taxonID	type	description	language	source
03CC0156FFF2FF8346DB16940B9F15EA.taxon	description	These lapilli have three pronounced angular-like points at the postero-mesial, antero-mesial, and antero-distal margins, making them appear triangular. Their antero-mesial projection (Amp) is subtle, and they have compact curved sulculus lapilli (Sl). Their mesial dents (Md) (the “ valleys ” along the outer rim of the mesial margin) are subtle, their mesial notch (Mn) is compressed, and they have deep incisura linea basalis (ILb).	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF2FF8346DB15BC0EE9143E.taxon	description	These otoliths also appear triangular and generally quite indistinguishable from A. maculatus. Though, their Md appears to be more pronounced. They have subtle Amp, and compact curved Sl. Their mesial notch is compressed, and they bear shallow ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF2FF8346DB140F0EA71489.taxon	description	The appearance of the otoliths is very similar to that of A. oetik and there are no obvious distinguishable features between the two species others than some of the A. venosus otoliths (n = 3) have deep bent ILb, while the others (n = 25) have shallow ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB10AF0EDA1729.taxon	description	The otolith appears to be triangular, but it has the most compact / shortest antero-distal margin among the “ triangular ” shaped taxa, making it seem almost like an arrowhead. Its Amp is quite prominent. It has compact curved Sl. Its Mn is compact. It has shallow ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB17F30C841641.taxon	description	The mesial shallow depression of these lapilli appears to be the widest out of all the taxa. Most of them have noticeable Md, particularly at the middle section of their mesial margin rim. Their mesial shallow depression (Msd) appears to be quite thick. Their Amp is quite prominent, and some (n = 12) have compact curved Sl, while the others (n = 10) have tubular curved Sl. They have compact Mn, and deep ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB15DF0B3F143D.taxon	description	Except for the antero-mesial margin of these otoliths, the mesial, posterior, and distal margins are well-rounded and convex. This results in a wide distal area and distal margin, and overall, makes the otoliths look circular shaped. They have wide Msd, and prominent Amp, and most (n = 8) show tubular curved Sl, while others (n = 6) have compact curved Sl. Their Mn is very prominent, and they have deeply bent ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB161B0C4E1695.taxon	description	The posterior margin of these otoliths can either be sharply or smoothly curved, but either way, they generally appear oval or egg-shaped. Their Amp is quite prominent, majority (n = 5) have compact curved Sl, with only one having tubular curved Sl. They have compact Mn and deep ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8746DB14060B32138A.taxon	description	Like H. sagor, the curvature of most of the otoliths’ margin making them appear circular. Although, the outermost part of their mesial margin is quite irregular, especially along the postero-mesial margin. Some of them (n = 8) have prominent Md, which causes a sharp slant that makes them appear “ swollen ” at the postero-mesial margin. They have prominent Amp and the majority of them (n = 19) bear tubular straight Sl, while the remaining ones (n = 5) have compact straight Sl. They have prominent mesial notch and their ILb is shallow.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF1FF8046DB156F0BEE1505.taxon	description	These otoliths lack curvature in their general appearance, making them look rectangular-shaped. Their Amp is quite small, and they have compact curved Sl. Their Mn is very compact, and they have shallow ILb.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF6FF8746DB125C0A4F12DE.taxon	description	These otoliths share a majority of the N. bilineata otoliths’ features, including having prominent Amp, Mn, and shallow ILb. This makes the two Netuma species quite indistinguishable from each other. However here, some N. thalassina otoliths tend to have compact straight Sl (n = 5), and most of them have compact curved Sl (n = 11).	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF6FF8746DB12A10A7D11F6.taxon	description	The opaque area towards their distal margin of the otoliths appears to be narrower than that of the Hexanematichthys and Netuma genera. With the addition of their mesial margin being symmetrically convex, the otoliths appear rather diamond shaped. Their Amp is prominent, and they have tubular curved Sl. They also have compressed Mn along with deep ILb. There is very minimal variation within the samples and their appearance seem to be constant.	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
03CC0156FFF6FF8746DB11C90FD810D6.taxon	description	These lapilli are quite similar to the P. argyropleuron ’ s, in terms of the features of their mesial and distal margins, that makes them also appear diamond shaped. However, here some of the otoliths tend to have small protrusions on the outermost part of their mesial margin, typically near to the middle but slightly towards the anterior margin. They have prominent Amp and some of them have tubular curved Sl (n = 22), and others are compactly curved (n = 11). Their Mn appears slightly compressed, and a few have shallow ILb (n = 13), while the remaining have deep ILb (n = 20).	en	Johari, Adibah, Kocsis, László, Roslim, Amajida, Haziq, Ilman (2025): Otolith morphometric analysis of North Bornean Ariidae (Siluriformes): new insights and taxonomic implications. Zootaxa 5660 (4): 451-474, DOI: 10.11646/zootaxa.5660.4.1, URL: https://doi.org/10.11646/zootaxa.5660.4.1
