745637/files/figure.png	https://doi.org/10.5281/zenodo.16745637	Fig. 24. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 24. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745614/files/figure.png	https://doi.org/10.5281/zenodo.16745614	Fig. 17. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), scanning electron micrographs of right leg II. A. Leg II, retrolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, ventroprolateral view. F. Detail of astragalus and calcaneus surfaces. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum; black arrow indicates a glandular opening. Scale bars: A = 300 µm; B = 200 µm; C, E = 100 µm; D = 40 µm; F = 10 µm.	Fig. 17. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), scanning electron micrographs of right leg II. A. Leg II, retrolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, ventroprolateral view. F. Detail of astragalus and calcaneus surfaces. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum; black arrow indicates a glandular opening. Scale bars: A = 300 µm; B = 200 µm; C, E = 100 µm; D = 40 µm; F = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745616/files/figure.png	https://doi.org/10.5281/zenodo.16745616	Fig. 18. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 18. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745653/files/figure.png	https://doi.org/10.5281/zenodo.16745653	Fig. 29. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 29. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745639/files/figure.png	https://doi.org/10.5281/zenodo.16745639	Fig. 25. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 25. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745659/files/figure.png	https://doi.org/10.5281/zenodo.16745659	Fig. 31. Metabiantes kivuensis sp. nov., sexual dimorphism. A–F. Paratype, ♀ (BE_RMCA_ARA. Opi.247664). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. G–H. Paratype, minor ♂ (BE_RMCA_ARA. Opi.247664), photo habitus. G. Dorsal view. H. Lateral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr= truncus. Scale bars: A–B, G–H = 1 mm; C–F = 100 µm.	Fig. 31. Metabiantes kivuensis sp. nov., sexual dimorphism. A–F. Paratype, ♀ (BE_RMCA_ARA. Opi.247664). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. G–H. Paratype, minor ♂ (BE_RMCA_ARA. Opi.247664), photo habitus. G. Dorsal view. H. Lateral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr= truncus. Scale bars: A–B, G–H = 1 mm; C–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFF2FF924437A15960D1F43A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745681/files/figure.png	https://doi.org/10.5281/zenodo.16745681	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745639/files/figure.png	https://doi.org/10.5281/zenodo.16745639	Fig. 25. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 25. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745641/files/figure.png	https://doi.org/10.5281/zenodo.16745641	Fig. 26. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 300 µm.	Fig. 26. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 300 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745646/files/figure.png	https://doi.org/10.5281/zenodo.16745646	Fig. 27. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of pedipalp and chelicera. A–E. Right pedipalp. A. Mesal view. B. Detail of femur. C. Detail of femoral granules. D. Tibia and tarsus, ventroectal view. E. Detail of aggregate pores of tarsus. F–H. Left chelicera. F. Mesal view. G. Ectal view. H. Frontal view. Scale bars: A = 200 µm; B = 20 µm; C, E = 10 µm; D, F–H = 100 µm.	Fig. 27. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of pedipalp and chelicera. A–E. Right pedipalp. A. Mesal view. B. Detail of femur. C. Detail of femoral granules. D. Tibia and tarsus, ventroectal view. E. Detail of aggregate pores of tarsus. F–H. Left chelicera. F. Mesal view. G. Ectal view. H. Frontal view. Scale bars: A = 200 µm; B = 20 µm; C, E = 10 µm; D, F–H = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745648/files/figure.png	https://doi.org/10.5281/zenodo.16745648	Fig. 28. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Detail of astragalus and calcaneus, prolateral view. F. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A = 400 µm; B = 200 µm; C = 100 µm; D = 20 µm; E = 40 µm; F = 10 µm.	Fig. 28. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Detail of astragalus and calcaneus, prolateral view. F. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A = 400 µm; B = 200 µm; C = 100 µm; D = 20 µm; E = 40 µm; F = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745653/files/figure.png	https://doi.org/10.5281/zenodo.16745653	Fig. 29. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 29. Metabiantes kivuensis sp. nov., holotype, major ♂ (BE_RMCA_ARA.Opi. 154162), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745657/files/figure.png	https://doi.org/10.5281/zenodo.16745657	Fig. 30. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of penis. A. Dorsal view. B. Lateral view. C. Ventral view. D. Detail of conductors, dorsal view. E. Anterior view. F. Detail of conductors, anterior view. Conductors in red, titillators in magenta, microsetae in yellow. Abbreviations: DF = dorsal fold; VF = ventral fold. Scale bars: A–C = 40 µm; D, F = 10 µm; E = 20 µm.	Fig. 30. Metabiantes kivuensis sp. nov., paratype, minor ♂ (MACN-Ar 45445), scanning electron micrographs of penis. A. Dorsal view. B. Lateral view. C. Ventral view. D. Detail of conductors, dorsal view. E. Anterior view. F. Detail of conductors, anterior view. Conductors in red, titillators in magenta, microsetae in yellow. Abbreviations: DF = dorsal fold; VF = ventral fold. Scale bars: A–C = 40 µm; D, F = 10 µm; E = 20 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745659/files/figure.png	https://doi.org/10.5281/zenodo.16745659	Fig. 31. Metabiantes kivuensis sp. nov., sexual dimorphism. A–F. Paratype, ♀ (BE_RMCA_ARA. Opi.247664). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. G–H. Paratype, minor ♂ (BE_RMCA_ARA. Opi.247664), photo habitus. G. Dorsal view. H. Lateral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr= truncus. Scale bars: A–B, G–H = 1 mm; C–F = 100 µm.	Fig. 31. Metabiantes kivuensis sp. nov., sexual dimorphism. A–F. Paratype, ♀ (BE_RMCA_ARA. Opi.247664). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. G–H. Paratype, minor ♂ (BE_RMCA_ARA. Opi.247664), photo habitus. G. Dorsal view. H. Lateral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; P = projection; S = seta; SrA = seminal receptacle area; Tr= truncus. Scale bars: A–B, G–H = 1 mm; C–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745662/files/figure.png	https://doi.org/10.5281/zenodo.16745662	Fig. 32. Metabiantes kivuensis sp. nov., sexual dimorphism, left legs II photos. A, D, G, J. Paratype, ♀ (BE_RMCA_ARA.Opi.247664). B, E, H, K. Paratype, minor ♂ (BE_RMCA_ARA.Opi.247664). C, F, I, L. Holotype, major ♂ (BE_RMCA_ARA.Opi.154162). A–C. Leg II, retrolateral view. D–F. Femur and patella, retrolateral view. G–I. Tibia, retrolateral view. J–L. Metatarsus and tarsus, retrolateral view. Scale bars = 500 µm.	Fig. 32. Metabiantes kivuensis sp. nov., sexual dimorphism, left legs II photos. A, D, G, J. Paratype, ♀ (BE_RMCA_ARA.Opi.247664). B, E, H, K. Paratype, minor ♂ (BE_RMCA_ARA.Opi.247664). C, F, I, L. Holotype, major ♂ (BE_RMCA_ARA.Opi.154162). A–C. Leg II, retrolateral view. D–F. Femur and patella, retrolateral view. G–I. Tibia, retrolateral view. J–L. Metatarsus and tarsus, retrolateral view. Scale bars = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745614/files/figure.png	https://doi.org/10.5281/zenodo.16745614	Fig. 17. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), scanning electron micrographs of right leg II. A. Leg II, retrolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, ventroprolateral view. F. Detail of astragalus and calcaneus surfaces. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum; black arrow indicates a glandular opening. Scale bars: A = 300 µm; B = 200 µm; C, E = 100 µm; D = 40 µm; F = 10 µm.	Fig. 17. Metabiantes herculeus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.173473), scanning electron micrographs of right leg II. A. Leg II, retrolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, ventroprolateral view. F. Detail of astragalus and calcaneus surfaces. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum; black arrow indicates a glandular opening. Scale bars: A = 300 µm; B = 200 µm; C, E = 100 µm; D = 40 µm; F = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745637/files/figure.png	https://doi.org/10.5281/zenodo.16745637	Fig. 24. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator. Scale bars: A = 500 µm; B–D = 100 µm.	Fig. 24. Metabiantes kaurii sp. nov., holotype, ♂ (MACN-Ar 46476), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. Stylus in green, conductors in brownish red, titillators in magenta. Abbreviations: Co = conductor; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator. Scale bars: A = 500 µm; B–D = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFCFFF994422A15960D1F37F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745681/files/figure.png	https://doi.org/10.5281/zenodo.16745681	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745664/files/figure.png	https://doi.org/10.5281/zenodo.16745664	Fig. 33. Metabiantes serratus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.223705), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 33. Metabiantes serratus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.223705), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745666/files/figure.png	https://doi.org/10.5281/zenodo.16745666	Fig. 34. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 200 µm.	Fig. 34. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 200 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745670/files/figure.png	https://doi.org/10.5281/zenodo.16745670	Fig. 35. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of pedipalp and chelicera. A–E. Left pedipalp. A. Mesal view. B. Trochanter and femur, mesal view. C. Tibia, ventromesal view. D. Detail of ectoproximal tibial spine. E. Detail of femur surface. F–H. Right chelicera. F. Ectal view. G. Mesal view. H. Frontal view. Scale bars: A = 200 µm; B–C, F–H = 100 µm; D–E = 10 µm.	Fig. 35. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of pedipalp and chelicera. A–E. Left pedipalp. A. Mesal view. B. Trochanter and femur, mesal view. C. Tibia, ventromesal view. D. Detail of ectoproximal tibial spine. E. Detail of femur surface. F–H. Right chelicera. F. Ectal view. G. Mesal view. H. Frontal view. Scale bars: A = 200 µm; B–C, F–H = 100 µm; D–E = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745673/files/figure.png	https://doi.org/10.5281/zenodo.16745673	Fig. 36. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Detail of calcaneus. G. Detail of astragalus. H. Detail of calcaneus surface. Black triangles indicate trichomes; white triangle indicates a sensillum chaeticum. Scale bars: A–C, E = 400 µm; D, F–G = 40 µm; H = 20 µm.	Fig. 36. Metabiantes serratus sp. nov., paratype, ♂ (MACN-Ar 45472), scanning electron micrographs of left leg II. A. Leg II, prolateral view. B. Femur, prolateral view. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Detail of calcaneus. G. Detail of astragalus. H. Detail of calcaneus surface. Black triangles indicate trichomes; white triangle indicates a sensillum chaeticum. Scale bars: A–C, E = 400 µm; D, F–G = 40 µm; H = 20 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745675/files/figure.png	https://doi.org/10.5281/zenodo.16745675	Fig. 37. Metabiantes serratus sp. nov., males, drawings and scanning electron micrographs of penis. A–E. Holotype (BE_RMCA_ARA.Opi.223705), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. E. Everted penis, dorsal view. F–J. Paratype (MACN- Ar 45474), scanning electron micrographs of penis. F. Dorsal view. G. Lateral view. H. Ventral view. I. Anterior view. J. Detail of conductors, anterior view. Stylus in green, conductors in brownish red, titillators in magenta, microsetae in yellow. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–E = 100 µm; F–G = 20 µm; H = 30 µm; I–J = 10 µm.	Fig. 37. Metabiantes serratus sp. nov., males, drawings and scanning electron micrographs of penis. A–E. Holotype (BE_RMCA_ARA.Opi.223705), penis drawings. A–B. Dorsal view (detail of conductors within box). C. Lateral view. D. Ventral view. E. Everted penis, dorsal view. F–J. Paratype (MACN- Ar 45474), scanning electron micrographs of penis. F. Dorsal view. G. Lateral view. H. Ventral view. I. Anterior view. J. Detail of conductors, anterior view. Stylus in green, conductors in brownish red, titillators in magenta, microsetae in yellow. Abbreviations: Co = conductor; DF = dorsal fold; LP = lateral projection; PB = pars basalis; PD = pars distalis; St = stylus; Ti = titillator; VF = ventral fold. Scale bars: A = 500 µm; B–E = 100 µm; F–G = 20 µm; H = 30 µm; I–J = 10 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745677/files/figure.png	https://doi.org/10.5281/zenodo.16745677	Fig. 38. Metabiantes serratus sp. nov., paratype, ♀ (MACN-Ar 45474). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A–B = 1 mm; C–F = 100 µm.	Fig. 38. Metabiantes serratus sp. nov., paratype, ♀ (MACN-Ar 45474). A–B. Habitus photos. A. Dorsal view. B. Lateral view. C–F. Ovipositor drawings. C–D. Dorsal view. E. Lateral view. F. Ventral view. Abbreviations: FG = furcal groove; Fu = furca; L = lobe; S = seta; SrA = seminal receptacle area; Tr = truncus. Scale bars: A–B = 1 mm; C–F = 100 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745679/files/figure.png	https://doi.org/10.5281/zenodo.16745679	Fig. 39. Metabiantes serratus sp. nov., sexual dimorphism, photos of left legs II. A, C, E, G. Paratype, ♀ (MACN-Ar 45474). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.223705). A–B. Leg II, retrolateral view. C–D. Detail of femur and patella, retrolateral view. E–F. Detail of tibia, retrolateral view. G–H. Detail of metatarsus, retrolateral view. Scale bars: A–B, D = 500 µm; C, E–H = 200 µm.	Fig. 39. Metabiantes serratus sp. nov., sexual dimorphism, photos of left legs II. A, C, E, G. Paratype, ♀ (MACN-Ar 45474). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.223705). A–B. Leg II, retrolateral view. C–D. Detail of femur and patella, retrolateral view. E–F. Detail of tibia, retrolateral view. G–H. Detail of metatarsus, retrolateral view. Scale bars: A–B, D = 500 µm; C, E–H = 200 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745590/files/figure.png	https://doi.org/10.5281/zenodo.16745590	Fig. 7. Metabiantes elongatus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.219865), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	Fig. 7. Metabiantes elongatus sp. nov., holotype, ♂ (BE_RMCA_ARA.Opi.219865), habitus photos. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745598/files/figure.png	https://doi.org/10.5281/zenodo.16745598	Fig. 10. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45425), scanning electron micrographs of left leg II. A. Femur, prolateral view. B. Detail of femur surface. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Distal metatarsus with division between astragalus and calcaneus visible. G. Detail of astragalus surface. H. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A, C, E = 200 µm; B, G = 20 µm; D, H = 10 µm; F = 30 µm.	Fig. 10. Metabiantes elongatus sp. nov., paratype, ♂ (MACN-Ar 45425), scanning electron micrographs of left leg II. A. Femur, prolateral view. B. Detail of femur surface. C. Tibia, prolateral view. D. Detail of tibia surface. E. Metatarsus, prolateral view. F. Distal metatarsus with division between astragalus and calcaneus visible. G. Detail of astragalus surface. H. Detail of calcaneus surface. Black triangle indicates a trichome; white triangle indicates a sensillum chaeticum. Scale bars: A, C, E = 200 µm; B, G = 20 µm; D, H = 10 µm; F = 30 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745608/files/figure.png	https://doi.org/10.5281/zenodo.16745608	Fig. 14. Metabiantes elongatus sp. nov., sexual dimorphism, left legs II photos. A, C, E, G. Paratype, ♀ (MACN-Ar 45440). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.219865). A–B. Leg II, retrolateral view. C–D. Femur, retrolateral view. E–F. Tibia, retrolateral view. G–H. Metatarsus, retrolateral view. Scale bars: A–B = 1 mm; C–H = 500 µm.	Fig. 14. Metabiantes elongatus sp. nov., sexual dimorphism, left legs II photos. A, C, E, G. Paratype, ♀ (MACN-Ar 45440). B, D, F, H. Holotype, ♂ (BE_RMCA_ARA.Opi.219865). A–B. Leg II, retrolateral view. C–D. Femur, retrolateral view. E–F. Tibia, retrolateral view. G–H. Metatarsus, retrolateral view. Scale bars: A–B = 1 mm; C–H = 500 µm.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16745681/files/figure.png	https://doi.org/10.5281/zenodo.16745681	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	Fig. 40. Geographical distribution of Clinobiantes paradoxus Roewer, 1927 and the new species of Metabiantes Roewer, 1915 herein described. Clinobiantes paradoxus (red hexagon) in Cameroon; M. kivuensis sp. nov. (green triangle), M. serratus sp. nov. (magenta circle), and M. elongatus sp. nov. (yellow inverted triangle) in Democratic Republic of Congo; M. herculeus sp. nov. (pink square) in Tanzania; M. kaurii sp. nov. (orange star) in Mozambique.	2025-08-01	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel		Zenodo	biologists	Mamani, Vanesa;Jocqué, Merlijn;Pérez-González, Abel			
