taxonID	type	description	language	source
03D4D13945108149FDBCFDF53075394E.taxon	discussion	Remark Subgeneric characters given by Barkalov (1983) are marked by an circumflex accent (^).	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D13945108149FDBCFDF53075394E.taxon	diagnosis	Diagnosis Eye pilose ^; frons swollen [especially so in the ♂] ^; antennal fossa separated by a median process of lunule ^; face haired only near eye marginv or bare; face and frons exceptionally wide ^ [in most of the species]; ♀ frons: inner margin of lateral channel strongly arched anteriorly; anterior portion of scutum often with a pair of submedian pruinose vittae [non-vittate in the ♀ of C. barovskii]; dorsal and ventral katepisternal hair patch at least posteriorly confluent or only narrowly separated on lower one third of sclerite; vein M 1 meeting vein R 4 + 5 at an acute angle; membrane of wing beyond the veins dm-cu and M 1 narrow; sclerites of distiphallus without ventral projections ^; dorsal lobe of gonostylus rudimentary or missing; right and left surstyli and gonostyli (slightly) asymmetrical.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D13945108146FDB6FBEE30EF3B5A.taxon	description	Figs 1, 5 C, H	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D13945108146FDB6FBEE30EF3B5A.taxon	materials_examined	Material examined FINLAND • 1 ♀; biogeographical province of Southern Häme [Etelä Häme EH], Valkeakoski, Sääksmäki; 15 Jun. 1932; E. Kivirikko leg.; http: // tun. fi / GAVK. 2281; JKA • 1 ♀; Tavastia australis [Etelä Häme EH], Valkeakoski, Sääksmäki; 27 Jun. 1935; E. Kivirikko leg.; http: // tun. fi / GAVK. 2280; JKA • 1 ♀; Southern Häme [Etelä Häme EH], Valkeakoski, Sääksmäki; 30 May 1937; E. Kivirikko leg.; http: // tun. fi / JX. 925264; MZH • 1 ♀ *; Uusimaa, Sipoo; 20 May 2013; T. Järveläinen leg.; TJA. RUSSIA • 1 ♀; Leningradskii oblast, Luschkij raion, Tolmatshevo; 18 May 1936; [? Stackelberg]; ZISP • 1 ♂ *; Khabarovskij krai, Lake Amut; 50.81059 ° N, 136.40042 ° E; 735 m a. s. l.; 11 Jun. 2013; J. H. Skevington leg.; CNC Diptera 229791.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D13945108146FDB6FBEE30EF3B5A.taxon	description	The specimens used for molecular analysis are indicated with an * (see Table 2). Description Male Although only the female of C. barovskii has been described, the molecular evidence presented here links the described female sex with a male specimen, thus confirming its identity, allowing us to here provide the description of the male sex of C. barovskii, based on a single specimen collected in the Russian Far East. LENGTH. Body 7 mm; wing 6.5 mm. HEAD. Eye brownish pilose, hairs on dorsal part of eye about as long as half width of 3 rd antennal segment. Anterior eye angle ca 90 °. Frons slightly swollen, with a shallow median furrow, shiny on anterior one-third, pruinose on posterior two thirds, with long, erect black hairs. Ocellar triangle about equilateral, long black pilose. Gena wide, viewed obliquely from below about as wide or slightly wider than oral cavity. Face not wide, ratio of width of face at level of facial tubercle to maximum width of head = 0.44: 1; face shiny, very faintly pruinose, with a broad fascia of dense pruinosity just below base of antennae. Sides of face bare, without hairs. Parafacia narrow, at level of facial tubercle about one third as wide as 3 rd antennal segment, fully developed only lateral facial tubercle, above level of facial tubercle compressed toward eye margin by a bulge, with black hairs, hairs in upper part slightly longer than in lower part. Lunule dark-brown to black. Antenna with scape, pedicel and 3 rd antennal segment black to dark brown, 3 rd antennal segment about as long as wide. Arista black, with very short hairs. THORAX. Scutum centrally faintly pruinose, anterior half with a pair of grey pruinose vittae, best visible if viewed from behind, laterally almost shiny; scutal hairs long black, erect, of about even length; hairs on scutellum similar to those on scutum, with marginal setae. Sub-scutellar fringe black and long. Pleurae long black pilose on posterior anepisternum, anepimeron and on katepisternum, dorsal and ventral katepisternal hair patches confluent. WING. Membrane entirely microtrichose, hyaline; veins in basal half bright yellow, in apical half dark, basal half of RS without long dark setulae; M 1 meeting R 4 + 5 at an acute angle; calypter whitish to yellowish, with orange or brownish rim, marginal fringe on dorsal lobe partly with brown setulate. Haltere with brownish stem and yellow capitulum. LEGS. Legs black, brownish-grey pruinose, but basal 1 / 4 of protibia, 1 / 7 of mesotibia, 1 / 10 of metatibia and extreme apices of femora usually reddish-yellow; hairs on femora nearly all black, but with a set of yellow hairs posteriorly on metafemur; ventral surface of metafemur with black setulate on basal 2 / 3, apical 1 / 3 with a set of antero-ventral black hairs. Tibiae mixed black and yellow pilose. Tarsi short black pilose or mixed black and reddish-brown pilose dorsally, ventral surface of pro- and metatarsi usually reddish-brown pilose, or intermixed with black hairs. Tarsi of mesoleg ventrally with black setulae. ABDOMEN. Tergites medially brownish pruinose (best seen if viewed obliquely from front), shiny on sides of tergites 2 – 4; tergites 1 – 3 with long mixed black and yellow hairs on disc, sides of tergite 1 and anterolateral corners of tergite 2 with long yellow hairs, remainder of lateral margin of tergite 2 and entire margin of tergite 3 with black hairs, tergite 4 with black hairs. Sternites faintly greyish-brown pruinose; sternite 1 yellow pilose; base and sides of sternite 2 with long mixed black and yellow hairs, sides of sternites 3 – 4 long black pilose, hairs on the median parts of sternites 2 – 4 black and appressed. GENITALIA. Right and left surstyli and gonostyli slightly asymmetrical; ventral margin of surstylus with a shallow basal convexity; hypandrium in lateral view about 1.2 times as long as wide; sclerite of distiphallus broadly fused dorsally, with long dorsal lobes. Female LENGTH. Body 6 – 7 mm; wing 5.4 – 6.2 mm. The ♀ differs from the ♂ in the following characters: integument more shiny; pilosity much shorter. HEAD. Eye hairs short, about ¼ or less of width of postpedicel. Frons not wide, at level of lunule about 0.3 times as wide as maximum width of head, with a shallow transverse sulcus at level of about lower 1 / 3 of frons; lateral channels narrow, widening from inner dorsal corner of eye towards transverse sulcus, enclosing a small semicircular area of dense grey pruinosity at about median 1 / 3 of frons, remainder of frons shiny; hairs on frons erect, as long as one third width of postpedicel, black. Occiput dorsally much wider than in male, grey pruinose, with short black hairs. Gena long yellow pilose. Face bare, at level of facial tubercle about 0.3 times as wide as maximum width of head, shiny; parafacia orange on dorsal half, at level of tentorial groove about half as wide as diameter of protibia, pale pruinose and with short yellow hairs. Lunule orange. Scape, pedicel and 3 rd antennal segment bright orange, 3 rd antennal segment larger than in male. THORAX. Scutum and scutellum shiny, pale pruinose sub-median vittae anterior on scutum reduced to spots; scutum with semi-erect short black hairs. Pleurae faintly grey pruinose, hairs yellow and black, shorter and less dense than in male. Calypter whitish, with orange rim and yellow fringe. Haltere with brownish or yellowish stem and yellow knob. LEGS. Legs black, but basal 1 / 4 of protibia, 1 / 3 of mesotibia and basal 1 / 10 of metatibia, and often extreme apices of tibiae and femora reddish-yellow; femora with both black and yellow hairs; ventral surface of metafemur basally without black setulae; tibial hairs and tarsi as in male. ABDOMEN. Tergites with less dense pruinosity compared to male, mainly with black hairs, except on tergite 1 and in anterolateral corners of tergite 2 with yellow hairs. Sternites lightly pruinose, short black and yellow pilose; hairs erect on sternites 1 – 2 and on anterior corners of sternite 3, remaining hairs more or less appressed.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D13945108146FDB6FBEE30EF3B5A.taxon	distribution	Distribution Finland, Russia.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D13945108146FDB6FBEE30EF3B5A.taxon	diagnosis	Differential diagnosis Similar in appearance to extralimital Cheilosia komabaensis, see Barkalov & Ichige (2016) for a differential diagnosis with that species. Similar in appearance to Cheilosia luteicornis and C. morio but differs as follows: body length shorter. Face narrower. Basal half of wing with yellow veins. Legs with less extensive yellowish parts. ♂: anterior eye angle smaller. Anterior part of frons shiny. Haltere knob yellow. Lateral margin of tergite 1 and anterolateral corner of tergite 2 with yellow hairs. Surstylus basally narrower than in C. morio, and shorter than in C. luteicornis. ♀: scape and pedicel orange. Postocular orbit dorsally with black hairs. Scutum with semi-erect black hairs. Tergites 3 – 4 with black hairs. Moreover, both sexes differ from C. morio by having sides of face not haired and from C. luteicornis by having parafacia above posterior tentorial pit strongly compressed.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	description	Figs 2, 5 A, D, F	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	materials_examined	Type localities: “ Juckasjarvi; Pello, Lapponiae Tornensis; Lycksele Lapponiae Umensis ” [Juckasjärvi and Lycksele in Swedish Lapland, and Pello in Finnish Lapland] (lectotype, selected by Claussen, designated here, ♀, MZLU).	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	materials_examined	Material examined FINLAND – Åland • 1 ♀; Åland, Lemland, Flaka; 60.006127 ° N, 20.138916 ° E; 20 May 2012; Raekunnas leg.; http: // id. luomus. fi / HT. 3488; MZH. – Pohjois-Pohjanmaa • 1 ♀; Rovaniemi; 66.27 ° N, 25.11 ° E; 19 June 1951; Stenberg leg.; http: // id. luomus. fi / GV. 93913; MZH. – South Hame • 1 ♀; Hattula; 61.03 ° N, 24.27 ° E; 1906; Wegelius leg.; http: // id. luomus. fi / GV. 93924; MZH 1 ♀; Forssa; 60.85 ° N, 23.62 ° E; 12 May 1962; Nylund leg.; http: // id. luomus. fi / GV. 93910; MZH • 1 ♀; Forssa; 60.85 ° N, 23.62 ° E; 14 May 1964; Käpylä leg.; http: // id. luomus. fi / GV. 93920; MZH • 1 ♂; Somero; 60.63 ° N, 23.46 ° E; 21 May 1964; Maaniitty leg.; http: // id. luomus. fi / GV. 93916; MZH. – South Karelia • 1 ♂; Kotka, Mussalo; 60.455875 ° N, 26.906058 ° E; 17 May 1918; Ulvinen leg.; http: // id. luomus. fi / GV. 23565; MZH • 1 ♂; Hamina; 60.67577 ° N, 27.079224 ° E; 10 May 1964; Tiensuu leg.; http: // id. luomus. fi / GV. 93911; MZH • 1 ♀; same data as for preceding; http: // id. luomus. fi / GV. 93921. – Uusimaa • 2 ♀♀; Loviisa; 60.44 ° N, 26.03 ° E; Nordström leg.; http: // id. luomus. fi / GV. 93922, http: // id. luomus. fi / GV. 93915; MZH • 1 ♀; Helsinki; 60.191 ° N, 24.875 ° E; Johansson leg.; http: // id. luomus. fi / GV. 93912; MZH • 1 ♀; Helsinki; 60.19 ° N, 25.02 ° E; Tuomikoski leg.; http: // id. luomus. fi / GV. 93927; MZH • 1 ♂; Helsinki; 60.19 ° N, 25.02 ° E; Tiensuu leg.; http: // id. luomus. fi / GV. 93909; MZH • 1 ♀; Vantaa; 60.27 ° N, 24.96 ° E; 1 Apr. – 31 May 1906; Frey leg.; http: // id. luomus. fi / GV. 93907; MZH • 1 ♂; Espoo, Bodom träsk; 60.256 ° N, 24.665 ° E; 16 May 1932; Frey leg.; http: // id. luomus. fi / GV. 93923; MZH • 1 ♀; Helsinki, Munksnäs; 60.198 ° N, 24.874 ° E; 6 May 1937; Hellén leg.; http: // id. luomus. fi / GV. 93917; MZH • 1 ♀; Helsinki; 60.191 ° N, 24.875 ° E; 25 May 1942; Frey leg.; http: // id. luomus. fi / GV. 93926; MZH • 2 ♀♀; Loviisa, Vahterpää; 60.369558 ° N, 26.462275 ° E; 10 May 1975; Albrecht leg.; http: // id. luomus. fi / GV. 23567, http: // id. luomus. fi / GV. 23566; MZH • 1 ♀; Loviisa, Vahterpää; 60.369327 ° N, 26.407907 ° E; 4 May 1975; Albrecht leg.; http: // id. luomus. fi / GV. 23568; MZH. – Varsinais-Suomi • 1 ♀; Varsinais-Suomi, Eriksberg; 60.374 ° N, 23.296 ° E; von Bonsdorff leg.; http: // id. luomus. fi / GV. 93914; MZH • 1 ♀; Lohja; 60.26 ° N, 24.01 ° E; 6 May 1918; Lindberg leg.; http: // id. luomus. fi / GV. 93918; MZH • 1 ♀; Salo; 60.189835 ° N, 22.876551 ° E; 17 May 1944; Hellman leg.; http: // id. luomus. fi / GV. 21350; MZH. GERMANY – Baden-Württemberg • 1 ♀; Baden-Württemberg, Schwarzwald, Rinken; 15 May 1992; Stuke leg.; CNC Diptera 101716 • 1 ♀; Baden-Württemberg, Schwarzwald, SE of Todtnauer Hütte; 15 May 1992; Stuke leg.; CNC Diptera 101715. – Lower Saxony • 1 ♀; Lopautal; 12 Apr. 1991; Kassebeer leg.; CNC Diptera 101818. NETHERLANDS – Drenthe • 1 ♂, 1 ♀; Drouwenerveld; 52.957 ° N, 6.766 ° E; 21 Apr. 2016; Bot leg.; SBA • 1 ♂, 2 ♀♀; Drouwenerveld; 52.957 ° N, 6.765 ° E; 6 Apr. 2019; Bot leg.; SBA • 1 ♀; Drouwenerveld; 52.957 ° N, 6.765 ° E; 17 Apr. 2019; Bot leg.; SBA • 1 ♂; Drouwenerveld; 52.957 ° N, 6.763 ° E; 5 Apr. 2020; Bot leg.; SBA • 1 ♂; Drouwenerveld; 52.952 ° N, 6.770 ° E; 23 Mar. 2022; Bot leg.; SBA. The molecular specimens are listed in Table 2.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	description	Description Male LENGTH. Body 9.1 – 12.0 mm, wing 8.1 – 9.1 mm. HEAD. Eye with brownish hairs. Anterior eye angle ca 110 ° – 120 °. Frons swollen, with a shallow median furrow, greyish-brown pruinose, with long, erect black hairs. Ocellar triangle about equilateral, with long black hairs. Gena wide, viewed obliquely from below about as wide or slightly wider than oral cavity. Face wide, ratio of width of face at level of facial tubercle to maximum width of head = 0.53 – 0.58: 1; face faintly pruinose, with a broad fascia of dense pruinosity just below base of antennae. Sides of face not haired. Parafacia at level of facial tubercle about half as wide as 3 rd antennal segment, with short black hairs. Facial surface at sides just above posterior tentorial pits weakly convex. Lunule dark-brown to black. Antenna with scape and pedicel usually black, 3 rd antennal segment usually reddish, but partly to entirely brownish to black in some specimens. Arista usually black, practically bare. THORAX. Scutum faintly pruinose, anterior half with a pair of pale pruinose vittae, best visible if viewed from behind; in well preserved specimens two less distinct lateral vittae are perceptible; scutal hairs long black, erect, of about even length; hairs of scutellum similar to those of scutum, scutellum without distinct marginal setae, but occasionally with some marginal hairs slightly stronger than on disc. Subscutellar fringe black and long. Pleurae with long black hairs on posterior anepisternum, anepimeron and on katepisternum, dorsal and ventral katepisternal hair patches confluent. WING. Membrane entirely microtrichose, more or less brownish tinged, especially so in anterior half; veins dark, basal half of RS usually without long dark setulae; calypter whitish to yellowish, with orange or brownish rim, marginal fringe on dorsal lobe partly with black setulae. Haltere with brownish stem and black knob. LEGS. Legs black, but basal 1 / 3 of pro- and metatibia, basal 2 / 5 of mesotibia and extreme apices of femora reddish-yellow, rarely also extreme apices of tibiae yellow; hairs of femora nearly all black, but with a set of yellow hairs posteriorly on metafemur; ventral surface of metafemur with black setulae on basal 2 / 3, apical 1 / 3 with a set of antero-ventral black hairs. Tibiae with anterior surface and basal part of ventral surface of protibia and ventral and posterior surface of metatibia with short reddish-yellow hairs, tibiae otherwise with mixed black and yellow hairs. Tarsi with short black hairs or mixed black and reddish-brown haired dorsally, ventral surface of pro- and metatarsi usually reddish-brown haired, or intermixed with black hairs. Tarsi of mesolegs ventrally with black setulae. ABDOMEN. Tergites entirely brownish pruinose, densely on disc, less densely on sides of tergites 1 – 3 and on tergite 4 (best seen when viewed obliquely from front); tergites 1 – 3 long yellow pilose on disc, sides of tergites 1 – 3 and tergites 4 with long black hairs. Sternites faintly greyish-brown pruinose; sternite 1 predominantly yellow haired, often with some black hairs intermixed medially and along posterior margin; base of sternite 2 and sides of sternites 2 – 4 usually with long black hairs, hairs on median parts of sternites 2 – 4 more or less appressed, yellow and / or black to varying extents. GENITALIA. Right and left surstylus and gonostyli slightly asymmetrical; ventral margin of surstylus with a shallow basal convexity; hypandrium in lateral view about 1.4 times as long as wide; sclerite of distiphallus broadly fused dorsally, with long dorsal lobes. Female LENGTH. Body 9.5 – 12.0 mm; wing 8.0 – 9.5 mm. The ♀ differs from the ♂ in the following characters: hairs much shorter and predominantly pale yellowish-brown. HEAD. Hairs of eyes short, about 1 / 4 or less width of postpedicel. Frons exceptionally wide, at level of lunule about 0.45 times as wide as maximum width of head, with a shallow transverse sulcus at level of about lower 1 / 3 of frons, and with a more or less faint median furrow between lunule and front ocellus; lateral channels narrow, widening from inner dorsal corner of eye towards transverse sulcus, enclosing a small semicircular area of dense grey pruinosity at about median 1 / 3 of frons; dorsal portion of frons, between lateral channels, transverse sulcus and ocellar triangle and a small area on each side of antennal base more or less shining, remaining parts of frons more or less pruinose; hairs of frons erect, slightly longer than half width of 3 rd antennal segment, varying in colour from predominantly yellow to predominantly black. Occiput dorsally much wider than in male, faintly grey pruinose, with short yellow hairs, or with a few black setulae intermixed dorso-laterally. Gena with long yellow hairs. Face not haired, at level of facial tubercle almost 0.5 times as wide as maximum width of head, faintly grey pruinose; parafacia at level of facial tubercle about as wide as diameter of protibia, pale pruinose and with short yellow hairs. 3 rd antennal segment larger than in male, most often bright orange, or with a black apico-dorsal rim. THORAX. Scutum and scutellum less densely pruinose, but with pale pruinose sub-median vittae anterior on scutum, usually distinct; hairs clearly unequal in length, erect, predominantly yellow, intermixed with scattered, somewhat longer black hairs; scutellum usually with 6 – 10 black marginal setae, sub-scutellar fringe short and yellow. Pleurae grey pruinose, with yellow hairs, but hairs shorter and less dense than in male. Calypter whitish, with orange rim and yellow fringe. Haltere with brownish stem and yellowish capitulum. LEGS. Legs black, but basal 1 / 3 of pro- and metatibia, and basal 2 / 5 of mesotibia, and often extreme apices of tibiae and femora reddish-yellow; femora predominantly with yellow hairs, but apices anteriorly and dorsally with short appressed black hairs, posteriorly occasionally with a few longer, thin black setae; ventral surface of metafemur basally without black setulae. ABDOMEN. Tergites entirely but faintly grey pruinose, especially tergites 2 – 3 on disc much less pruinose than in male, hairs of tergites entirely yellow, short, erect. Sternites greyish pruinose to varying extents, from entirely matt to slightly shining, with short yellow hairs; hairs erect on sternites 1 – 2 and on anterior corners of sternite 3, remaining hairs more or less appressed.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	distribution	Distribution Austria (H. Heimburg pers. com.), Belgium, Bosnia and Herzegovina, Bulgaria, Czechia, Finland, France, Germany, Italy, Lithuania, Montenegro, Netherlands, Norway, Poland, Serbia, Slovenia, Sweden, Switzerland, Ukraine. Distribution reported here for Belgium and Switzerland were not reported before in the IUCN Red List assessment (Vujić & Likov 2021).	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	diagnosis	Differential diagnosis Very similar to Cheilosia morio, but differing as follows: sides of face not haired. The parafacia width is about half of the width of the postpedicel. Face and frons are a little less wide than in C. morio. The basal 1 / 3 – 2 / 5 of the tibiae are yellow in both sexes. Calypter is pale yellowish, fringe in the male partly dark brown. ♀ with scutal pilosity of unequal length, with numerous black hairs intermixed. ♂: abdominal tergites 2 – 3 at least partly yellow haired on disc, tergite 3 often predominantly yellow haired. Genitalia: surstylus with a shallow basal convexity; hypandrium about 1.4 times as long as wide.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D139451F8141FE7AF9FA36223CF0.taxon	discussion	Remarks Zetterstedt received the description of “ Eristl. lineata nov. spec. ♂ ” in a letter from P. Wahlberg, but he did not study the underlying specimen (s), nor other specimens of E. lineata (1843: 807 – 808). Subsequently Zetterstedt (1849: 3173, in a note under “ 31. E. melanopa ”) synonymized E. lineata under his C. morio. Among the Cheilosia materials in the Wahlberg and Boheman collections, now incorporated in the general Diptera collection of the NRM (H. Bartsch personal communication), the following historical specimens under the name Cheilosia morio could be located: 1) Wahlberg collection: 3 ♀♀ C. luteicornis, 1 ♀ C. morio; the specimens are without locality and other data but marked by a small coloured tag. These specimens cannot be syntypes, because of the female gender. 2) Boheman collection: under the name C. morio, there are 1 ♂ and 2 ♀♀ from “ Holmiae ” (= Stockholm), collected by Wahlberg and labelled by Boheman “ Hlm. / P. Wg. ”. The label of the male agrees well with the type locality and the characters of the specimen agree with those given in the original description of E. lineata Wahlberg, and it is here accepted as the lectotype.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D1394518815BFDC6FE0436313B0E.taxon	description	Figs 3 – 4, 5 B, E, G	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D1394518815BFDC6FE0436313B0E.taxon	materials_examined	Material examined FINLAND – Åland • 1 ♂; Jomala; 60.158572 ° N, 19.829543 ° E; 10 May 2005; Milankov and Ståhls leg.; http: // id. luomus. fi / GJ. 6062; MZH. – Etelä Häme • 1 ♂; Somero; 60.63 ° N, 23.46 ° E; 19 May 1962; Nurminen leg.; http: // id. luomus. fi / GV. 93934; MZH • 1 ♀; same data as for preceding; http: // id. luomus. fi / GV. 93933; MZH • 1 ♂; Urjala; 61.08 ° N, 23.49 ° E; 28 Apr. 1964; Brander leg.; http: // id. luomus. fi / GV. 93931; MZH • 1 ♀; Forssa; 60.85 ° N, 23.62 ° E; 14 May 1964; Käpylä leg.; http: // id. luomus. fi / GV. 93945; MZH • 1 ♂; Urjala; 61.08 ° N, 23.49 ° E; 30 April 1965; Brander leg.; http: // id. luomus. fi / GV. 93943; MZH. – Etelä-Savo • 1 ♂; Savonlinna; 61.781651 ° N, 29.36578 ° E; 29 May 1936; http: // id. luomus. fi / GV. 21422; MZH. – Ladoga Karelia • 3 ♂♂; Rautjärvi; 61.41 ° N, 29.39 ° E; 15 May 1944; Tiensuu leg.; http: // id. luomus. fi / GV. 93930, http: // id. luomus. fi / GV. 93938, http: // id. luomus. fi / GV. 93941; MZH. – South Karelia • 1 ♀; Summajoki, Pitkäkoski; 60.67577 ° N, 27.079224 ° E; 10 May 1964; Tiensuu leg.; http: // id. luomus. fi / GV. 93936; MZH • 1 ♂; Hamina, Leluntie; 60.558704 ° N, 27.33415 ° E; 10 May 1970; Tiensuu leg.; http: // id. luomus. fi / GV. 93929; MZH • 1 ♂; Hamina Kitula, Tupenmäki; 60.734 ° N, 27.211 ° E; 11 May 1976; Tiensuu leg.; http: // id. luomus. fi / GV. 93940; MZH. – Uusimaa • 1 ♂; Helsinki; 60.191 ° N, 24.875 ° E; Johansson leg.; http: // id. luomus. fi / GV. 93935; MZH • 1 ♂; Hanko, Tvärminne; 59.843 ° N, 23.206 ° E; Häyren leg.; http: // id. luomus. fi / GV. 93932; MZH • 1 ♂; Vantaa; 60.27 ° N, 24.96 ° E; 1 – 31 May 1905; Frey leg.; http: // id. luomus. fi / GV. 93946; MZH • 1 ♀; Vantaa; 60.27 ° N, 24.96 ° E; 1920; Wegelius leg.; http: // id. luomus. fi / GV. 93937; MZH • 1 ♀; Helsinki; 12 May 1945; Nuorteva, Matti leg.; http: // id. luomus. fi / GV. 93942; MZH • 1 ♀; Mäntsälä, Sahajärvi; 60.7159 ° N, 25.4429 ° E; 10 May 2005; Ståhls leg.; http: // id. luomus. fi / GJ. 6063; MZH • 1 ♂ *; Sipoo, Hindsby; 60.3508 ° N, 25.2001 ° E; 18 May 2022; Neuvonen leg.; http: // id. luomus. fi / GJ. 6271; MZH • 1 ♀ *; same data as for preceding; http: // id. luomus. fi / GJ. 6270; MZH. – Varsinais-Suomi • 1 ♀; Salo, Eriksberg; 60.374 ° E, 23.296 ° N; von Bonsdorff leg.; http: // id. luomus. fi / GV. 93939; MZH • 1 ♂; Lieto; 60.54 ° N, 22.5 ° E; Niemelä leg.; http: // id. luomus. fi / GV. 93944; MZH • 1 ♀; Salo; 60.189835 ° N, 22.876551 ° E; 16 May 1944; Hellman leg.; http: // id. luomus. fi / GV. 21348; MZH • 1 ♀; Salo; 60.189835 ° N, 22.876551 ° E; 17 May 1944; Hellman leg.; http: // id. luomus. fi / GV. 21349; MZH. GERMANY – Baden-Württemberg • 1 ♂; Schwarzwald, Zastler Hütte; 1256 m a. s. l.; 15 May 1992; Stuke leg.; CNC Diptera 101714. – Lower Saxony • 1 ♂; Lopautal; 12 Apr. 1991; Kassebeer leg.; CNC Diptera 101817. ITALY – Val Pusteria • 1 ♀; Monguelfo, Rienzi; 1000 m a. s. l.; 14 May 1985; Verlinden leg.; • 1 ♀; same data as for preceding; 16 May 1985. Most of Verlindens collection is destroyed now by Anthrenus museorum (Linnaeus, 1761) and not accessible anymore. SWEDEN – Småland • 1 ♂ *; Stenbrohult; 26 Apr. 2010; Nilsson leg.; MZH. – Uppland • 1 ♀; Uppsala, Dalkarlskärret; 1 May 1997; J. van Steenis leg.; JSB • 1 ♀; Halstavik, Pansarudden; 8 May 1997; J. van Steenis leg.; JSB • 1 ♀; Uppsala, Kodöden; 11 May 1997; J. van Steenis leg.; JSB • 1 ♀; Uppsala, Nåsten; 9 May 2002; J. van Steenis leg.; JSB.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D1394518815BFDC6FE0436313B0E.taxon	description	Specimens used for molecular analysis are indicated with an asterisk (*) and listed in Table 2. Description Male LENGTH. Body 7.5 – 9.5 mm, wing 6 – 8.1 mm. HEAD. Eye with brownish hairs. Anterior eye angle 120 ° – 130 °. Frons swollen, with median furrow running over entire length, obscured by faint pruinosity, with long, erect black hairs. Ocellar triangle equilateral, with long black hairs. Gena wide, viewed obliquely from below about as wide as 3 / 5 length of protibia. Face wide, ratio of width of face at level of facial tubercle to maximum width of head = 0.60 – 0.66: 1; face slightly shining, obscured by faint pruinosity and a narrow streak of pruinosity just below antennal insertion. Sides of face with long black hairs, spreading from level of antennal insertion towards level of facial tubercle. Parafacia narrow, fully developed only lateral to facial tubercle, about as wide as one third width of 3 rd antennal segment, above posterior tentorial pit strongly compressed toward eye margin by a haired bulge. Antennal pits separated. Lunule dark-brown to black. Antenna with scape and pedicel usually black, pedicel reddish to varying extents in individual specimens. 3 rd antennal segment sub-circular, usually velvet-black, but partly or entirely reddish in some specimens. Arista usually black, but occasionally more or less reddish in correspondence with colour of basoflagellomere; hairs on arista much shorter than maximum thickness of arista. THORAX. Scutum faintly brownish pruinose, anteriorly with a pair of contrasting sub-median pale pruinose vittae, best visible if viewed from behind; in well preserved specimens two less distinct lateral vittae perceptible; occasionally scutal vittae more or less confluent, displaying a joined pruinose area; hairs long black, erect, of about even length. Colour and hairs of scutellum similar to those of scutum, without distinct marginal setae, but occasionally with some marginal hairs slightly stronger than on disc. Sub-scutellar fringe black and long. Pleurae with long black hairs on posterior anepisternum, anepimeron and on katepisternum, dorsal and ventral katepisternal hair patches confluent. Metasternum haired. WING. Membrane entirely microtrichose, more or less brownish tinged, especially so in anterior half; veins dark, basal half of RS usually with a few dark setulae. Calypter brownish-grey to blackish, with dark rim; marginal fringe on dorsal lobe partly with black setae. Haltere with stem brownish and black knob. LEGS. Pro- and metalegs usually black, mesotibia often reddish basally, but sometimes up to basal one third and extreme apices of all tibiae, and apices of femora may be more or less reddish-yellow to brownish in individual specimens. Femora with black to brownish hairs, occasionally intermixed with a few paler hairs, especially so posteriorly on metafemur; ventral surface of metafemur with black setae on basal two thirds, but without setae apically. Tibiae covered in short, dark reddish-brown and black hairs, but hairs on anterior surface of protibia entirely and densely reddish brown. Tarsi with hairs on dorsal surface short black or mixed black and reddish-brown, hairs on ventral surface of pro- and metatarsi usually reddish-brown, but often intermixed with black hairs. Tarsi of mesolegs ventrally with black setulae. ABDOMEN. Tergite 4 and sides of tergites 1 – 3 slightly shiny, tergites 2 – 3 velvet-black on disc; pruinosity of tergites – viewed obliquely from front – dark brown; hairs of tergites long black, erect. Sternites moderately shining, faintly greyish-brown pruinose, with black hairs, hairs long and erect on sternites 1 – 2 and laterally on sternites 3 – 4, remaining hairs short, appressed or semi-appressed. GENITALIA. Right and left surstylus and gonostyli slightly asymmetrical. Ventral margin of surstylus with a strong basal convexity; hypandrium in lateral view about as long as wide; sclerite of distiphallus broadly fused dorsally, with short dorsal lobes. Female LENGTH. Body 7 – 8 mm, wing 7 – 8 mm. The ♀ differs from the ♂ in the following characters: hairs much shorter and predominantly pale yellowish-brown. HEAD. Hairs of eyes obviously shorter than width of postpedicel. Frons exceptionally wide, at level of lunule about half as wide as maximum width of head, with a distinct transverse sulcus at level of about lower one third of frons and with remnant of a median furrow between lunule and transverse sulcus; transverse sulcus and median furrow faintly grey pruinose, frontal areas above and below transverse sulcus otherwise shining; lateral channels narrow, widening from inner dorsal corner of eye toward transverse sulcus, where inner margin of channel encloses a small semicircular area of dense grey pruinosity; hairs of frons erect, not longer than half width of 3 rd antennal segment, varying in colour from entirely yellow to entirely black. Occiput dorsally much wider than in male, faintly grey pruinose, with short yellow hairs, or with a mixture of yellow and black hairs. Gena with yellow hairs. Face at level of facial tubercle about 0.5 times as wide as maximum width of head; faintly grey pruinose, except for a narrow non-pruinose stripe, between dorsal margin of facial tubercle and upper margin of oral cavity, spreading toward lower corner of oral cavity. Facial hairs yellow, short, occasionally inconspicuous. Parafacia usually less strongly compressed above level of facial tubercle. 3 rd antennal segment larger, most often bright orange-coloured, or with a black apico-dorsal rim. THORAX. Scutum and scutellum less densely pruinose, but with pair of pale pruinose sub-median vittae anterior on scutum distinct; hairs short, about as long as distance between rear ocelli, yellow, intermixed with scattered, somewhat longer black hairs. Scutellum usually with a few (2 – 4) black or yellow marginal setae, sub-scutellar fringe short and yellow. Pleurae with yellow hairs, pattern of hairs as in male. Calypter whitish-orange, with darker rim and yellow fringe. Haltere stem brownish, knob yellow. LEGS. Legs black, but basal 1 / 3 – 2 / 5 of tibiae and often extreme apices of tibiae and femora reddish-yellow. Femora predominantly with yellow hairs, but apices anteriorly and dorsally with short appressed black hairs, posteriorly occasionally with a few longer, thin black setae. Ventral surface of metafemur basally without black setulae. ABDOMEN. Tergites slightly shiny, especially tergites 2 – 3 much less pruinose on disc, hairs on tergites entirely yellow, short, erect. Sternites greyish pruinose to varying extents, from entirely pruinose to slightly shiny, with short yellow hairs; hairs erect on sternites 1 – 2 and on anterior corners of sternite 3, remaining hairs more or less appressed.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D1394518815BFDC6FE0436313B0E.taxon	distribution	Distribution Austria, Czech Republic, Finland, France, Germany, Greece, Italy, Montenegro, Norway, Poland, Russia, Slovakia, Sweden, Switzerland. Distribution reported here for Austria, Finland, Norway and Switzerland were not reported before in the IUCN Red List assessment (Vujić et al. 2021).	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D1394518815BFDC6FE0436313B0E.taxon	diagnosis	Differential diagnosis In its overall appearance similar to Cheilosia luteicornis, with the following differences: Sides of face haired, ♂ with long black hairs, ♀ with yellowish hairs which are less conspicuous. Parafacia narrow, about 1 / 3 of the width of postpedicel. Face and frons slightly wider than in C. luteicornis. ♀ with hairs on scutum of about equal length, with only individual black hairs intermixed. ♂: face with a slight, but distinct haired bulge on each side, just dorsal to the facial prominence. Bases of pro- and metatibiae usually blackish or obscured reddish. Abdomen entirely black-haired. Calypter brownish-grey to blackish. Genitalia: surstylus with strong basal convexity; hypandrium about as long as wide.	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
03D4D1394518815BFDC6FE0436313B0E.taxon	discussion	Remarks The holotype specimen bears labels with what could seem to be contradicting information on the locality, but there is no doubt that this specimen is the holotype collected in Lapland. The label mentioning E. lineata and Holmia is similar to other bottom-drawer labels used by Zetterstedt for species of which he had no material, and most likely this label was added to the specimen by Zetterstedt himself when he synonymized E. lineata with E. morio. The type accords well with the original diagnosis and morphological description and with its subsequent additions (Zetterstedt 1843: 795), but the description of the facial hairs does not: The face of the male is described as “ hypostoma nudum [!], nitidum, atrum [face bare, shiny, black] and later (Zetterstedt 1843: 795) as “ epistomate bituberculate, nudo [!], nitido ” [face bituberculate, bare, shiny black], by this giving the impression of a species with a face lacking hairs. Actually, the face of the holotype clearly has long, black hairs laterally below the antennal insertion, with individual hairs reaching the level of facial tubercle. As a consequence of this discrepancy between the description and the characters actually observed, C. morio was erroneously and almost consistently regarded as a species with a bare face throughout the central European literature. The nomenclatural consequences will be discussed below. DNA barcoding A total of 18 COI barcodes of 633 bp length representing four species of the subgenus Neocheilosia and additionally the barcodes of two species of the subgenus Convocheila (Barkalov 2002) and one species of the subgenus Montanocheila Barkalov, 2002 were used for the NJ distance tree (for specimen data and GenBank accession numbers see Table 2). The NJ tree (Fig. 6) consistently resolved the included male and female individuals of Cheilosia barovskii, C. luteicornis, C. morio and C. shiranesana in separate clusters, respectively. All the bootstrap values supporting the nodes of the species clusters were high (99 – 100 %). The uncorrected interspecific pairwise distances among the included species of the subgenus Neocheilosia ranged from 2.96 % (between C. luteicornis and C. morio) to 7.57 % (between C. barovskii and C. shiranesana).	en	Bot, Sander, Ståhls, Gunilla, Bygebjerg, Rune, Doczkal, Dieter, Vujić, Ante (2025): Revision of the European species of the subgenus Neocheilosia Barkalov (Diptera, Syrphidae: Cheilosia). European Journal of Taxonomy 997: 1-27, DOI: 10.5852/ejt.2025.997.2917, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2917/13205
