taxonID	type	description	language	source
03D78450FFE2D5738A1250C7C0F77942.taxon	materials_examined	Material examined: 28 animals and 15 eggs. Specimens mounted on microscope slides in Hoyer’s medium (24 animals + 10 eggs), fixed on SEM stubs (0 + 5), and processed for DNA sequencing (4 + 0). Type locality: 44 ° 02 ' N, 114 ° 49 ' E; 100 m asl: Malaysia: Sarawak, Borneo, Gunung Mulu; epiphyllous moss on the tree leaf in the primary tropical forest; coll. 27 July 2016 by Piotr Gąsiorek. Type depositories: Holotype (slide MY. 098.01 with 4 paratypes) and 19 paratypes (slides: MY. 098. *, where the asterisk can be substituted by any of the following numbers 02, 04 – 05) and 15 eggs (slides: MY. 098. *: 03, 06; SEM stub: 18.13) are deposited at the Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Kraków, Poland. Etymology: We take great pleasure in dedicating this new species to Filip Dudziak, son of the second author. Description: Animals (measurements and statistics in Table 4): Before mounting in Hoyer’s medium, body almost transparent in juveniles and white in adults, eyes absent; after fixation on microscope slides body transparent (Fig. 9 A). Body cuticle covered with fine granulation clearly visible on the dorsal side of the caudal body region (Fig. 9 B). On legs I – III, a patch of fine granulation is placed on the external surface of the legs, near the claws (Fig. 9 C), whereas granulation on legs IV extends from the claws onto the entire dorsolateral surface of the legs, being denser towards the claws (Fig. 9 E). A pulvinus is present on the internal surface of legs I – III (Fig. 9 D). Claws slender, of the hufelandi type. Primary branches with distinct accessory points, a long common tract, and with an evident stalk connecting the claw to the lunula (Fig. 10 A – B). Lunulae on all legs smooth (Fig. 10 A – B). Bars under claws absent. Mouth antero-ventral, bucco-pharyngeal apparatus of the Macrobiotus type (Fig. 11 A). The oral cavity armature well-developed and composed of three bands of teeth (Fig. 11 B – E). The first band of teeth comprises numerous small granules arranged in a several rows situated anteriorly in the oral cavity, just behind the bases of the peribuccal lamellae (Fig. 11 B – E). The second band of teeth is situated between the ring fold and the third band of teeth, and is composed of ridges parallel to the main axis of the buccal tube (Fig. 11 B – E). The teeth of the third band are located within the posterior portion of the oral cavity, between the second band of teeth and the buccal tube opening (Fig. 11 B – E). The third band of teeth is divided into the dorsal and the ventral portion. Under PCM, both dorsal and ventral teeth are visible as two lateral and one median transverse ridges (Fig. 11 B – E). The ventro-median tooth is divided into two roundish teeth of which one is sometimes larger (Fig. 11 C, E). Pharyngeal bulb spherical, with triangular apophyses, three rod-shaped macroplacoids and a microplacoid clearly distant (more than its length) from the third macroplacoid (Fig. 11 A, F – G). The macroplacoid length sequence is 2 <1 <3. The first macroplacoid is anteriorly narrowed and the third has a subterminal constriction (Fig. 11 F – G). Eggs (measurements and statistics in Table 5): Laid freely, white, spherical with conical processes with the elongated terminal portion terminated with a small concave disc with an irregular edge (Figs. 12 A – F, 13 A – F). The labyrinthine layer between the process walls is visible under PCM as a reticular pattern with slightly sinuous margins (Fig. 12 A – B). Eight to ten areoles are present around each process (Figs. 12 A – B, 13 A – B). The surface of the areoles is sculptured and porous (Figs. 12 A – B, 13 A – D). Pores large and visible both under PCM and SEM (Figs. 12 A – B, 13 A – D, respectively). The ridges separating each areole are narrower than the areole diameter (Figs. 12 A – B, 13 A – D). Reproduction: The new species is probably parthenogenetic since no spermathecae or testis filled with spermatozoa were found in specimens freshly mounted in Hoyer’s medium. DNA sequences: We obtained sequences for three out of the four DNA markers which we had tried to sequences. We did not get the ITS- 2 sequences for the species as the reads were always of bad quality. Out of these three successfully sequenced markers 18 S rRNA and 28 S rRNA was represented by a single haplotype, whereas COI was represented by two haplotypes: The 18 S rRNA sequence (GenBank: MT 261913), 1017 bp long; The 28 S rRNA sequence (GenBank: MT 261904), 780 bp long; The COI haplotype 1 sequence (GenBank: MT 260372), 658 bp long; COI haplotype 2 sequence (GenBank: MT 260373), 658 bp long. Remarks: Paramacrobiotus filipi sp. nov. is only the fourth species reported from Malaysia and the fourth specifically from Borneo (Pilato et al. 2004; Gąsiorek 2018; Gąsiorek et al. 2020; Gąsiorek and Michalczyk 2020).	en	Stec, Daniel, Dudziak, Magdalena, Michalczyk, Łukasz (2020): Integrative Descriptions of Two New Macrobiotidae Species (Tardigrada: Eutardigrada: Macrobiotoidea) from French Guiana and Malaysian Borneo. Zoological Studies (Zool. Stud.) 59 (23): 1-25, DOI: 10.6620/ZS.2020.59-23, URL: http://dx.doi.org/10.5281/zenodo.12822655
03D78450FFEAD571898750C7C73979C2.taxon	materials_examined	Material examined: Two slides (TH. 001.01 and TH. 001.02) with 1 paratype and 6 eggs from the type series mounted in Faure medium (these slides are now deposited at the Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Kraków, Poland). PCM photomicrographs of the holotype and another paratype as well as two eggs from the type series. Amended description of the species: According to the original description, the granulation is absent on the first three pairs of legs and lunules IV are faintly dentate. However, our re-examination of the type material revealed the presence of faint granulation present on the external surface of legs I – III in larger animals (Fig. 14 A) whereas in smaller specimens the granulation can be hardly or even not visible. Moreover, we found that lunules on all the legs are smooth (Fig. 14 A – B). The original description also states that indistinct reticular sculpture is present within the areolae. We confirmed that the areolae surface is sculptured, however only wrinkles are present whereas reticulation or pores are absent or not visible under PCM (Fig. 14 C – D). We also confirmed multiple divisions of the medio-ventral tooth in the third band of teeth into several roundish teeth (Fig. 14 E – F) and the absence or invisibility of the body granulation under PCM.	en	Stec, Daniel, Dudziak, Magdalena, Michalczyk, Łukasz (2020): Integrative Descriptions of Two New Macrobiotidae Species (Tardigrada: Eutardigrada: Macrobiotoidea) from French Guiana and Malaysian Borneo. Zoological Studies (Zool. Stud.) 59 (23): 1-25, DOI: 10.6620/ZS.2020.59-23, URL: http://dx.doi.org/10.5281/zenodo.12822655
