identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CC0156FFF2FF8346DB16940B9F15EA.text	03CC0156FFF2FF8346DB16940B9F15EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arius maculatus	<div><p>Arius maculatus (n= 12) (Fig. 5, Supplementary data 5)</p><p>These lapilli have three pronounced angular-like points at the postero-mesial, antero-mesial, and antero-distal margins, making them appear triangular. Their antero-mesial projection (Amp) is subtle, and they have compact curved sulculus lapilli (Sl). Their mesial dents (Md) (the “valleys” along the outer rim of the mesial margin) are subtle, their mesial notch (Mn) is compressed, and they have deep incisura linea basalis (ILb).</p></div>	https://treatment.plazi.org/id/03CC0156FFF2FF8346DB16940B9F15EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF2FF8346DB15BC0EE9143E.text	03CC0156FFF2FF8346DB15BC0EE9143E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arius oetik	<div><p>Arius oetik (n= 7) (Fig. 5, Supplementary data 5)</p><p>These otoliths also appear triangular and generally quite indistinguishable from A. maculatus . Though, their Md appears to be more pronounced. They have subtle Amp, and compact curved Sl. Their mesial notch is compressed, and they bear shallow ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF2FF8346DB15BC0EE9143E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF2FF8346DB140F0EA71489.text	03CC0156FFF2FF8346DB140F0EA71489.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arius venosus	<div><p>Arius venosus (n= 28) (Fig. 5, Supplementary data 5)</p><p>The appearance of the otoliths is very similar to that of A. oetik and there are no obvious distinguishable features between the two species others than some of the A. venosus otoliths (n= 3) have deep bent ILb, while the others (n= 25) have shallow ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF2FF8346DB140F0EA71489	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB10AF0EDA1729.text	03CC0156FFF1FF8046DB10AF0EDA1729.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrachocephalus mino	<div><p>Batrachocephalus mino (n= 1) (Fig. 5, Supplementary data 5)</p><p>The otolith appears to be triangular, but it has the most compact/shortest antero-distal margin among the “triangular” shaped taxa, making it seem almost like an arrowhead. Its Amp is quite prominent. It has compact curved Sl. Its Mn is compact. It has shallow ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF1FF8046DB10AF0EDA1729	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB17F30C841641.text	03CC0156FFF1FF8046DB17F30C841641.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptarius truncatus	<div><p>Cryptarius truncatus (n= 22) (Fig. 5, Supplementary data 6)</p><p>The mesial shallow depression of these lapilli appears to be the widest out of all the taxa. Most of them have noticeable Md, particularly at the middle section of their mesial margin rim. Their mesial shallow depression (Msd) appears to be quite thick. Their Amp is quite prominent, and some (n= 12) have compact curved Sl, while the others (n= 10) have tubular curved Sl. They have compact Mn, and deep ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF1FF8046DB17F30C841641	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB15DF0B3F143D.text	03CC0156FFF1FF8046DB15DF0B3F143D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexanematichthys sagor	<div><p>Hexanematichthys sagor (n= 14) (Fig. 6, Supplementary data 7)</p><p>Except for the antero-mesial margin of these otoliths, the mesial, posterior, and distal margins are well-rounded and convex. This results in a wide distal area and distal margin, and overall, makes the otoliths look circular shaped. They have wide Msd, and prominent Amp, and most (n= 8) show tubular curved Sl, while others (n= 6) have compact curved Sl. Their Mn is very prominent, and they have deeply bent ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF1FF8046DB15DF0B3F143D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB161B0C4E1695.text	03CC0156FFF1FF8046DB161B0C4E1695.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kyataphisa nenga	<div><p>Kyataphisa nenga (n= 6) (Fig. 5, Supplementary data 6)</p><p>The posterior margin of these otoliths can either be sharply or smoothly curved, but either way, they generally appear oval or egg-shaped. Their Amp is quite prominent, majority (n= 5) have compact curved Sl, with only one having tubular curved Sl. They have compact Mn and deep ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF1FF8046DB161B0C4E1695	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8746DB14060B32138A.text	03CC0156FFF1FF8746DB14060B32138A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Netuma bilineata	<div><p>Netuma bilineata (n= 24) (Fig. 6, Supplementary data 7)</p><p>Like H. sagor, the curvature of most of the otoliths’ margin making them appear circular. Although, the outermost part of their mesial margin is quite irregular, especially along the postero-mesial margin. Some of them (n= 8) have prominent Md, which causes a sharp slant that makes them appear “swollen” at the postero-mesial margin. They have prominent Amp and the majority of them (n= 19) bear tubular straight Sl, while the remaining ones (n= 5) have compact straight Sl. They have prominent mesial notch and their ILb is shallow.</p></div>	https://treatment.plazi.org/id/03CC0156FFF1FF8746DB14060B32138A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB156F0BEE1505.text	03CC0156FFF1FF8046DB156F0BEE1505.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Osteogeneiosus militaris	<div><p>Osteogeneiosus militaris (n= 14) (Fig. 6, Supplementary data 6)</p><p>These otoliths lack curvature in their general appearance, making them look rectangular-shaped. Their Amp is quite small, and they have compact curved Sl. Their Mn is very compact, and they have shallow ILb.</p></div>	https://treatment.plazi.org/id/03CC0156FFF1FF8046DB156F0BEE1505	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF6FF8746DB125C0A4F12DE.text	03CC0156FFF6FF8746DB125C0A4F12DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Netuma thalassina	<div><p>Netuma thalassina (n= 16) (Fig. 6, Supplementary data 7)</p><p>These otoliths share a majority of the N. bilineata otoliths’ features, including having prominent Amp, Mn, and shallow ILb. This makes the two Netuma species quite indistinguishable from each other. However here, some N. thalassina otoliths tend to have compact straight Sl (n= 5), and most of them have compact curved Sl (n= 11).</p></div>	https://treatment.plazi.org/id/03CC0156FFF6FF8746DB125C0A4F12DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF6FF8746DB12A10A7D11F6.text	03CC0156FFF6FF8746DB12A10A7D11F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plicofollis argyropleuron	<div><p>Plicofollis argyropleuron (n= 39) (Fig. 6, Supplementary data 8)</p><p>The opaque area towards their distal margin of the otoliths appears to be narrower than that of the Hexanematichthys and Netuma genera. With the addition of their mesial margin being symmetrically convex, the otoliths appear rather diamond shaped. Their Amp is prominent, and they have tubular curved Sl. They also have compressed Mn along with deep ILb. There is very minimal variation within the samples and their appearance seem to be constant.</p></div>	https://treatment.plazi.org/id/03CC0156FFF6FF8746DB12A10A7D11F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF6FF8746DB11C90FD810D6.text	03CC0156FFF6FF8746DB11C90FD810D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plicofollis nella	<div><p>Plicofollis nella (n= 33) (Fig. 6, Supplementary data 8)</p><p>These lapilli are quite similar to the P. argyropleuron ’s, in terms of the features of their mesial and distal margins, that makes them also appear diamond shaped. However, here some of the otoliths tend to have small protrusions on the outermost part of their mesial margin, typically near to the middle but slightly towards the anterior margin. They have prominent Amp and some of them have tubular curved Sl (n= 22), and others are compactly curved (n= 11). Their Mn appears slightly compressed, and a few have shallow ILb (n= 13), while the remaining have deep ILb (n= 20).</p></div>	https://treatment.plazi.org/id/03CC0156FFF6FF8746DB11C90FD810D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Johari, Adibah;Kocsis, László;Roslim, Amajida;Haziq, Ilman	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
