identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D187A9761DFFDCFCAEFCE8A5C466CF.text	03D187A9761DFFDCFCAEFCE8A5C466CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatoma Woodward 1859	<div><p>Genus Anatoma Woodward, 1859</p><p>Type species: Scissurella crispata Fleming,</p><p>1828, subsequently designated by Geiger (2012).</p></div>	https://treatment.plazi.org/id/03D187A9761DFFDCFCAEFCE8A5C466CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nekhaev, I. O.;Krol, E. N.	Nekhaev, I. O., Krol, E. N. (2020): A review of the genus Anatoma in the Eurasian Arctic seas (Gastropoda: Vetigastropoda: Anatomidae). Zoosystematica Rossica (China) 29 (1): 128-137, DOI: 10.31610/zsr/2020.29.1.128, URL: https://doi.org/10.31610/zsr/2020.29.1.128
03D187A9761DFFDAFCAEFC48A2CB6042.text	03D187A9761DFFDAFCAEFC48A2CB6042.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatoma crispata (Fleming 1828)	<div><p>Anatoma crispata (Fleming, 1828)</p><p>(Figs 1A–G; 2A–F; 3A–F; 4C)</p><p>Scissurella crispata Fleming, 1828: 385, pl. 6, fig. 3.</p><p>Scissurella angulata Lovén, 1846: 20 .</p><p>Anatoma schioettei HØisaeter et Geiger, 2011: 103– 105, figs 48–60; Geiger, 2012: 1086–1089, figs 886–889, syn. nov.</p><p>Scisurella crispata: Herzenstein, 1885: 67; Thiele, 1928: 565.</p><p>Anatoma crispata: Golikov et al., 2001: 104 (partim); Kantor &amp; Sysoev, 2006: 24, fig. 9A (partim); HØisaeter &amp; Geiger, 2011: 90–94, figs 1–18; Nekhaev, 2014: 78; Geiger, 2012: 832–847, figs 674–680.</p><p>Anatoma sp.: Nekhaev &amp; Krol, 2017: 2283.</p><p>Material examined. Norway, Norwegian Sea, Fin-</p><p>nmark, S. Lovén leg., 1 shell (SMNH 4394, lecto-</p><p>type of Scissurella angulata). Russia: Barents Sea:</p><p>60 m, 71°54.26′N, 47°51.52′E, 18 Aug. 2006, R / V</p><p>Dal’nie Zelentsy, 2 spms, 1 shell; Motovskiy Bay, 197</p><p>m, 69°36.87′N, 32°16.43′E, 26 May 1996, M/S GS-</p><p>440, 4 shells (ZIN 62141/36); Murman Coast, 42 m,</p><p>69°40.82′N, 31°37.00′E, 6 July 2005, R / V Dal’nie</p><p>Zelentsy, 1 shell; Murman Coast, 18–20 m, 69°36.5′N,</p><p>32°29.5′E, 29 May 1901, R / V Andrey Pervozvannyy,</p><p>3 shells (ZIN 1607 /7); Ura Inlet, 17 m, 69°22.70′N,</p><p>32°54.88′E, 6 Oct. 2006, 4 shells; Ura Inlet, 158 m,</p><p>69°27′N, 33°08′E, 2 Oct. 2006, 1 spm; Ura Inlet, mouth of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=33.133335&amp;materialsCitation.latitude=69.45" title="Search Plazi for locations around (long 33.133335/lat 69.45)">Shalimskaya Bay</a>, 24–40 m , 13 Aug. 1887, 1 shell (ZIN 1602 /1); Ivanovskaya Inlet, 54 m , 68°22.46′N, 38°32.30′E, 30 July 2008, R / V Dal’nie Zelentsy, 1 spm; Kola Inlet, 22 m, 69°16.81′N, 33°32.99′E, 28 May 2013, Yu.A. Zuev &amp; S. V. Goldin leg., 4 shells (ZIN 32142 /37); off Teriberka, 150–170 m, 6 Aug. 1884, 1 shell (ZIN 1603 /2); Dal’ne-Zelenetskaya Bay, 66 m , 69°08.41′N, 36°04.54′E, 4 June 2009, R / V Dal’nie Zelentsy, 2 spms; Franz Joseph Land: 251 m, 80°44.19′N, 53°36.85′E, 23 Aug. 2006, R / V Dal’nie Zelentsy, 2 shells (ZIN 62140 /35), 76 m, 80°31.60′N, 52°34.12′E, 29 Aug. 2007, R / V Dal’nie Zelentsy, 1 shell, 102 m, 79°52.05′N, 51°49.13′E, 24 Aug. 2006, R / V Dal’nie Zelentsy, 3 spms, 53 m, 80°14.5′N, 73°51′E, 14 Sept. 1936, R / V Sadko, 4 spms (ZIN 1615 /15), Hooker I., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=73.85&amp;materialsCitation.latitude=80.24167" title="Search Plazi for locations around (long 73.85/lat 80.24167)">Tikhaya Bay</a>, 120 m , 25 Aug. 1992, R / V Pomor, 2 spms (ZIN 58716 /34); Novaya Zemlya, Chernaya Inlet, 46 m, 70°35′N, 54°55′E, 8 Oct. 2000, R / V Dal’nie Zelentsy, 2 spms.; Kara Sea: 446 m , 80°58.5′N, 73°32′E, 9 Sept. 1936, R / V Sadko, 4 shells (ZIN 1614 /14); 41 m, 80°11′N, 75°02′E, 18 Aug. 1934, R / V Sedov, 21 shells (ZIN 37030 /20) .</p><p>Redescription. Shell small, globular, fragile, matt or slightly shiny, from white to yellowish, semitransparent or opaque. Spire low; upper whorls from rounded to stepped; body whorl relatively large, lenticular; shell base convex. Protoconch of 0.75–1.0 whorls, with sculpture of coarse flocculae which usually tending to form two or three spiral riblets. Diameter of protoconch from 210 to 320 µm, nucleus 70–180 µm wide. Teleoconch I of 0.25–1.0 whorls. Teleoconch II of 0.7– 2.0 whorls. Upper parts of body whorl with 3–14 thin spirals riblets, shell base with 7–21 ones. Teleoconch I with 13–25 axial riblets, sometimes weak spiral cord present in position of selenizone (Fig. 2C). First whorl of teleoconch II with 33–65 axial riblets. Body whorl with 43–80 axials. Axials irregularly spaced (Fig. 1F). Spirals weaker than axials. Spirals and axials on base forming reticulate pattern. Aperture rounded; outer lip above selenizone arciform or almost straight, out- er lip under selenizone and columellar lip round- ed. Umbilicus open, deep with distinct funiculus. Space between selinizone (Fig. 1D, s) and suture of subsequent whorl variable (Fig. 1G).</p><p>Mean values of measurements of 34 specimens: NW 2.9 ± 0.04, SH 1.38 ± 0.06 mm, BWH 1.25 ± 0.05 mm, AH 0.93 ± 0.03 mm, SW 1.73 ± 0.06 mm, AW 0.90 ± 0.03 mm. Measurements of the largest specimen (Franz Josef Land, 76 m): NW 3, SH 2.05 mm, BWH 1.82 mm, AH 1.30 mm, SW 2.34 mm, AW 1.17 mm.</p><p>Radula (Fig. 3A–F): rachidian tooth trapezoidal, with prominent central denticle and 3–6 narrow pointed denticles on either side. Five lateral teeth with narrow pointed denticles. Lateral teeth 1–3 usually with 5–8 denticles. Fourth lateral tooth reduced, with distinct terminal denticle. Fifth lateral tooth with approximately 7–9 denticles. Tips of lateral teeth directed towards inner margin, rachidian and lateral teeth with slightly serrate edges. Inner marginal teeth with triangular tips directed towards outer margin, bearing approximately 7–9 narrow denticles. Outer marginal teeth spoon-shaped, with approximately 15–20 thin sharp denticles.</p><p>Distribution and habitat. Anatoma crispata is known from Baffin Bay, Greenland, Shetland Islands, Scandinavian coast, Barents Sea and northwestern Kara Sea; the depth range is from 10 to 1083 m (HØisaeter &amp; Geiger, 2011; Geiger, 2012). The records of A. crispata from the northeastern Pacific (Sirenko et al., 2013) probably refer to another species and need confirmation. Also, we did not find any specimens identified as A. crispata in ZIN collection. The findings of A. crispata in the Laptev Sea and East Siberian Sea (e.g. Gorbunov, 1946; Golikov et al., 2001) actually belong to A. schanderi . Anatoma crispata was found on different substrates. The minimum observed temperature at the collection site of living specimens was −0.8°C (Franz Josef Land).</p><p>Remarks. HØisaeter &amp; Geiger (2011) described A. schioettei from the upper slope of Greenland. This species differs from A. crispata in having a wider sutsel (space between the beginning of selenizone and the suture of the subsequent whorl) and in having about 1.5 times as many or more distinct axials (e.g. 22 vs 15 on teleoconch I) (HØisaeter &amp; Geiger, 2011). However, these characters were variable in our material and numerous specimens displayed significant non-discrete variation: they differed in the position of selenizone (Fig. 1G), in the number of axial riblets and in the distance between them (Fig. 1F); the variability occurred even between the different parts of the same shell.</p><p>The number of axial ribs varied from 8 to 25 (mean 14.64 ± 0.74, n = 22) on teleoconch I and from 33 to 65 (mean 46.86 ± 1.09, n = 36) on the first whorl of teleoconch II. The hypothesis of normal distribution cannot be statistically rejected for both values (teleoconch I: Shapiro-Wilk test = 0.919, p = 0.073; first whorl of teleoconch II: Shapiro–Wilk test = 0.971, p = 0.45).</p><p>Both the forms also do not substantially differ from each other by other characters such as shell shape, sculpture and size of protoconch, and</p><p>I.O. Nekhaev &amp; E.N. Krol. Review of the genus Anatoma radular morphology (see Figs 1B, 2B, 2E, 2F, 3D for typical A. crispata; Figs 1A, 2A, 2C, 2D, 3A– C, 3E, 3F for typical A. schioettei; and Figs 1C–D for transitional forms). Therefore, we consider A. schioettei as a synonym of A. crispata .</p></div>	https://treatment.plazi.org/id/03D187A9761DFFDAFCAEFC48A2CB6042	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nekhaev, I. O.;Krol, E. N.	Nekhaev, I. O., Krol, E. N. (2020): A review of the genus Anatoma in the Eurasian Arctic seas (Gastropoda: Vetigastropoda: Anatomidae). Zoosystematica Rossica (China) 29 (1): 128-137, DOI: 10.31610/zsr/2020.29.1.128, URL: https://doi.org/10.31610/zsr/2020.29.1.128
03D187A9761BFFD5FF09FAC7A162677E.text	03D187A9761BFFD5FF09FAC7A162677E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatoma golikovi Nekhaev & Krol 2020	<div><p>Anatoma golikovi sp. nov.</p><p>(Fig. 4A–B, D–F)</p><p>Holotype. Norway, Barents Sea, Svalbard, near Hopen I., 49 m, 76°11.04′N, 23°11.78′E, 21 Aug. 2008, R / V Dal’nie Zelentsy (ZIN 62138 /1).</p><p>Diagnosis. Anatoma golikovi sp. nov. differs from all Arctic species of the genus in its planispiral shell with a depressed spire, the presence of distinct funiculus, and embryonic shell with sculpture of loose flocculae.</p><p>Description. Shell small, fragile, slightly shiny, cream, semitransparent. Spire flat; body whorl relatively large, lenticular; shell base convex. Protoconch of 0.8 whorls, with sculpture of loose flocculae, which tending to form irregular spiral riblets. Diameter of protoconch 240 µm, nucleus 70 µm wide. Teleoconch I of 0.7 whorls. Teleoconch II of 1.1 whorls. Six thin spirals riblets on upper parts of body whorls and 17 on shell base. Teleoconch I with 16 axial riblets. Teleoconch II with 51 strong axial ribs. Shell base with 60 axial riblets. Spirals and axials forming reticulate sculpture around umbilicus.</p><p>Aperture rounded; outer lip almost straight above selenizone and rounded under selenizone and columellar lip. Umbilicus open, deep, with distinct funiculus. Space between selinizone and suture of subsequent whorl almost equal to width of selenizone.</p><p>Measurements of holotype: NW 2.6, SH 0.93 mm, AH 0.86 mm, SW 1.50 mm, AW 0.79 mm.</p><p>Distribution. Known only from the type locality.</p><p>Habitat. The holotype was collected from the sea bottom with sand, shellrock and pebbles. The temperature at the collecting site was +2.17°C, salinity 33.84‰.</p><p>Remarks. Anatoma crispata differs from A. golikovi sp. nov. in having an elevated spire (see Fig. 4B–C for comparison) which is visible even in young specimens (with less than one whorl on teleoconch II). Anatoma schanderi can be distinguished from the new species by its larger size, elevated spire and reticulate pattern on the embryonic shell. Anatoma tenuisclupta (Seguenza, 1880) which is known from the Norwegian coast differs in the lacking funiculus and elevated spire.</p><p>Etymology. The species is named after Alexan- der Golikov, a prominent malacologist and benthic ecologist.</p></div>	https://treatment.plazi.org/id/03D187A9761BFFD5FF09FAC7A162677E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nekhaev, I. O.;Krol, E. N.	Nekhaev, I. O., Krol, E. N. (2020): A review of the genus Anatoma in the Eurasian Arctic seas (Gastropoda: Vetigastropoda: Anatomidae). Zoosystematica Rossica (China) 29 (1): 128-137, DOI: 10.31610/zsr/2020.29.1.128, URL: https://doi.org/10.31610/zsr/2020.29.1.128
03D187A97614FFD5FF06FD1BA39764CA.text	03D187A97614FFD5FF06FD1BA39764CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anatoma schanderi Hoisaeter & Geiger 2011	<div><p>Anatoma schanderi HØisaeter et Geiger, 2011</p><p>(Fig. 5A–C)</p><p>Scissurella crispata: Gorbunov, 1946: 45.</p><p>Anatoma crispata: Golikov et al., 2001: 104 (partim).</p><p>Anatoma schanderi HØisaeter et Geiger, 2011: 106–109,</p><p>figs 68–83; Geiger, 2012: 1081–1085, figs 881–885.</p><p>Material examined. Russia: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.21666&amp;materialsCitation.latitude=78.21667" title="Search Plazi for locations around (long 142.21666/lat 78.21667)">East Siberian Sea</a>, 57 m, 78°13′N, 142°13′E, 19 Dec. 1937, R / V <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.68333&amp;materialsCitation.latitude=76.8" title="Search Plazi for locations around (long 129.68333/lat 76.8)">Sadko</a>, 1 shell (ZIN 37032); Laptev Sea, 68 m, 76°48′N, 129°41′E, 8 Nov. 1937, R / V Sadko, 1 shell (ZIN 37031) .</p><p>Redescription. Shell large, towering, fragile, slightly shiny, semitransparent. Spire high; whorls stepped; shell base convex. Protoconch of 0.75 whorls, with reticulate sculpture. Diameter of protoconch 190 µm, nucleus have 100 µm width. Teleoconch I of 1.0 whorl. Teleoconch II of 3.2 whorls. Upper parts of body whorls with 14 thin spirals riblets, shell base with 31 ones. Teleoconch I with 30 axial riblets; strong spiral cord present in position of selenizone. First whorl of teleoconch II with 41 axial riblets. Body whorl with 81 axials. Axials irregularly spaced. Spirals and axials of same size, forming reticulate sculpture. Aperture rounded. Umbilicus open, deep, with small funiculus. Space between selinizone and suture of subsequent whorl as wide as selenizone.</p><p>Measurements of largest specimen examined: NW 4.9, SH 4.20 mm, BWH 3.70 mm, AH 2.30 mm, SW 4.25 mm, AW 2.70 mm.</p><p>Remarks. The two empty shells collected from the waters around the New Siberian Islands (the Laptev Sea and the East Siberian Sea) clearly correspond to the description of A. schanderi, which was previously known only from the extreme North of the Atlantic: from Nova Scotia to the Norwegian Sea. The examined specimens are the only representatives of the genus known from the region, and apparently all the records of A. crispata from the Laptev Sea and the East Siberian Sea by the Russian authors (e.g. Gorbunov, 1946; Golikov, 1995; Golikov et al., 2001; Kantor &amp; Sysoev, 2006) in fact refer to these specimens of A. schanderi .</p></div>	https://treatment.plazi.org/id/03D187A97614FFD5FF06FD1BA39764CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nekhaev, I. O.;Krol, E. N.	Nekhaev, I. O., Krol, E. N. (2020): A review of the genus Anatoma in the Eurasian Arctic seas (Gastropoda: Vetigastropoda: Anatomidae). Zoosystematica Rossica (China) 29 (1): 128-137, DOI: 10.31610/zsr/2020.29.1.128, URL: https://doi.org/10.31610/zsr/2020.29.1.128
