taxonID	type	description	language	source
03C2879BFFEDFFB1440EA42563F1F6A3.taxon	description	Includes species Monotypic.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFB1440EA42563F1F6A3.taxon	distribution	Distribution Cameroon (Southwest Province), Equatorial Guinea (Corisco Island).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFB1440EA42563F1F6A3.taxon	discussion	Remarks Santos & Prieto (2009) recorded a Clinobiantes sp. for Equatorial Guinea, but the identity of this species has not been clarified yet.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	description	Figs 1, 3 – 6; Table 1	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	materials_examined	Type material Lectotype here designated CAMEROON • ♂; Bibundi; SMF 9900064 (examined) (Fig. 2). Paralectotype CAMEROON • ♀; same data as for lectotype; SMF 9900064 (examined) (Fig. 2).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	discussion	Remarks A second female is morphologically different and not conspecific with the male lectotype of Clinobiantes paradoxus or the female paralectotype herein designated (Fig. 2).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	description	Redescription Male (lectotype, SMF 9900064) BODY MEASUREMENTS. Total body length 2.48, carapace length 0.76, scutum magnum length 2.17, carapace maximum width 1.36, abdominal scutum maximum width 1.8. Appendage measurements in Table 1. DORSUM. Outline slightly hourglass-shaped (almost rectangular) with Eta (η) shape, with a very slight constriction at sulcus I level (Figs 1 A, 3 A). Carapace granulated, wider than long, anterior border slightly convex and unarmed (Figs 1 A, 3 A). Cheliceral sockets not marked (Figs 1 A, 3 A). Eyes separated near sulcus I (Figs 1 A, 3 A); interocular area granulated (Figs 1 E, 3 A). Carapace in lateral view straight at anterior region and slightly higher posteriorly (Fig. 1 E). Abdominal scutum in lateral view convex (Fig. 1 E). Sulcus I deep and well-marked, in dorsal view curved to the anterior body region (Figs 1 A, E, 3 A). Mesotergal areas coarsely granulated and well-defined, with sulci II – V marked but shallower than sulcus I; medially sulci II – III slightly curved to the anterior body region; sulci IV – V straight (Figs 1 A, E, 3 A). Mesotergal areas III – IV medially with two conical and pointed tubercles, strongly inclined backward (Figs 1 A, E – F, 3 A). Mesotergal area V granulated, medially with three conical and pointed tubercles. Lateral borders of abdominal scutum with rows of granules (Figs 1 A, E, 3 A). Free tergites granulated; free tergite I medially with three small conical tubercles (Figs 1 A, E, 3 A). VENTER. Coxa I with some setiferous tubercles in anterior margin (Fig. 1 C); coxa II incrassated, of same size as (or slightly larger than) coxa IV (Fig. 1 B – D); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Fig. 1 B – D). Posterior border of spiracular area and free sternites I – V with a row of granules; anal operculum with few granules (Fig. 1 D, F). Spiracles not concealed (Fig. 1 B, D). CHELICERA. Basichelicerite unarmed, with an elongated and slightly marked bulla (Figs 1 E, 3 D). Cheliceral hand with sparse setae (Fig. 3 D – E). Fixed and movable fingers with small triangular-shaped teeth (Fig. 3 E). PEDIPALP. Coxa elongated (i. e., remarkably longer than trochanter), proximally with one dorsoectal and one ectoventral granule (Figs 1 A, C, 3 A). Femur straight, proximally with a slight ventral narrowing followed by a small ventral spine (Figs 1 E, 3 B – C). Patella elongated, club-shaped, with a distal ventromesal spine (Figs 1 E, 3 B – C). Tibia with two ventromesal and two ventroectal long spines (Fig. 3 B – C). Tarsus with two ventromesal and two ventroectal long spines (Figs 1 E, 3 B – C). LEGS. Femur II fusiform dorsally swollen (Figs 1 A, B, 6 B). Tibia II slender, slightly distally swollen, with ventral small triangular tubercles (Fig. 6 D). Metatarsus II with a ventrally swollen astragalus bearing two rows of triangular tubercles; astragalus-calcaneus boundary marked by a strong constriction, and slightly thickened calcaneus giving a peculiar form to the distal part of the metatarsus (Fig. 6 F). Tarsi III – IV with a dense scopula. Tarsal formula: 3 (2): 5 (3): 5: 5. COLOR (in 80 % ethanol). Body and appendages uniformly yellowish-brown (Figs 1 A – F, 6 B, D, F). MALE GENITALIA. Penis with clearly defined boundaries between pars basalis and pars distalis (Fig. 4 A). Pars basalis tubular, basally thin, broadens medially, with distal constriction (Fig. 4 A). Pars distalis swollen, with maximum width at titillator level (Fig. 4 A – B, D). Apical edge laminar (i. e., dorsoventrally flat) with a medial U-shaped cleft dividing it into two rounded halves (Fig. 4 B, D); halves less chitinous apically, and could possibly be inflated by hemolymph pressure (Fig. 4 C – D). Pars distalis with a distal depression in the ventromedial region (Fig. 4 D). Each side of pars distalis armed with short, conical microsetae arranged irregularly, extending from the dorsolateral to the ventrodistal region (Fig. 4 B – D). Capsula externa with two broad titillators covering most capsula interna (Fig. 4 B – C). Capsula interna with two complex conductors and one stylus, basally fused. Each conductor with two medial laminar folds apically, one short dorsal and one ventral longer, visible ventrally within the U-shaped cleft; each conductor also with one broad lateral projection (Fig. 4 B); stylus tubular, with its free tip fully covered by conductors (Fig. 4 B – D). Female (paralectotype, SMF 9900064) BODY MEASUREMENTS. Total body length 2.82, carapace length 0.76, scutum magnum length 2.31, maximum carapace width 1.38, abdominal scutum maximum width 1.88. Appendage measurements in Table 1. BODY. Female resembles male in armature of scutum magnum, but with tubercles of mesotergal areas III – IV, posterior border, and free tergite I slightly smaller (Fig. 5 A – B vs Fig. 1 E). Leg II with femur, tibia, and metatarsus neither swollen nor armed (Fig. 6 A, C, E vs Fig. 6 B, D, F); metatarsus II thin and unarmed, with pseudoarticular rings (Fig. 6 E vs Fig. 6 F). Tarsal formula 3 (2): 5 (3): 5: 5. FEMALE GENITALIA. Ovipositor cylindrical (Fig. 5 C), distally bearing two lobes (furca) (Fig. 5 C – D, F). Each furcal lobe with five long, pointed setae (Fig. 5 E) – three dorsal and two ventral – resulting in a total of six setae on the dorsal region (Fig. 5 D) and four on the ventral region (Fig. 5 F). External surface of furcal lobes with several short, pointed projections, irregularly distributed (Fig. 5 E). Receptacle chambers located near the base of the furcal groove (Fig. 5 D – F).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFEDFFBB4475A6FA60D1F5D0.taxon	distribution	Distribution Known only from the type locality (Fig. 40).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE7FFB84410A64F6735F439.taxon	description	Included species Metabiantes armatus Lawrence, 1962, M. barbertonensis Lawrence, 1963, M. basutoanus Kaury, 1961, M. caracticus Kaury, 1961, M. convexus Roewer, 1949, M. elongatus sp. nov., M. filipes (Roewer, 1912), M. flavus Lawrence, 1949, M. hanstroemi Kauri, 1961, M. herculeus sp. nov., M. incertus Kauri, 1961, M. insulanus (Roewer, 1949), M. jeanneli (Roewer, 1913), M. kakololius Kauri, 1985, M. kaurii sp. nov., M. kivuensis sp. nov., M. kosibaiensis Kauri, 1961, M. lawrencei Starega, 1992, M. leighi (Pocock, 1902), M. litoralis Kauri, 1961, M. longipes (Kauri, 1985), M. machadoi Lawrence, 1957, M. meraculus (Loman, 1898), M. minutus Kauri, 1985, M. montanus Kauri, 1985, M. obscurus Kauri, 1961, M. parvulus Kauri, 1985, M. perustus Lawrence, 1963, M. pumilio Roewer, 1927, M. punctatus (Sørensen, 1910), M. pusulosus (Loman, 1898), M. rudebecki Kauri, 1961, M. serratus sp. nov., M. stanleyi Kauri, 1985, M. submontanus Kauri, 1985, M. teres Lawrence, 1963, M. teretipes Lawrence, 1962, M. traegardhi Kauri, 1961, M. trifasciatus Roewer, 1915, M. ulindinus Kauri, 1985, M. unicolor (Roewer, 1912), M. urbanus Kauri, 1961, M. varius Kauri, 1961, M. zuluanus Lawrence, 1937, and M. zuurbergianus Kauri, 1961.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE7FFB84410A64F6735F439.taxon	distribution	Distribution Afrotropical: Angola, Cameroon, Democratic Republic of the Congo, Equatorial Guinea *, Kenya, Lesotho, Mozambique, São Tomé and Príncipe, South Africa, Tanzania, and Uganda.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE7FFB84410A64F6735F439.taxon	discussion	Remarks * Santos & Prieto (2009) recorded Metabiantes spp. for Equatorial Guinea (Muni River, Motora River (Mitong), Congüe River, and Etembue), but the identity of this species has not been clarified yet.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE4FFA64421A41460D1F3D9.taxon	description	urn: lsid: zoobank. org: act: CCA 055 CD- 6549 - 420 A-BD 70 - 415692 EE 1280 Figs 7 – 14; Table 2	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE4FFA64421A41460D1F3D9.taxon	diagnosis	Diagnosis Metabiantes elongatus sp. nov. differs from the rest of the species of the genus (except M. serratus sp. nov., M. litoralis, and M. zuluanus) by the following combination of characteristics: presence of two tubercles on mesotergal areas III – V (Figs 7 A, 8 A, 13 A) and, in males, having a slender femur II (Figs 10 A, 14 B, D) and metatarsus II with tubercles on the ventral region (Figs 10 E, 14 H). Metabiantes elongatus and M. serratus share a remarkably similar male genital morphology but M. elongatus can be easily differentiated from M. serratus in lacking tubercles on free tergites I – II (Fig. 7 C vs Figs 33 C, 34 C) and in possessing a thinner femur II (Figs 10 A, 14 B, D vs Figs 36 A – B, 39 B, D), unswollen tibia (Figs 10 C, 14 F vs Figs 36 C, 39 F), and unswollen metatarsus of leg II (Figs 10 E, 14 H vs Figs 36 E, 39 H). Additionally, M. elongatus lacks a pronounced constriction at astragalus-calcaneus junction in metatarsus of leg II as observed in M. serratus (Fig. 10 E – F vs Fig. 36 E – F). Metabiantes elongatus can be differentiated from M. litoralis by having granules on free tergites instead of the presence of a row of small tubercles in M. litoralis (Fig. 7 C vs Kauri 1961: fig. 33 a – b). Males of Metabiantes elongatus have a non-enlarged trochanter II (Fig. 7 A). In contrast, males of M. zuluanus exhibit a remarkably swollen trochanter II (Lawrence 1937 a: fig. 26 a). Regarding male genital morphology, M. elongatus has a penis with a deeper U-shaped cleft of lamina apicalis, wider titillators, and remarkably smaller basal setae, easily differentiated from the penis of M. litoralis and M. zuluanus with a shallow cleft, narrow titillators, and bigger basal setae (Figs 11 B – D, 12 A – C vs Kauri 1961: figs 22 a – b, 34 a – b).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE4FFA64421A41460D1F3D9.taxon	etymology	Etymology The species epithet is derived from the Latin word ‘ longatus ’, meaning ‘ elongate’ and refers to the elongated femur II in males of this species.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE4FFA64421A41460D1F3D9.taxon	materials_examined	Type material Holotype CONGO • ♂; Bas-Congo, Mayombe, Luki Forest Reserve; 5.63333 ° S, 13.06667 ° E; 12 Nov. 2006; D. De Bakker and J. P. Michiels leg.; primary rainforest; fogging; RMCA, BE _ RMCA _ ARA. Opi. 219865. Paratypes CONGO • 1 ♂, 1 ♀; same data as for holotype; RMCA, BE _ RMCA _ ARA. Opi. 247662 • 1 ♂ (SEM voucher); same data as for holotype; MACN-Ar 45425 • 1 ♂ (SEM voucher); same data as for holotype; MACN-Ar 45430 • 9 ♂♂ (1 photo voucher), 13 ♀♀ (1 photo voucher, 2 SEM vouchers); same data as for holotype; MACN-Ar 45440 • 9 ♂♂, 6 ♀♀; same data as for holotype; 13 Nov. 2006; RMCA, BE _ RMCA _ ARA. Opi. 219866 • 10 ♂♂, 16 ♀♀; same data as for holotype; 5 Nov. 2006; RMCA, BE _ RMCA _ ARA. Opi. 219858.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE4FFA64421A41460D1F3D9.taxon	description	Description Male (holotype, BE _ RMCA _ ARA. Opi. 219865) BODY MEASUREMENTS. Total body length 1.95, carapace length 0.66, scutum magnum length 1.84, carapace maximum width 1.08, abdominal scutum maximum width 1.62. Appendage measurements in Table 2. DORSUM. Outline slightly hourglass-shaped with Eta (η) shape, with a very slight constriction located at sulcus I level (Figs 7 A, 8 A). Carapace with scattered granules, wider than long; anterior border slightly convex and unarmed (Figs 7 A, 8 A). Cheliceral sockets not marked (Fig. 8 A). Eyes separated near sulcus I; interocular area with scattered granules (Figs 7 A, C, 8 A, C). Carapace in lateral view straight at anterior region and slightly higher posteriorly (Figs 7 C, 8 C). Sulcus I deep and complete, medially slightly curved toward the posterior body region (Fig. 8 A). Mesotergal areas I – IV granulated and well-defined, with sulci II – IV marked but shallower than sulcus I; medially sulcus II slightly curved to the anterior body region; sulci III – V straight (Fig. 8 A). Mesotergal areas III – IV medially with two conical and pointed setiferous tubercles (Figs 7 A, 8 A). Mesotergal area V with two irregular rows of granules and four medial tubercles in the posterior margin (Fig. 8 A). Lateral margins with a row of granules (Figs 7 A, 8 A). Ozopore with an oval and narrow orifice with a descending channel that extends toward the ventroposterior region (Fig. 8 C). Free tergites granulated (Fig. 7 C). VENTER. Coxa I with few, small, medial setiferous granules; coxa II incrassated, slightly smaller than coxa IV (Figs 7 B, 8 B); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Fig. 8 B). Posterior border of spiracular area, free sternites I – V with a row of granules; anal operculum granulated (Fig. 8 B) Spiracles not concealed (Fig. 8 B). CHELICERA. Basichelicerite unarmed with slightly marked bulla (Fig. 9 G). Cheliceral hand with sparse setae (Fig. 9 G – H). Fixed finger with spaced, triangular-shaped teeth; movable finger with a row of small, rounded teeth (Fig. 9 H). PEDIPALP. Coxa elongated (i. e., remarkably longer than trochanter), proximally with three granules – one dorsoectal, one ectal (Fig. 8 A), and one ventroectal (Fig. 8 B). Trochanter with one small ventroectal setiferous granule (Fig. 8 B). Femur straight, proximally with a strong ventral narrowing followed by a small ventromesal spine (Fig. 9 A – B); ventrally with scattered pores and surface texture with scales-like appearance (Fig. 9 B, D). Patella elongated, club-shaped, and armed with a short mesodistal spine (Fig. 9 A). Tibia with two ventroectal and two long ventromesal spines (Fig. 9 A, C); tibia ventrally with scattered pores and scales-like surface (Fig. 9 E). Tarsus with two ventromesal and two ventroectal spines; proximal spines longer than distal spines (Fig. 9 A, C). Setae of spines with a basal portion smooth, then with scattered microtrichia (Fig. 9 C, E); microtrichia with a wide base and rounded tip (Fig. 9 F). LEGS. Coxa II with a dorsal row of granules (Fig. 8 C). Femur II elongated and slightly fusiform, swelling dorsally, slightly more pronounced, unarmed (Figs 7 C, 10 A – B, 14 D). Tibia II elongated ventrally with small triangular-shaped tubercles (Figs 10 C – D, 14 F). Metatarsus II with elongated astragalus, armed with equidistant transverse rows of triangular-shaped tubercles (Figs 10 E – G, 14 H). Calcaneus occupies a reduced distal portion of the metatarsus; calcaneus with scattered long sensilla chaetica and trichomes distributed along all surfaces, with higher density on the ventral region; trichomes of variable length with a wider ovate-shaped base and pointed tip (Fig. 10 F, H). Tarsi III – IV with a dense scopula. Tarsal formula: 3 (2): 5 (4): 5: 5. COLOR (specimen preserved in 80 % ethanol). Body brown yellowish; anterior border and lateral margins of carapace, lateral area to eyes until anterolateral margin of mesotergal area I with brown reticulations (Fig. 7 A – C). Appendages light brown yellowish; metatarsi and tarsi I – IV darker (Figs 7 A – C, 14 B, D, F, H). MALE GENITALIA. Penis with clearly defined boundaries between pars basalis and pars distalis (Fig. 11 A). Pars basalis basally thin, broadens medially, with distal constriction (Fig. 11 A). Pars distalis swollen with maximum width at titillator level (Figs 11 B, D, 12 A, C); apical edge laminar (i. e., dorsoventrally flat) with a medial U-shaped cleft dividing it into two rounded halves (Figs 11 B, D, 12 A, C). Halves apically curved towards ventral side, less chitinous and irregularly deformed, probably signifying that could be inflated by hemolymph pressure (Figs 11 C – D, 12 B – C, F). Pars distalis with a slight distal depression in the ventromedial region (Figs 11 D, 12 C, F). Each side of pars distalis armed with short, conical microsetae irregularly arranged, extending basally from dorsolateral to ventrodistal region (Figs 11 B – D, 12 A – C). Capsula externa with two broad titillators that cover almost all the capsula interna (Figs 11 B, 12 A). Capsula interna with two complex conductors and one stylus, basally fused. Each conductor with two medial laminar folds apically, one short dorsal and one ventral longer, visible ventrally within the U-shaped cleft (Figs 11 B, D, 12 A, C – G). Ventral folds with apical edges in contact, waved, medially free edges dorsally (Fig. 12 D) and ventrally folded, with a strongly curved and shorter subapical edge hoodshaped (Fig. 12 C, E, G). Each conductor also bearing one broad lateral projection (Fig. 11 B, D); stylus tubular, S-shaped in lateral view, with its free tip fully covered by conductors (Fig. 11 B – D). Female (paratype, MACN-Ar 45440) BODY MEASUREMENTS. Total body length 1.97, carapace length 0.67, scutum magnum length 1.81, carapace maximum width 1.04, abdominal scutum maximum width 1.71. Appendage measurements in Table 2. BODY. Resembles that of males in the armature of the scutum magnum (Fig. 13 A – B vs Figs 7 A, C, 8 A, C). Leg II not dimorphic (Fig. 14 A); femur II not swollen as in male (Fig. 14 A, C vs Fig. 14 B, D); tibia II thin as in male but without ventral tubercles (Fig. 14 E vs Fig. 14 F); metatarsus II thin and unarmed (Fig. 14 G vs Fig. 14 H). Tarsal formula 3 (2): 5 (4): 5: 5. FEMALE GENITALIA. Ovipositor cylindrical (Fig. 13 C), distally bearing two lobes (furca) (Fig. 13 C – D, F). Each furcal lobe with five long, pointed setae (Fig. 13 E) – three dorsal and two ventral – resulting in a total of six setae on the dorsal region (Fig. 13 D) and four on the ventral region (Fig. 13 F). External surface of dorsal and ventral furcal lobes medially with several short, pointed projections, irregularly distributed (Fig. 13 D, F). Receptacle chambers located near the base of the furcal groove (Fig. 13 D – F).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFE4FFA64421A41460D1F3D9.taxon	distribution	Distribution Known only from the type locality (Fig. 40).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFFAFFA94421A47460D1F7B3.taxon	description	urn: lsid: zoobank. org: act: D 394 CC 39 - C 444 - 49 BC- 8707 - 824153 F 3 ED 9 A Figs 15 – 20; Table 3	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFFAFFA94421A47460D1F7B3.taxon	diagnosis	Diagnosis Metabiantes herculeus sp. nov. differs from its congeners (except M. kaurii sp. nov., M. kivuensis sp. nov., M. machadoi, M. obscurus, M. pusulosus, and M. zuurbergianus) by the following combination of traits: absence of tubercles on mesotergal areas III – V and free tergites; sexually dimorphic male leg II with a thickened femur and patella, a broad tibia, and a metatarsus ventrally armed with tubercles (Figs 17 A – F, 20 B, D, F, H). Males of M. herculeus differ from those of M. kaurii and M. kivuensis by having ventral tubercles on tibia II, absent in the latter species (Fig. 17 C – D vs Figs 23 C – D, 28 C – D). The penis of M. herculeus is distinctive with conductors closely together and lacking lateral projections, unlike the widely separated conductors in M. kaurii (Fig. 18 B vs Fig. 24 B) and the presence of lateral projections in M. kivuensis (Fig. 18 B vs Fig. 29 B). Additionally, the shorter, non-contacting halves of the lamina apicalis in M. herculeus contrast with the longer, contacting halves in M. kaurii (Fig. 18 B, D vs Fig. 24 B, D). Unlike M. kivuensis, M. herculeus lacks a longitudinal division of mesotergal area IV into two halves, which is characteristic of females and major males of M. kivuensis (Figs 15 A, 16 A, 19 A vs Figs 25 A, 31 A). Furthermore, males of M. herculeus lack an enlarged trochanter II, distinguishing them from M. pusulosus (Fig. 15 A vs Kauri 1961: fig. 5 a). Also, males of M. herculeus are easily distinguished from those of M. machadoi by their abruptly thickened femur II (Fig. 17 A – B vs Lawrence 1957: fig. 3 b) and from those of M. zuurbergianus by the presence of ventral tubercles on tibia II (Fig. 17 C – D vs Kauri 1961: 25). The penis of M. herculeus features a deeper U-shaped cleft of the lamina apicalis, wider titillators, and smaller basal setae, which distinguish it from the shallow cleft, narrow titillators, and larger basal setae in M. obscurus and M. zuurbergianus (Fig. 18 B – D vs Kauri 1961: figs 7 a – b, 11 a – b). Additionally, the rounded pars distalis with wide titillators in M. herculeus differs from the triangular lateral edges of pars distalis and narrow titillators in M. pusulosus (Fig. 18 B – D vs Kauri 1961: fig. 1 a – b).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFFAFFA94421A47460D1F7B3.taxon	etymology	Etymology The species name is derived from the Latin word Hercules, the Roman counterpart of the Greek hero Heracles, the most popular figure in ancient Greek mythology. It can be understood as something related to or characteristic of Hercules, often indicating strength and power qualities, specifically referring to the strongly thickened femur II of the species.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFFAFFA94421A47460D1F7B3.taxon	materials_examined	Type material Holotype TANZANIA • ♂; Mbeya Region, Tukuyu, Ushirika area, Kayuki tea estate; 9.41667 ° S, 34.66667 ° E; 1200 m a. s. l.; 29 Nov. 1991; R. Jocqué leg.; litter; RMCA, BE _ RMCA _ ARA. Opi. 173473. Paratype TANZANIA • 1 ♀; same data as for holotype; RMCA, BE _ RMCA _ ARA. Opi. 247663.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFFAFFA94421A47460D1F7B3.taxon	description	Description Male (holotype, BE _ RMCA _ ARA. Opi. 173473) BODY MEASUREMENTS. Total body length 2.52, carapace length 0.86, scutum magnum length 2.07, carapace maximum width 1.43, abdominal scutum maximum width 1.68. Appendage measurements in Table 3. DORSUM. Outline slightly hourglass-shaped with Eta (η) shape, with a very slight constriction located at sulcus I level (Figs 15 A, 16 A); carapace with scattered granules, wider than long, with a small and rounded frontal hump (Figs 15 A, 16 A); anterior border slightly convex and unarmed (Figs 15 A, 16 A). Cheliceral sockets not marked (Fig. 16 A). Eyes separated near sulcus I; interocular area finely granulated (Figs 15 A, E, 16 A). Carapace in lateral view straight posterior to frontal hump and becoming slightly higher toward the posterior region (Fig. 15 E). Abdominal scutum in lateral view convex (Fig. 15 E). Sulcus I deep and complete (Fig. 16 A). Mesotergal areas granulated and well-defined, with sulci II – V marked but shallower than sulcus I; medially sulci II – III slightly curved to anterior body region; sulci IV – V straight (Fig. 16 A). Mesotergal area V granulated (Fig. 16 A). Lateral margins of abdominal scutum with two rows of granules (Fig. 16 A). Free tergites granulated (Figs 15 E, 16 A). VENTER. Coxa I with few small setiferous granules (Fig. 15 C); coxa II incrassated, bigger than coxa IV (Fig. 15 B – D); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Fig. 15 C – D). Posterior border of spiracular area, free sternites I – V with a row of granules; anal operculum granulated (Fig. 15 D – F). Spiracles not concealed (Fig. 15 D). CHELICERA. Basichelicerite unarmed, with not marked bulla (Fig. 16 G). Cheliceral hand with sparse setae (Fig. 16 G – H). Fixed and movable fingers with small triangular-shaped teeth. PEDIPALP. Coxa elongated (i. e., remarkably longer than trochanter), dorsoproximally with one mesal and one ectal granule (Figs 15 A, 16 A). Trochanter smooth (Fig. 16 B – C). Femur straight, proximally with one small ventromesal spine, ventral surface with granules (Fig. 16 B – F). Patella elongated, club-shaped, with a small mesodistal spine (Fig. 16 B – C). Tibia with two ventroectal and two ventromesal spines; distal ventroectal spine with the highest socket and longest seta (Fig. 16 B – C). Tarsus inflated, with oval shape, armed with two ventromesal and two ventroectal spines, proximal spines longer than distal spines (Figs 15 E, 16 B – C); tarsus ventrally with granules on the medial surface (Fig. 16 C). LEGS. Femur II proximally thin, followed by an abrupt strong thickness, then tapering gradually (Figs 15 E, 17 A – B, 20 B, D). Patella II long, thickened (Figs 17 A, 20 B). Tibia II widened with ventral small triangular-shaped tubercles (Figs 17 C – D, 20 F). Metatarsus II with a very long calcaneus that occupies more than half of the ventral metatarsus (Fig. 17 E); astragalus short, with equidistant transverse rows of tubercles (Fig. 17 E); calcaneus with scattered low and rounded trichomes, long sensilla chaetica and glandular pores on the ventral surface (Fig. 17 E – F). Tarsi III – IV with a dense scopula. Tarsal formula: 3 (2): 5 (4): 5: 5. COLOR (specimen preserved in 80 % ethanol). Body brown-yellowish (Fig. 15 A, C – E); carapace, coxae I – IV, and appendages with brown reticulations (Figs 15 A – E); lateroanterior, lateral, and lateroposterior borders of mesotergal areas dark brown; mesotergal areas I – III with two medial dark brown patches, closer in mesotergal area III; mesotergal area IV with one posterior medial dark brown patch; mesotergal area V with two lateral dark brown patches; free tergites I – II with two lateral dark brown patches and one medial rounded spot; free tergite III dark brown (Fig. 15 A, E); posterior border of spiracular area and free sternites I – IV with a line of dark brown patches (Fig. 15 B – F); anal operculum dark brown (Fig. 15 F). MALE GENITALIA. Penis with clearly defined boundaries between pars basalis and pars distalis (Fig. 18 A). Pars basalis basally thin, broadens medially, with distal constriction (Fig. 18 A). Pars distalis slightly swollen with maximum width at titillator level (Fig. 18 B). Apical edge laminar (i. e., dorsoventrally flat) with a medial U-shaped cleft that divides it into two rounded and elongated halves (Fig. 18 B, D); these halves are apically less chitinous and could possibly be inflated by hemolymph pressure (Fig. 18 D). Pars distalis with a distal depression in the ventromedial region (Fig. 18 D). Each side of pars distalis armed with short, conical microsetae irregularly arranged, extending basally from the dorsolateral to the ventroapical side (Fig. 18 B – D). Capsula externa with two broad titillators separated by a dorsal cleft basally narrow (Fig. 18 B). Capsula interna formed by two conductors and one stylus, basally fused. Each one with two laminar folds apically, one small pointed dorsal fold and one longer ventral fold, ventrally visible within the U-shaped cleft; stylus tubular, and irregular S-shaped in lateral view with its free tip fully covered by conductors (Fig. 18 B – D). Female (paratype, BE _ RMCA _ ARA. Opi. 247663) BODY MEASUREMENTS. Total body length 2.16, carapace length 0.67, scutum magnum length 1.83, carapace maximum width 1.12, abdominal scutum maximum width 1.64. Appendage measurements in Table 3. BODY. Female resembles male in the armature of the scutum magnum (Fig. 19 A – B vs Fig. 15 A, E) but differs from male by having a thinner pedipalp tarsus (Fig. 19 B vs Fig. 15 E). Also, female differs by having a thin leg II (Fig. 20 A vs Fig. 20 B); femur and patella II thin (Fig. 20 A, C vs Fig. 20 B, D); tibia II thin and unarmed (Fig. 20 E vs Fig. 20 F); metatarsus II and basitarsus thin and unarmed, with pseudoarticular rings (Fig. 20 A, G vs Fig. 20 B, H). Tarsal formula 3 (2): 5 (4): 5: 5. FEMALE GENITALIA. Ovipositor cylindrical (Fig. 19 C), distally bearing two long lobes (furca) (Fig. 19 C – D, F). Each furcal lobe with five long, pointed setae (Fig. 19 E) – three dorsally and two ventrally – resulting in a total of six setae on the dorsal region (Fig. 19 D) and four on the ventral region (Fig. 19 F). Receptacle chambers located near the base of the furcal groove (Fig. 19 D, F).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFFAFFA94421A47460D1F7B3.taxon	distribution	Distribution Known only from the type locality (Fig. 40).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFF2FF924437A15960D1F43A.taxon	description	urn: lsid: zoobank. org: act: 44 F 823 B 9 - 361 E- 4 C 7 C-B 792 - D 685 D 7 DEC 49 F Figs 21 – 24; Table 4	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFF2FF924437A15960D1F43A.taxon	diagnosis	Diagnosis Metabiantes kaurii sp. nov. differs from its congeners (except M. herculeus sp. nov., M. kivuensis sp. nov., M. machadoi, M. obscurus, M. pusulosus, and M. zuurbergianus) by the following combination of traits: absence of tubercles on mesotergal areas III – V and free tergites; sexually dimorphic male leg II with a thickened femur, a broad tibia, and a metatarsus ventrally armed with tubercles (Figs 21 A, 22 A, 23 B, D, F). Male of M. kaurii differs from M. herculeus by the absence of ventral tubercles on tibia II, present in the latter species (Fig. 23 C – D vs Fig. 17 C – D). The penis of M. kaurii is distinctive with widely separated conductors and lacking lateral projections, unlike the close-together conductors in M. herculeus (Fig. 24 B vs Fig. 18 B) and the presence of lateral projections in M. kivuensis (Fig. 24 B vs Fig. 29 B). Additionally, the longer, contacting halves of the lamina apicalis in M. kaurii contrast with the shorter, non-contacting halves in M. herculeus (Fig. 24 B, D vs Fig. 18 B, D). Unlike M. kivuensis, M. kaurii lacks a longitudinal division of mesotergal area IV into two halves, which is characteristic of females and major males of M. kivuensis (Figs 21 A, 22 A vs Figs 25 A, 31 A). Furthermore, the male of M. kaurii lacks an enlarged trochanter II, distinguishing it from those of M. pusulosus (Fig. 21 A vs Kauri 1961: fig. 5 a). Also, the male of M. kaurii is distinguished from that of M. machadoi by its abruptly thickened femur II (Fig. 23 A vs Lawrence 1957: fig. 3 b). The penis of M. kaurii features a deeper U-shaped cleft of the lamina apicalis, wider titillators, and smaller basal setae, which distinguish it from the shallow cleft, narrow titillators, and larger basal setae in M. obscurus and M. zuurbergianus (Fig. 24 B – D vs Kauri 1961: figs 7 a – b, 11 a – b). Additionally, the slight triangular lateral edges of the pars distalis and wide titillators in M. kaurii differ from the stronger triangular lateral edges of the pars distalis and narrow titillators in M. pusulosus (Fig. 24 B – D vs Kauri 1961: fig. 1 a – b).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFF2FF924437A15960D1F43A.taxon	etymology	Etymology The species epithet is a patronym, honoring the Estonian arachnologist Hans Kauri (1906 – 1999) for his contributions to opilionology, particularly that of the African fauna.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFF2FF924437A15960D1F43A.taxon	materials_examined	Type material Holotype MOZAMBIQUE • ♂; Niassa; 12.38278 ° S, 35.33369 ° E; 1724 m a. s. l.; 14 Nov. 2006; L. Geeraert and M. Jocqué leg.; montane forest; pitfall trap; MACN-Ar 46476.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFF2FF924437A15960D1F43A.taxon	description	Description Male (holotype, MACN-Ar 46476) BODY MEASUREMENTS. Total body length 2.41, carapace length 0.71, scutum magnum length 2.28, carapace maximum width 1.08, abdominal scutum maximum width 1.53. Appendage measurements in Table 4. DORSUM. Outline hourglass-shaped with Eta (η) shape, with a slight constriction posteriorly at sulcus I level (Figs 21 A, 22 A). Carapace granulated, wider than long, anterior border convex, and unarmed (Figs 21 A, 22 A). Cheliceral sockets not marked (Fig. 22 E). Eyes separated near sulcus I; interocular area granulated (Figs 21 E, 22 A). Carapace in lateral view straight in anterior region and slightly higher posteriorly (Fig. 21 E). Abdominal scutum in lateral view convex (Fig. 21 E). Sulcus I deep and complete, curving slightly posteriorly at midline in dorsal view (Fig. 22 A). Mesotergal areas granulated and well-defined, with sulci II – V marked but shallower than sulcus I; sulci II – III slightly curved anteriorly; sulci IV – V straight. Mesotergal area V granulated (Figs 21 A, 22 A). Lateral margins of abdominal scutum with two rows of granules (Fig. 22 A). Free tergites with two rows of granules (Figs 21 E – F, 22 A). VENTER. Coxa I with some small setiferous granules; coxa II incrassated, of same size as (or slightly smaller than) coxa IV (Fig. 21 C); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Fig. 21 C). Posterior border of spiracular area, free sternites I – V with a row of granules (Fig. 21 D – F); anal operculum granulated (Fig. 21 D – F). Spiracles not concealed (Fig. 21 D). CHELICERA. Basichelicerite unarmed with a slightly marked bulla (Fig. 22 E). Cheliceral hand with sparse setae (Fig. 22 E – F). Fixed and movable fingers with small triangular-shaped teeth (Fig. 22 F). PEDIPALP. Coxa elongated (i. e., remarkably longer than trochanter), dorsoproximally with one mesal and one ectal granule (Figs 21 A, 22 A). Trochanter unarmed. Femur straight, proximally with one small ventromesal spine (Fig. 22 B – D). Patella elongated, club-shaped, and armed with a small distal ventromesal spine (Fig. 22 B). Tibia with two ventromesal and two ventroectal spines, with distal ventroectal spine longest (Fig. 22 B – C). Tarsus inflated, of spheroid shape, armed with two ventromesal and two ventroectal spines; proximal spines longer than distal spines; ventroectal spine with the highest elevated socket; tarsus ventromedially with small granules (Fig. 22 B – C). LEGS. Femur II unarmed, proximally thin, followed by an abrupt strong thickness, then tapering gradually (Fig. 23 A – B). Patella II short and unarmed. Tibia II widened and unarmed (Fig. 23 C – D). Metatarsus III with astragalus occupying the half region (Fig. 23 E – G), ventrally with triangular-shaped tubercles (Fig. 23 F – I). Calcaneus occupies half of metatarsus, with scattered low and rounded trichomes and long sensilla chaetica along the lateral surface (Fig. 23 H – J). Tarsi III – IV with a dense scopula. Tarsal formula: 3 (2): 5 (4): 5: 5. COLOR (specimen preserved in 80 % ethanol). Body brown yellowish (Fig. 21 A – F); carapace with brown reticulations on anterior and lateral regions (Fig. 21 A). Lateroanterior, lateral, and lateroposterior borders of mesotergal areas I – III and anal operculum dark brown (Fig. 21 A, F); lateroanterior, lateral, and posterior borders of mesotergal area IV with dark brown patches (Fig. 21 A). Mesotergal area V and free tergites with two lateral dark brown patches (Fig. 21 A); coxae I – IV with brown reticulations; free sternites with anterior dark brown patches (Fig. 21 B, D). Appendages with brown reticulations or dark patches (Fig. 21 A – B, E). MALE GENITALIA. Pars basalis and pars distalis with indistinct boundaries, defined by the start of the most basal macrosetae marking the beginning of pars distalis (Fig. 24 A – D). Pars basalis tubular, basally thin, gradually widening toward distal region (Fig. 24 A); pars distalis slightly swollen, with maximum width at titillator level (Fig. 24 B), and ventrally with slight angular lateral edges (Fig. 24 D). Apical edge laminar (i. e., dorsoventrally flat) with a deep medial U-shaped cleft dividing it into two elongated, rounded halves, apically in contact (Fig. 24 A – B, D). Pars distalis with a small distal depression in the ventromedial region (Fig. 24 D). Each side of pars distalis bearing small conical microsetae irregularly arranged, extending from the dorsal to ventroapical region (Fig. 24 B – D). Capsula externa with two broad titillators covering most of the capsula interna; titillators separated by a U-shaped cleft at the base, narrowing apically (Fig. 24 B – C). Capsula interna formed by two laminar conductors and one stylus basally fused. Conductors apically curved toward medial region; conductor tips ventrally visible within U-shaped cleft. Stylus tubular, basally wide, apically thin, and with an irregular S-shaped curve in lateral view (Fig. 24 C), with a rounded tip (Fig. 24 B, D). Female Unknown.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFF2FF924437A15960D1F43A.taxon	distribution	Distribution Known only from the type locality (Fig. 40).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFCFFF994422A15960D1F37F.taxon	description	urn: lsid: zoobank. org: act: 364 EA 7 D 6 - 9176 - 4 A 30 - BE 56 - A 53 A 50 C 64 FD 5 Figs 25 – 32; Table 5	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFCFFF994422A15960D1F37F.taxon	diagnosis	Diagnosis Major males and females (excluding minor males) of Metabiantes kivuensis sp. nov. can be distinguished from those of all other species of Metabiantes by the presence of a longitudinal division of mesotergal area IV into two halves (Figs 25 A, 31 A). Additionally, M. kivuensis differs from its congeners (except M. herculeus sp. nov., M. kaurii sp. nov., M. machadoi, M. obscurus, M. pusulosus, and M. zuurbergianus) by the following combination of traits: absence of tubercles on mesotergal areas III – V and free tergites; sexually dimorphic male leg II with a thickened femur, a broad tibia, and a metatarsus ventrally with tubercles (Figs 26 A, 28 A – F, 32 B – C, E – F, H – I, K – L). Males of M. kivuensis differ from those of M. herculeus by the absence of ventral tubercles on tibia II, present in the latter species (Fig. 28 C – D vs Fig. 17 C – D). The penis of M. kivuensis sp. nov. is distinctive with closely together conductors and the presence of lateral projections, unlike the widely separated conductors in M. kaurii (Fig. 29 B vs Fig. 24 B) and the absence of lateral projections in M. herculeus (Fig. 29 B vs Fig. 24 B). Furthermore, males of M. kivuensis lack an enlarged trochanter II, distinguishing them from those of M. pusulosus (Fig. 25 A, E vs Kauri 1961: fig. 5 a). Also, major males of M. kivuensis are distinguished from those of M. machadoi by their abruptly thickened femur II (Fig. 25 C vs Lawrence 1957: fig. 3 b). The penis of M. kivuensis features a deeper U-shaped cleft of the lamina apicalis and smaller basal setae, which distinguish it from the shallow cleft and larger basal setae in M. obscurus and M. zuurbergianus (Fig. 29 B – D vs Kauri 1961: figs 7 a – b, 11 a – b). Additionally, the rounded pars distalis in M. kivuensis differs from the stronger angular lateral edges of pars distalis in M. pusulosus (Fig. 29 B – D vs Kauri 1961: fig. 1 a – b).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFCFFF994422A15960D1F37F.taxon	etymology	Etymology The species epithet ‘ kivuensis ’ means ‘ of or from Kivu’, a reference to the species type locality in North Kivu, a province of the Democratic Republic of the Congo.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFCFFF994422A15960D1F37F.taxon	materials_examined	Type material Holotype CONGO • major ♂; Kivu-N, Kilindera, “ Face N. du Ruwenzori, camp de Kilindera ” [North face of Ruwenzori, Kilindera camp]; 0.38333 ° N, 29.91667 ° E; 2950 m a. s. l.; Jul. – Aug. 1974; M. Lejeune leg.; “ litière du bambusetum du Musoso ” [litter of bamboo vegetation along the Musoso]; RMCA, BE _ RMCA _ ARA. Opi. 154162. Paratypes CONGO • 1 major ♂, 1 minor ♂ (photo voucher), 1 ♀ (photo voucher); same data as for holotype; RMCA, BE _ RMCA _ ARA. Opi. 247664 • 1 minor ♂ (SEM voucher); same data as for holotype; MACNAr 45445 • 1 major ♂; same data as for holotype; MACN-Ar 45437. Other material examined CONGO • 1 major ♂, 2 ♀♀, 2 juvs; Kivu-C, Kanzuiri, “ Face N. du Ruwenzori, camp de Kanzuiri, crête du Kanzuiri ” [North face of Ruwenzori, Kilindera camp, Kanzuiri Ridge]; 0.41667 ° N, 29.9 ° E; 3500 m a. s. l.; Jul. – Aug. 1974; M. Lejeune leg.; “ dans mousses ” [in mosses]; RMCA, BE _ RMCA _ ARA. Opi. 154244 • 1 minor ♂, 1 ♀; same data as for preceding; MACN-Ar 45470.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFCFFF994422A15960D1F37F.taxon	description	Description Male (holotype, BE _ RMCA _ ARA. Opi. 154162) BODY MEASUREMENTS. Total body length 2.95, carapace length 0.94, scutum magnum length 2.58, carapace maximum width 1.43, abdominal scutum maximum width 2.09. Appendage measurements in Table 5. DORSUM. Outline slightly hourglass-shaped with Eta (η) shape, with a very slight constriction at sulcus I level (Figs 25 A, 26 A). Carapace wider than long with a row of granules on the posterior margin and granules concentrated laterally around eyes, anterior border slightly convex and unarmed (Fig. 26 A); Cheliceral sockets not marked (Fig. 26 A). Eyes separated near sulcus I; interocular area smooth (Figs 25 A, 26 A). Carapace in lateral view straight at anterior region and slightly higher posteriorly (Figs 25 C, 26 C). Abdominal scutum in lateral view convex (Figs 25 C, 26 C). Sulcus I deep, complete, and slightly curved to the anterior body region (Figs 25 A, 26 A). Mesotergal areas granulated and well-defined, with sulci II – V marked but shallower than sulcus I; sulci II – III medially arched to the anterior body region; sulcus IV medially arched to posterior body region (Fig. 25 A); sulcus V medially invaginated towards mesotergal area IV dividing it into two (Fig. 25 A). Mesotergal area V with two rows of granules (Fig. 26 A). Lateral borders of abdominal scutum with two rows of granules (Fig. 26 A). Ozopore with an oval and narrow orifice with a descending channel that extends toward the ventroposterior region (Fig. 26 C). Free tergites granulated (Fig. 26 A, C). VENTER. Coxa I with few small medial setiferous granules (Fig. 26 B); coxa II incrassated, slightly bigger than coxa IV (Fig. 25 B); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Fig. 26 B). Lateroposterior border of spiracular area and lateral border of free sternites I – V with rows of granules (Fig. 26 B – C); anal operculum granulated (Fig. 26 B – C). Spiracles not concealed (Fig. 26 B). CHELICERA. Basichelicerite unarmed with a slightly marked bulla (Figs 26 C, 27 F – G). Cheliceral hand with sparse setae (Fig. 27 F – H). Movable fingers with small, square-shaped teeth (Fig. 27 H). PEDIPALP. Coxa elongated (i. e., remarkably longer than trochanter), proximally with one dorsoectal granule and three ventroectal granules – one proximal and two distal ones (Figs 25 A, 26 A – B). Trochanter unarmed (Fig. 27 A). Femur robust, thick, and straight, ventroproximally with one mesal spine (Fig. 27 A); ventral surface with small granules and scattered pores (Fig. 27 B – C). Patella elongated, club-shaped, with a small mesodistal spine (Fig. 27 A). Tibia with two ventroectal and two ventromesal spines (Fig. 27 A). Tarsus inflated, spheroid in shape, with two ventromesal and two ventroectal spines; proximal spines longer than distal ones (Fig. 27 A, D); ventral surface with conspicuous tubercles containing aggregate pores (Fig. 27 D – E). LEGS. Coxa II with a row of dorsal granules (Fig. 26 C). Femur II proximally thin, followed by an abrupt strong thickness, then tapering gradually (Figs 25 C, 32 C, F). Patella II short and thickened (Figs 25 C, 32 C, F). Tibia II dorsally widened (Figs 25 C, 32 C, I) and unarmed (Figs 28 C – D, 32 C, I). Metatarsus II with elongated astragalus, ventrally slightly swollen and armed with small triangular-shaped tubercles (Fig. 28 A, E). Calcaneus occupies less than the last third of metatarsus, with trichomes and scattered long sensilla chaetica distributed along all surfaces; trichomes with wider ovate-shaped bases, pointed tips, and variable length (Fig. 28 E – F). Tarsi III – IV with a dense scopula. Tarsal formula: 3 (2): 5 (4): 5: 5. COLOR (specimen preserved in 80 % ethanol). Body yellowish; carapace with light brown reticulations on the anterior and lateral sides (Fig. 25 A, C). Anterior, lateral, and lateroposterior borders of mesotergal area I dark brown (Fig. 25 A). Lateroanterior, lateral, and posterior borders of mesotergal area II dark brown (Fig. 25 A). Lateroposterior border of mesotergal areas III – IV with dark brown patches. Mesotergal areas II – III medially with four medial dark points – two anterior and two posterior ones. Mesotergal area IV with two medial dark points in each half (Fig. 25 A). Posterior border and free tergites with a row of dark points (Fig. 25 A). Free sternite V and anal operculum with dark brown patches (Fig. 25 B). Appendages with light brown reticulations (Figs 25 A – C, 32 C, F, I, L). MALE GENITALIA. Penis with distinguishable limits between pars basalis and pars distalis (Fig. 29 A). Pars basalis tubular, thin at the base, slightly broadened apically, and ending in a slight constriction (Fig. 29 A). Pars distalis swollen, with maximum width at titillator level (Figs 29 B, D, 30 A). Apical edge, laminar (i. e., dorsoventrally flat) with a large, open U-shaped cleft medially dividing into two rounded halves (Figs 29 B, D, 30 A, C); halves apically curved ventrally, less chitinous, and potentially inflatable by hemolymph pressure (Fig. 30 C). Pars distalis with a small apical depression in the ventromedial region (Figs 29 D, 30 C). Each side of pars distalis armed with irregularly arranged microsetae, extending from dorsolateral to the ventrodistal region (Figs 29 B – D, 30 A – C). Capsula externa with two broad titillators ending in rounded tips, separated by a close U-shaped cleft; tips with inner rounded projections (Fig. 30 E – F). Capsula interna formed by two complex conductors, each one with one laminar medial small dorsal fold and one longer ventral fold, with a lateral extension covering partially the dorsal fold, visible within the U-shaped cleft (Figs 29 B, 30 A, D – F); each conductor also with one broad lateral projection (Fig. 29 B); stylus long and tubular, with its tip covered by conductors (Fig. 29 B – D). Minor male (paratype, BE _ RMCA _ ARA. Opi. 247664) BODY MEASUREMENTS. Total body length 2.74, carapace length 0.96, scutum magnum length 2.5, carapace maximum width 1.4, abdominal scutum maximum width 1.98. Appendage measurements in Table 5. BODY. Minor male resembles major male in the armature of the scutum magnum, except for mesotergal area IV, which is not divided (Figs 26 A, 31 G vs Fig. 25 A). Minor male has a dimorphic leg II but differs from major male by having remarkably thinner femur, patella, and tibia (Figs 31 H, 32 B, E, H vs Figs 25 C, 32 C, F, I); metatarsus II slightly thinner than that of major males (Fig. 32 K vs Fig. 32 L). Tarsal formula: 3 (2): 5 (3): 5: 5. Female (paratype, BE _ RMCA _ ARA. Opi. 247664) BODY MEASUREMENTS. Total body length 2.91, carapace length 0.89, scutum magnum length 2.56, carapace maximum width 1.4, abdominal scutum maximum width 2.2. Appendage measurements in Table 5. BODY. Female resembles both minor and major males in the armature of the scutum magnum (Fig. 31 A – B vs Figs 25 A, C, 31 G – H). Female differs from the minor male, but not from the major male, by having the mesotergal area IV divided (Fig. 31 A vs Figs 26 A, 31 G). Female differs from both major and minor males by having a thin pedipalp tarsus and lacking a dimorphic leg II (Fig. 31 B vs Figs 25 C, 31 H), with femur and patella thinner than in minor and major males (Fig. 32 D vs Fig. 32 E – F); tibia with similar width to that of the minor male and thinner than in the major male (Fig. 32 G vs Fig. 32 H – I); metatarsus II slightly thinner than that of minor and major males (Fig. 32 J vs Fig. 32 K – L). Tarsal formula 3 (2): 5 (4): 5: 5. FEMALE GENITALIA. Ovipositor cylindrical (Fig. 31 C), distally bearing two lobes (furca) (Fig. 31 C – D, F). Each furcal lobe with five long, pointed setae (Fig. 31 E) – three dorsal and two ventral ones – resulting in a total of six setae on the dorsal region (Fig. 31 D) and four on the ventral region (Fig. 31 F). External surface of furcal lobes with several short, pointed lateral projections, irregularly distributed (Fig. 31 E). Receptacle chambers located near the base of the furcal groove (Fig. 31 D – F).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFCFFF994422A15960D1F37F.taxon	distribution	Distribution Known only from the type locality (Fig. 40).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	description	urn: lsid: zoobank. org: act: 99512 C 0 D- 7 DDB- 411 F-BA 7 F- 57 D 07 D 853187 Figs 33 – 39; Table 6	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	diagnosis	Diagnosis Metabiantes serratus sp. nov. differs from the rest of the species of Metabiantes (except M. elongatus sp. nov., M. litoralis and M. zuluanus) by the following combination of characteristics: presence of tubercles on mesotergal areas III – V and sexually dimorphic male leg II with thickened femur, and metatarsus with tubercles on the ventral region (Figs 33 A, C, 34 A, C, 38 A – B, 36 A – H, 39 B, D, F, H). Metabiantes serratus and M. elongatus share a remarkably similar male genital morphology, but M. serratus can be easily differentiated from M. elongatus by the presence of tubercles on free tergites I – II in contrast with granules in M. elongatus (Fig. 34 A, C vs Fig. 7 C); M. serratus also differs from M. elongatus by having a strong, thickened femur and swollen tibia and metatarsus of leg II (Figs 36 A – C, E, 39 B, D, F, H vs Figs 10 A, C, E, 14 B, D, F, H). Additionally, M. serratus has a pronounced constriction at the astragalus-calcaneus junction, which is absent in M. elongatus (Figs 36 E – F, 39 H vs Figs 10 E – F, 14 H). Males of Metabiantes serratus lack an enlarged trochanter II, distinguishing them from those of M. zuluanus (Fig. 33 A, C vs Lawrence 1937 a: fig. 26). Regarding male genital morphology, M. serratus has the penis with a deeper U-shaped cleft of lamina apicalis, wider titillators, and remarkably smaller basal setae, easily differentiated from the penis of M. litoralis and M. zuluanus with shallow cleft, narrow titillators, and larger basal setae (Fig. 37 B – H vs Kauri 1961: fig. 34 a – b, 22 a – b).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	etymology	Etymology The species epithet ‘ serratus ’, from the Latin meaning ‘ toothed like a saw’ refers to the serrated ventral transverse rows of triangular tubercles of the metatarsus II in males of this species.	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	materials_examined	Type material Holotype CONGO • ♂; Bas-Congo, Mayombe, Luki Forest Reserve; 5.63333 ° S, 13.06667 ° E; 27 Sep. 2007; D. De Bakker and J. P. Michiels leg.; along trail in primary rainforest; sieving; RMCA, BE _ RMCA _ ARA. Opi. 223705. Paratypes CONGO • 1 ♂, 3 ♀♀; same data as for holotype; RMCA, BE _ RMCA _ ARA. Opi. 247665 • 1 ♂ (SEM voucher); same data as for holotype; MACN-Ar 45472 • 2 ♂♂ (1 SEM voucher), 1 ♀ (photo voucher); same data as for holotype; MACN-Ar 45474 • 6 ♂♂, 6 ♀♀; same data as for holotype; 1 Oct. 2007; RMCA, BE _ RMCA _ ARA. Opi. 223775	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	description	Description Male (holotype, BE _ RMCA _ ARA. Opi. 223705) BODY MEASUREMENTS. Total body length 1.73, carapace length 0.61, scutum magnum length 1.56, carapace maximum width 0.97, abdominal scutum maximum width 1.27. Appendage measurements in Table 6. DORSUM. Outline slightly hourglass-shaped with Eta (η) shape, with a very slight constriction at sulcus I level (Figs 33 A, 34 A). Carapace with scattered medial granules, wider than long, anterior border slightly convex and unarmed, with a small and rounded frontal hump (Figs 33 C, 34 C); interocular area with scattered granules (Fig. 34 A). Cheliceral sockets not marked (Fig. 34 A). Eyes separated near sulcus I. Carapace in lateral view straight posterior to frontal hump and becoming slightly higher toward the posterior region (Figs 33 C, 34 C). Abdominal scutum in lateral view convex (Figs 33 C, 34 C). Sulcus I deep, complete, and straight (Fig. 34 A). Mesotergal areas granulated and well-defined; sulci II – III medially curved to the anterior body region; sulci IV – V straight (Fig. 34 A). Mesotergal areas III – IV medially with two conical and pointed tubercles (Figs 33 A, C, 34 A, C). Mesotergal area V with two irregular rows of granules and medially with three conical tubercles (Figs 33 C, 34 A, C). Lateral borders of abdominal scutum with a row of granules (Fig. 34 A, C). Ozopore with an oval and narrow orifice with a descending channel that extends toward the ventroposterior region (Fig. 34 C). Free tergite I with a row of granules; free tergites I – II medially with three conical tubercles; free tergite III with a row of medial conspicuous granules (Fig. 34 A). VENTER. Coxa I with few small medial granules (Fig. 34 B); coxa II incrassated, of same size as (or slightly larger than) coxa IV (Figs 33 B, 34 B); anteroposterior borders of coxa III with a row of strong granules connecting with coxae II and IV, respectively (Figs 33 B, 34 B). Posterior border of spiracular area and free sternites I – V with a row of granules (Fig. 34 B); anal operculum granulated (Fig. 34 B – C). Spiracles not concealed (Fig. 34 B). CHELICERA. Basichelicerite unarmed, with a slightly marked bulla (Fig. 35 F – G). Cheliceral hand with sparse setae (Fig. 35 F – H). Movable fingers with small square-shaped teeth (Fig. 35 H). PEDIPALP. Coxa elongated (i. e., remarkably longer than trochanter), proximally with two granules – one dorsoectally and one ventroectally (Fig. 34 A – B). Trochanter unarmed (Fig. 35 A – B). Femur straight, proximally with a slight ventral narrowing followed by a small ventromesal spine (Fig. 35 A – B), ventral surface with granules and pores (Fig. 35 B, E). Patella elongated, club-shaped, and armed with a small mesodistal spine (Fig. 35 A). Tibia with two ventromesal and two ventroectal long spines (Fig. 35 A). Tarsus thin, with two ventromesal and two ventroectal spines; proximal spines longer than distal spines (Fig. 35 A, C). Setae of spines proximally smooth, then covered by scattered microtrichia (Fig. 35 C – D). LEGS. Femur II unarmed, thin proximally, followed by an abrupt thickness, and then tapering gradually (Figs 33 C, 36 A – B, 39 B, D). Patella II long, thickened, and unarmed (Figs 33 C, 36 A, 39 B, D). Tibia II distally widened, with ventral triangular-shaped tubercles that increase in size towards the distal region (Figs 33 C, 36 C – D, 39 B, F). Metatarsus II with astragalus ventrally swollen and armed with equidistant transverse rows of triangular-shaped tubercles (Figs 33 C, 36 E – G, 39 B, H). Limit astragalus-calcaneus defined by a strong constriction with a slightly incrassate calcaneus giving a peculiar form to distal region of the metatarsus (Figs 33 C, 36 E – F, 39 H). Calcaneus ventrally mostly with ovate-shaped trichomes and some ovate-shaped base and pointed tip trichomes of variable length; lateral and dorsal surfaces of calcaneus with long thin-pointed trichomes and scattered sensilla chaetica and glandular pores (Fig. 36 F, H). Tarsi III – IV with a dense scopula. Tarsal formula: 3 (2): 5 (4): 5: 5. COLOR (specimen preserved in 80 % ethanol). Body dark brown; medial and posterior regions of carapace yellowish (Fig. 33 A); ventral body region and appendages brown-yellowish with light brown reticulations (Fig. 33 B – C); metatarsus II dark brown (Figs 33 C, 39 B, H). MALE GENITALIA. Penis with distinct boundaries between pars basalis and pars distalis (Fig. 37 A). Pars basalis tubular, slightly broadened apically, and ends in a constriction (Fig. 37 A). Pars distalis swollen with maximum width at basal level of titillators (Fig. 37 B, D, F). Apical edge, laminar (i. e., dorsoventrally flat), with a medial U-shaped cleft dividing into two rounded halves (Fig. 37 B, D, F, H); halves apically less chitinous and curved ventrally (Fig. 37 C, G – H). Pars distalis with a distal depression in the ventromedial region (Fig. 37 D, H). Each side of pars distalis armed with short, conical microsetae, irregularly arranged, extending basally from dorsolateral to the ventrodistal region (Fig. 37 B, C, F – H). Capsula externa with two broad titillators separated by a narrow cleft (Fig. 37 B – C, E – G). Capsula interna is formed by two complex conductors and one stylus, basally fused. Each conductor apically with one small medial dorsal fold and one longer ventral fold, ventrally visible within the U-shaped cleft (Fig. 37 B, D, I – J); each conductor also with one broad lateral projection, visible in the everted condition (Fig. 37 B, D – E). Stylus with a rounded tip (Fig. 37 E), wider basally, narrow distally, and with an irregular S-shaped curve in lateral view (Fig. 37 B – E). Female (paratype, MACN-Ar 45474) BODY MEASUREMENTS. Total body length 1.83, carapace length 0.52, scutum magnum length 1.5, carapace maximum width 0.89, abdominal scutum maximum width 1.24. Appendage measurements in Table 6. BODY. Female resembles males in the armature of the scutum magnum (Fig. 38 A – B vs Fig. 33 A, C) but differs by lacking dimorphic leg II; femur and patella not swollen as in male (Fig. 39 A, C vs Fig. 39 B, D); tibia thinner and unarmed (Fig. 39 E vs Fig. 39 F); metatarsus not swollen and unarmed (Fig. 39 G vs Fig. 39 H). Tarsal formula 3 (2): 4 (3): 5: 5. FEMALE OVIPOSITOR. Ovipositor cylindrical (Fig. 38 C), distally bearing two lobes (furca) (Fig. 38 C – D, F). Each furcal lobe with five long, pointed setae (Fig. 38 E) – three dorsal and two ventral – resulting in a total of six setae on the dorsal region (Fig. 38 D) and four on the ventral region (Fig. 38 F). Receptacle chambers located near the base of the furcal groove (Fig. 38 D – F).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
03C2879BFFC5FF86442BA32F60D1F3D7.taxon	distribution	Distribution Known only from the type locality (Fig. 40).	en	Mamani, Vanesa, Jocqué, Merlijn, Pérez-González, Abel (2025): Notes on African Biantinae with sexual dimorphism in leg II: five new Metabiantes species and redescription of Clinobiantes paradoxus (Opiliones: Laniatores: Biantidae). European Journal of Taxonomy 1006: 1-58, DOI: 10.5852/ejt.2025.1006.2989, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2989/13441
