taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896560/files/figure.png	https://doi.org/10.5281/zenodo.16896560	Figure 2. Mitochondrial tree of Tylototriton obtained from the ML phylogenetic analysis of the concatenated ND2 and 16S rRNA gene fragments (1665 bp). See Table 2 for voucher specimen information and GenBank accession numbers. Red, blue and green colors denote the new species Tylototriton koliaensis sp. nov., T. ziegleri s. str., and T. thaiorum, respectively (color codes as in Fig. 1). Numbers in brackets correspond to locality numbers (Fig. 1, Table 1). Numbers at tree nodes correspond to UFBS/PP support values. Photographs by Nikolay A. Poyarkov.	Figure 2. Mitochondrial tree of Tylototriton obtained from the ML phylogenetic analysis of the concatenated ND2 and 16S rRNA gene fragments (1665 bp). See Table 2 for voucher specimen information and GenBank accession numbers. Red, blue and green colors denote the new species Tylototriton koliaensis sp. nov., T. ziegleri s. str., and T. thaiorum, respectively (color codes as in Fig. 1). Numbers in brackets correspond to locality numbers (Fig. 1, Table 1). Numbers at tree nodes correspond to UFBS/PP support values. Photographs by Nikolay A. Poyarkov.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896566/files/figure.png	https://doi.org/10.5281/zenodo.16896566	Figure 5. Cranial morphology of Tylototriton koliaensis sp. nov. (ZMMU A-8157, holotype; A–C) and T. ziegleri s. str. (ZMMU A-8169; D–F) visualized via micro-CT scanning. Skulls are shown in dorsal (A, D), ventral (B, E) and lateral (C, F) aspects. Abbreviations: a.f.sq. – fronto-squamosal arc; art. – articular; c.int. – internasal fontanelle; c.occ. – occipital condyle; den. – dentary; exoc.+prot. – prootic–exoccipital; f.a.m. – antero-medial fenestra; f.o. – foramen opticum; fr. – frontal; mx. – maxilla; nas. – nasal; oper. – operculum; osph. – orbitosphenoid; p.cor. – coronoid process; par. – parietal; pfr. – prefrontal; pmx. – premaxilla; prart. – prearticular; psph. – parasphenoid; pter. – pterygoid; q. – quadrate; sq. – squamosal; vom. – vomer.	Figure 5. Cranial morphology of Tylototriton koliaensis sp. nov. (ZMMU A-8157, holotype; A–C) and T. ziegleri s. str. (ZMMU A-8169; D–F) visualized via micro-CT scanning. Skulls are shown in dorsal (A, D), ventral (B, E) and lateral (C, F) aspects. Abbreviations: a.f.sq. – fronto-squamosal arc; art. – articular; c.int. – internasal fontanelle; c.occ. – occipital condyle; den. – dentary; exoc.+prot. – prootic–exoccipital; f.a.m. – antero-medial fenestra; f.o. – foramen opticum; fr. – frontal; mx. – maxilla; nas. – nasal; oper. – operculum; osph. – orbitosphenoid; p.cor. – coronoid process; par. – parietal; pfr. – prefrontal; pmx. – premaxilla; prart. – prearticular; psph. – parasphenoid; pter. – pterygoid; q. – quadrate; sq. – squamosal; vom. – vomer.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896568/files/figure.png	https://doi.org/10.5281/zenodo.16896568	Figure 6. Holotype of Tylototriton koliaensis sp. nov. (ZMMU A-8157, female) in preservative. A, dorsal view; B, ventral view; C, head, dorsal view; D, head, ventral view; E, head, lateral view; F, palmar view of right hand; G, plantar view of left foot; H, ventral view of cloacal area. Photographs by Nikolay A. Poyarkov.	Figure 6. Holotype of Tylototriton koliaensis sp. nov. (ZMMU A-8157, female) in preservative. A, dorsal view; B, ventral view; C, head, dorsal view; D, head, ventral view; E, head, lateral view; F, palmar view of right hand; G, plantar view of left foot; H, ventral view of cloacal area. Photographs by Nikolay A. Poyarkov.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896570/files/figure.png	https://doi.org/10.5281/zenodo.16896570	Figure 7. Schematic drawing of vomers and vomerine tooth series of Tylototriton koliaensis sp. nov. holotype (ZMMU A-8157). Drawing by Dmitriy V. Arkhipov.	Figure 7. Schematic drawing of vomers and vomerine tooth series of Tylototriton koliaensis sp. nov. holotype (ZMMU A-8157). Drawing by Dmitriy V. Arkhipov.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896572/files/figure.png	https://doi.org/10.5281/zenodo.16896572	Figure 8. Specimens of Tylototriton koliaensis sp. nov. in life from Kolia Mountain Pass, Phia Oac-Phia Den National Park, Nguyen Binh, Cao Bang Province, Vietnam. A, holotype ZMMU A-8157, female, in situ; B, paratype ZMMU A-8158, male, in situ. Photographs by Nikolay A. Poyarkov.	Figure 8. Specimens of Tylototriton koliaensis sp. nov. in life from Kolia Mountain Pass, Phia Oac-Phia Den National Park, Nguyen Binh, Cao Bang Province, Vietnam. A, holotype ZMMU A-8157, female, in situ; B, paratype ZMMU A-8158, male, in situ. Photographs by Nikolay A. Poyarkov.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896574/files/figure.png	https://doi.org/10.5281/zenodo.16896574	Figure 9. Specimens of Tylototriton ziegleri s. str. in life. A, male ZMMU A-8167 from Thanh Van Village, Quang Ba, Bat Dai Son Nature Reserve, Ha Giang Province, Vietnam, in situ; B, male from Chieu Lau Thi Mt., Ha Giang Province, Vietnam, in situ (not collected); C, male from Wenshan, Yunnan Province, China, in situ (not collected). Photographs by Nikolay A. Poyarkov (A), Loi Phuoc Tran (B), and Yinpeng Zhang (C).	Figure 9. Specimens of Tylototriton ziegleri s. str. in life. A, male ZMMU A-8167 from Thanh Van Village, Quang Ba, Bat Dai Son Nature Reserve, Ha Giang Province, Vietnam, in situ; B, male from Chieu Lau Thi Mt., Ha Giang Province, Vietnam, in situ (not collected); C, male from Wenshan, Yunnan Province, China, in situ (not collected). Photographs by Nikolay A. Poyarkov (A), Loi Phuoc Tran (B), and Yinpeng Zhang (C).	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896578/files/figure.png	https://doi.org/10.5281/zenodo.16896578	Figure 10. Topotype of Tylototriton ziegleri (ZMMU A-8167, male) Thanh Van Village, Quang Ba, Bat Dai Son Nature Reserve, Ha Giang Province, Vietnam, in life. A, dorsal view; B, ventral view. Photographs by Nikolay A. Poyarkov.	Figure 10. Topotype of Tylototriton ziegleri (ZMMU A-8167, male) Thanh Van Village, Quang Ba, Bat Dai Son Nature Reserve, Ha Giang Province, Vietnam, in life. A, dorsal view; B, ventral view. Photographs by Nikolay A. Poyarkov.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896558/files/figure.png	https://doi.org/10.5281/zenodo.16896558	Figure 1. Distribution of Tylototriton koliaensis sp. nov. (red), T. ziegleri s. str. (blue) and T. thaiorum (green) in northern Indochina. A, map of Indochina, the inset shows the area of the studied species. B, localities of the studied species; see Table 1 for locality details. Dots show type localities. Locality 4 was not confirmed with molecular data and should be regarded as Tylototriton cf. koliaensis sp. nov.	Figure 1. Distribution of Tylototriton koliaensis sp. nov. (red), T. ziegleri s. str. (blue) and T. thaiorum (green) in northern Indochina. A, map of Indochina, the inset shows the area of the studied species. B, localities of the studied species; see Table 1 for locality details. Dots show type localities. Locality 4 was not confirmed with molecular data and should be regarded as Tylototriton cf. koliaensis sp. nov.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
03CF87D8FFA3FFC0D22CFEEC9298FE6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16896580/files/figure.png	https://doi.org/10.5281/zenodo.16896580	Figure 11. The type locality of Tylototriton koliaensis sp. nov. in Phia Oac-Phia Den National Park, Cao Bang Province, Vietnam. A, macrohabitat. B, microhabitat. Photographs by Son Xuan Le.	Figure 11. The type locality of Tylototriton koliaensis sp. nov. in Phia Oac-Phia Den National Park, Cao Bang Province, Vietnam. A, macrohabitat. B, microhabitat. Photographs by Son Xuan Le.	2024-02-19	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan		Zenodo	biologists	Poyarkov, Nikolay A.;Nguyen, Tan Van;Le, Son Xuan;Le, Dac Xuan			
