identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D187CA7073B861FEBBFA02FA516423.text	03D187CA7073B861FEBBFA02FA516423.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cenomanisega Lucena & Melo 2025	<div><p>Genus † Cenomanisega gen. nov.</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: E4DA9E 8E-6EA3-4BCC-8CC1-62C92BCD3B54</p><p>Type species: † Cenomanisega krombeini sp. nov.</p><p>Diagnosis</p><p>Male mandible elongated, flattened, broadly foliaceous, lacking subapical tooth, apparently truncate apically; female mandible delicate, tapering apically, without subapical teeth on laminar edge; general body shape long and flattened dorsoventrally; pronotum long, slightly longer than scutum in mid-line, posterolateral lobe well separated from tegula, lacking posterolateral and posteromedial pits; mesopleuron anteriorly delimited with carina, mostly evenly convex, with broad posterior depression below scrobal pit, without transverse sulcus; posterolateral corner of propodeum round, lacking posterior propodeal projection, posterior surface declivitous.</p><p>Description</p><p>Head: Pyriform, front well developed and protruding; eye bulging, occupying much of lateral head, following contour of frons anteriorly; lower gena 0.4 × eye width, upper gena narrowed, nearly indistinct; malar space slightly shorter than 0.4 × eye height, crossed by longitudinal malar sulcus; pre-hypostomal foveae absent; occipital carina extended dorsally, fading gradually laterally; scapal basin shallowly excavated, disc smooth, borders smoothly round, without carina or sharp edges; scape length 4.1 × maximum width; labial and maxillary palpomeres with three and four articles, respectively; flagellum fusiform, with intermediate articles broader than long, progressively shortened from F4 to F10 and with flattened ventral areas on F4–F11; mandible strongly tapering apically, main axis straight, lacking subapical teeth on laminar edge.</p><p>Mesosoma: Elongated and somewhat fusiform, slightly flattened dorsally; pronotum long, slightly longer than scutum in mid-line, posterior border concave, crossed anteriorly and posteriorly by transverse grooves, posterolateral lobe well separated from tegula, lacking posterolateral and posteromedial pits; notaulus deeply impressed, strongly converging posteriorly; parapsidal line well marked posteriorly; scutellum slightly shorter than half length of scutum in mid-line, simple, evenly convex, without anterior transverse groove, pits, or foveae; mesopleuron anteriorly bordered with carina, evenly convex, with wide posterior shallow depression below scrobal pit, without transverse sulcus; metanotum about one-third of scutellum in mid-line, expanded and slightly depressed laterally, without differentiated disc; propodeum box-like, dorsum longer than high, with distinguishable lateral, dorsal, and posterior surfaces; dorsal and posterior surfaces medially crossed by contiguous longitudinal groove; dorsum and posterior surface delimited laterally by confluent grooves, metapostnotum indicated anteriorly as a distinct transverse raised band, posterolateral corner round, lacking posterior propodeal projection, posterior surface declivitous; tarsal claw with short, perpendicular, submedial tooth; inner metatibial spur distinctly shorter than the outer; dorsum of metacoxa with strongly produced longitudinal carina; dorsum of metafemur evenly convex, without anterior expansion; costal cell narrow, first abscissa of M diverging before 1cu-a, gently and smoothly curved submedially; Rs+M arising apically on M; medial cell indicated by spectral and nebulous veins; R1 slightly shorter than pterostigma; Rs smoothly curved, distal trace fading gradually towards costal margin.</p><p>Metasoma: Four exposed terga, internalized sclerites partly exposed, composing a very slender and delicate metasomal tube; sterna convex and undivided medially.</p><p>Male: Same as female, except: mandible elongated, flattened, broadly foliaceous, lacking subapical or distal teeth, apparently truncate apically; lower medial margin of clypeus evenly convex; inner margin of antennal sockets touching; scape and pedicel comparatively shorter; flagellum cylindrical, without ventral flattened areas; frons with medial longitudinal groove, extending from median ocellus towards scapal basin; metabasitarsus without two rows of spines; T1–T4 bordered laterally by confluent strong carinae, delimiting the laterotergites; distal elements of genitalia partly exposed, with typical long, slender, and undifferentiated paramere, volsella, and aedeagus.</p><p>Etymology</p><p>The name is a combination of the words ‘Cenomanian’ (geological age); and - sega, from Amisega Cameron. The name is feminine.</p><p>† Cenomanisega krombeini sp. nov.</p><p>(Figs 1–3)</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: 6D1 07F33-753B-4E15-9588-595DC70ADEDC</p><p>Description</p><p>Holotype female (DZUP Bur-283). Body length, ~ 4.1 mm.</p><p>Head: LID very narrow, ~0.3 × eye width; upper limit of eye surpassing lateral ocellus, inner ocular margins strongly converging toward the upper frons and diverging near posterior margin of head; ocelli deeply inserted, with well-marked small sulci around them, lateral ocelli nearly touching inner ocular margin, closer together than with median ocellus; inner margin of antennal sockets nearly touching; pedicel about three-quarters of F1 length; F1 longer than combined length of F2 + F3; F2 and F3 distinctly shortened, F11 acuminate, slightly longer than F10; subantennal distance ~1 × MOD, clypeus thickened, lower medial margin slightly concave; head finely and sparsely punctate, becoming denser below on face and gena; setation nearly indistinct, short and sparse, becoming slightly longer and denser below on face and occiput; eye glabrous.</p><p>Mesosoma: Lower posterior corner of propleuron smoothly round; basisternum not enlarged, disc slightly depressed, posterior process short and broadly round, lateral process long and acute; anterior portion of pronotum short, ~0.3 × total length after transverse groove in mid-line, lateral depression smoothly excavated; profemur thoroughly cylindrical and elongated; metabasitarsus with two rows of short, suberect, thickened spines; dorsum finely and regularly punctate, interspaces slightly rugose, particularly on pronotum and scutum; propodeum mostly smooth, with scarce marginal punctures; metapleuron– propodeum with large medial polished band; dorsum nearly glabrous; some short, suberect, gold setation, becoming denser and longer on legs and mesopleuron; wing membrane entirely setose.</p><p>Metasoma: Metasomal terga finely punctate, becoming denser marginally on T3; laterotergites of T1–T2 polished; with short, suberect, gold setation, becoming denser marginally on metasomal sclerites.</p><p>Male: See above for sexually dimorphic characters provided in the description of the genus.</p><p>Etymology</p><p>The species is named after Dr Karl V. Krombein, whose contributions to the biology and systematics of the Amiseginae established the modern foundations for studies involving these fascinating wasps.</p><p>Type material</p><p>Holotype female in amber piece DZUP Bur-283. The specimen is well preserved and complete. Syninclusion: a nematoceran mosquito.</p><p>Paratypes: Two males in the same amber piece, DZUP Bur-505, syninclusion: a wasp; one female, DZUP Bur-1019; one female, DZUP Bur-410, syninclusions: a beetle, five dipterans, and a cricket; one female, DZUP Bur-577; one female, DZUP Bur-771; one female, DZUP Bur-074; one female, DZUP Bur-1683; and one female, DZUP Bur-1574, syninclusions: four beetles and a wasp .</p><p>Additional examined material (non-type)</p><p>One male in amber piece DZUP Bur-1772; one female, DZUP Bur-1367; and one male, DZUP Bur-1249 .</p><p>Remarks</p><p>The new genus does not resemble any extant genus of Amiseginae, but it is superficially similar to † Protamisega Evans (known only from the male holotype of Taimyr amber, Russia; Late Santonian, 85.8–84.9 Mya). It can be distinguished from † Protamisega by the combination of the following characters: malar space crossed by longitudinal malar sulcus (lacking in † Protamisega); pronotum crossed anteriorly and posteriorly by transverse grooves (crossed anteriorly by transverse carina that curves towards midline in † Protamisega); notaulus deeply impressed, reaching anterior margin of scutum (notaulus evanescent anteriorly in † Protamisega); parapsidal line well marked (parapsidal line absent or indistinct in † Protamisega); anterior band of scutellum simple, lacking transverse groove (anterior band of scutellum with transverse groove medially, divided by longitudinal carina in † Protamisega); dorsum of propodeum moderately long (dorsum of propodeum short, shelf-like in † Protamisega), lacking longitudinal carinae delimiting a series of foveae (with several longitudinal carinae delimiting a series of foveae in † Protamisega), posterolateral corner round, lacking posterior propodeal projection (posterolateral corner obtusely angulate in † Protamisega); posterior declivity of propodeum without carinae radiating outwards (posterior declivity of propodeum with several carinae radiating outwards in † Protamisega); and dentate tarsal claw (simple in † Protamisega).</p></div>	https://treatment.plazi.org/id/03D187CA7073B861FEBBFA02FA516423	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lucena, Daercio A. A.;Melo, Gabriel A. R.	Lucena, Daercio A. A., Melo, Gabriel A. R. (2025): Expanding the Amiseginae: remarkable new Cretaceous-age fossils shed light on the early diversification of a highly specialized clade of cuckoo wasps (Hymenoptera: Chrysididae). Zoological Journal of the Linnean Society 204 (2), DOI: 10.1093/zoolinnean/zlaf054, URL: https://doi.org/10.1093/zoolinnean/zlaf054
03D187CA7074B86CFC6CFA6FFD706742.text	03D187CA7074B86CFC6CFA6FFD706742.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretamisega Lucena & Melo 2025	<div><p>Genus † Cretamisega gen. nov.</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: 336 23697-EAAB-47DF-B154-85B9B1F31C7B</p><p>Type species: † Cretamisega mandibularis sp. nov.</p><p>Diagnosis</p><p>Female mandible distinctly elongated, relatively robust (compared with other Amiseginae) and lacking subapical teeth on laminar edge; male mandible elongated, broadly flattened, apically sharp, culminating in a short distal tooth, laminar edge lacking tooth; head not pyriform, somewhat round in profile; general body shape short and stout; malar space moderately short, 0.3 × eye height, longitudinally crossed by malar sulcus; posterolateral lobe of pronotum reaching tegula, lacking posterolateral and posteromedial pits; posterolateral corner of propodeum round, without distinct projection.</p><p>Description</p><p>Head: Not pyriform, somewhat round in profile, with frons not protruding and broad face; frons without longitudinal groove; eye bulging, ovoid, occupying much of lateral head; labial and maxillary palpomeres with three and four articles, respectively; gena wide, becoming broadened below; malar space 0.3 × eye height, longitudinally crossed by discrete malar sulcus; pre-hypostomal foveae absent; occipital carina indistinct, apparently only extended dorsally; scapal basin shallowly excavated, disc smooth, borders smoothly round, lacking carina or sharp edges; scape length 3.8 × maximum width, gently curved submedially; pedicel short; flagellum fusiform, with intermediate articles broader than long, progressively shortened from F4 to F10 and with ventral flattened areas on F4–F11; mandible huge, 0.8 × eye height, tapering apically, gently curved backwards, lacking subapical teeth on laminar edge.</p><p>Mesosoma: Evenly convex, not flattened dorsally, somewhat shortened anteroposteriorly; pronotum about as long as scutum in mid-line,anteriorly crossed by transverse groove;posterolateral lobe of pronotum reaching tegula, lacking posterolateral and posteromedial pits; notaulus deeply impressed, parallel-sided posteriorly; parapsidal line well marked posteriorly; scutellum slightly shorter than half length of scutum in mid-line, simple, evenly convex, without anterior transverse groove, pits, or foveae; mesopleuron simple, evenly convex, without transverse groove, anterior margin evenly round; metanotum about half length of scutellum in mid-line, slightly expanded and depressed laterally, without differentiated disc; propodeum somewhat round, with distinguishable lateral, dorsal, and posterior surfaces; dorsum relatively short, simple, lacking transverse posterior groove or carina, smoothly round, metapostnotum undistinguishable, posterolateral corner evenly round, lacking posterior propodeal projection; dorsum and posterior declivity delimited laterally by discrete, confluent groove; posterior declivity gradually declivitous, simple, lacking medial longitudinal groove; tarsal claw with short, perpendicular, submedial tooth; dorsum of metacoxa with longitudinal carina; dorsum of metafemur evenly convex, without anterior expansion; costal cell very narrow, first abscissa of M diverging before 1cu-a, gently curved submedially; Rs+M arising apically on M; medial cell indicated by spectral veins; R1 well developed, longer than stigma; Rs smoothly curved, nebulous distal trace fading gradually towards costal margin.</p><p>Metasoma: Four exposed terga; sterna convex and undivided medially; internalized sclerites partly exposed, composing a very slender and delicate metasomal tube.</p><p>Male: Same as female, except: mandible broadly flattened, apically sharp, culminating in a distal tooth, laminar edge lacking subapical tooth; inner margin of eye more strongly convergent on upper frons; lateral ocellus nearly touching inner margin of eye; LID and subantennal distance comparatively narrower than in female; scape cylindrical, not curved submedially, flagellum more cylindrical and elongated, without flattened ventral areas; metanotum evenly convex; metasoma with five exposed terga.</p><p>Etymology</p><p>The genus name is a combination of cret-, from Cretaceous, and Amisega Cameron, the type genus of the subfamily. The name is feminine.</p><p>† Cretamisega mandibularis sp. nov.</p><p>(Figs 4–6)</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: 46E5 E089-61B1-4449-89EF-78C490B6D352</p><p>Description</p><p>Holotype female (DZUP Bur-1679). Body length, ~ 3.6 mm.</p><p>Head: Upper limit of eye surpassing dorsal margin of lateral ocellus; inner margin converging gently on upper frons; LID wide, slightly shorter than eye width; ocelli inserted smoothly, without small sulci around them, lateral ocelli closer to inner eye margin and median ocellus than to each other; POL 0.4 × OOL, 0.7 × OL; inner margin of antennal sockets nearly touching; F11 gently acuminate, slightly longer than F10; F2 and F3 distinctly shortened, F1 longer than combined length of F2 + F3; subantennal distance ~1 × MOD, lower medial margin of clypeus evenly convex.</p><p>Mesosoma: Lower posterior corner of propleuron smoothly round;anteriorportionofpronotumshorterthanhalfofposterior band, posterior margin without transverse groove; anterobasal margin of procoxa with transverse carina; metabasitarsus lacking two rows of spines.</p><p>Metasoma: Lateral margins broadly round, without carina or sharp edge delimiting the lateral fold. Details of the integument (sculpturing, vestiture, and coloration) are not discernible.</p><p>Male: See above for sexually dimorphic characters provided in description of the genus.</p><p>Etymology</p><p>The species is named after the long mandible, characteristic of the female.</p><p>Type material</p><p>Holotype female in amber piece DZUP Bur-1679. The specimen is well preserved and almost intact. Part of distal flagellomeres of the left antenna is lacking. There are some cracks in the amber piece, blocking most of the frontal view of the head and lateral right of the specimen.</p><p>Paratypes: One male in amber piece DZUP Bur-883; one male, DZUP Bur-1653; and female, DZUP Bur-1202, syninclusion: a cockroach .</p><p>Additional examined material (non-type) One male, DZUP Bur-1867 .</p><p>Remarks</p><p>The new genus superficially resembles the extant genera Amisega Cameron and Duckeia Costa Lima, based on the short and stout body, simple and round propodeum, lacking marginal carina or groove, round posterolateral corner lacking process, and the general shape of the head (particularly Amisega). The relatively broadened metanotum is like Duckeia, but in † Cretamisega it is further expanded and depressed laterally. The huge and specialized female mandible of † Cretamisega is unique within the Amiseginae s.l. Additionally, the new genus is readily distinguished from Amisega and Duckeia based on the combination of the following characters: male with strongly modified mandibles (males of Amisega and Duckeia lack dimorphic mandibles); pronotum anteriorly crossed by transverse groove (absent in Amisega and Duckeia), posterolateral lobe reaching tegula (widely separated from tegula in Duckeia; faintly separated in Amisega), and lacking posterolateral and posteromedial pits (with pits in Duckeia; with only posterolateral pit in Amisega); notaulus deeply impressed, parallel-sided posteriorly (vaguely indicated in Amisega and Duckeia); dorsum of propodeum with undifferentiated metapostnotum (disc of dorsal propodeum with distinguishable metapostnotum in Duckeia and Amisega), and posterior margin simple (delimited posteriorly by transverse groove in Amisega and transverse carina in Duckeia).</p></div>	https://treatment.plazi.org/id/03D187CA7074B86CFC6CFA6FFD706742	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lucena, Daercio A. A.;Melo, Gabriel A. R.	Lucena, Daercio A. A., Melo, Gabriel A. R. (2025): Expanding the Amiseginae: remarkable new Cretaceous-age fossils shed light on the early diversification of a highly specialized clade of cuckoo wasps (Hymenoptera: Chrysididae). Zoological Journal of the Linnean Society 204 (2), DOI: 10.1093/zoolinnean/zlaf054, URL: https://doi.org/10.1093/zoolinnean/zlaf054
03D187CA7079B86EFF42F9B9FAB2609E.text	03D187CA7079B86EFF42F9B9FAB2609E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Miracorium Lucena & Melo 2018	<div><p>Genus † Miracorium Lucena &amp; Melo, 2018</p><p>Type species: † Miracorium tetrafoveolatum Lucena &amp; Melo, 2018 .</p><p>Included species: † Miracorium amica sp. nov. and † M. tetrafoveolatum Lucena &amp; Melo, 2018 .</p><p>Amended diagnosis</p><p>The genus was described originally from male characters only. Herein, we described the first female and extend the generic diagnosis to include sexually dimorphic characters exclusive to females as the following: scapal basin well excavated, borders smoothly round, without carina or sharp edges; head lacking occipital carina; pronotum shorter than scutum in mid-line, gradually declivitous anteriorly, crossed transversely by discrete anterior groove, anterior margin of pronotal flange projecting backwards, posterior margin concave, without transverse groove, posterolateral lobe well developed and reaching tegula, lacking posterolateral and posteromedial pits; labial and maxillary palpomeres with three and four articles, respectively, in females (three and five articles in males); female metanotum 0.8 × as long as scutellum along mid-line, anterior border deepened transversely with medial septum, lateral margin depressed, disc evenly raised (male metanotum as long as scutellum along mid-line, with an anterior pair of medial deep foveae followed by a pair of even larger and deeper posterolateral foveae); dorsal surface of propodeum short, shelf-like, with some distinct enclosures of irregular size, posteriorly bordered by transverse carina, disc slightly depressed; posterolateral corner produced into pointed, short process; posterior surface suddenly sloped, medially crossed by longitudinal median carina; forewing with broad costal cell; tarsal claw simple.</p><p>† Miracorium amica sp. nov.</p><p>(Fig. 7)</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: 72A6 27D3-6749-4D5C-A560-88F6E2D73992</p><p>Description</p><p>Holotype female (DZUP Bur-564). Body length, ~ 2.9 mm. Head: Pyriform, front not protruding, somewhat flattened; frons with discrete medial longitudinal groove, briefly interrupted below; eye bulging, occupying much of lateral head; upper limit surpassing posterior margin of lateral ocellus, inner margin converging on upper frons, continuing parallel dorsally at ocellar area; LID 0.8 × eye width; gena narrow; maxillary palpomeres with four countable articles on right side; malar space ~0.4 × eye height, crossed by longitudinal groove; ocelli inserted smoothly, without well-marked small sulci around them, lateral ocelli closer to inner eye margin and median ocellus than to each other; occipital carina indistinct, apparently absent; scapal basin well excavated, borders smoothly round, without carina or sharp edges; scape length 4.6 × maximum width; flagellum fusiform, with intermediate articles broader than long, F1 longer than combined length of F2 + F3, F4–F6 progressively shortened, F4–F6 with ventral flattened areas; inner margin of antennal sockets touching; mandible apparently thin and delicate; integument finely punctulate, denser on upper frons and vertex; with short, suberect setation; eye glabrous.</p><p>Mesosoma: Convex, not flattened dorsally, shortened anteroposteriorly; pronotum relatively short, subequal with scutum in mid-line, gradually declivitous anteriorly, crossed transversely by discrete anterior groove, anterior margin of pronotal flange projecting backwards, posterior margin concave, without transverse groove, posterolateral lobe reaching tegula, lacking posterolateral and posteromedial pits; notaulus deeply impressed, parallel sided posteriorly, parapsidal line faintly marked posteriorly; scutellum slightly longer than half length of scutum in mid-line, evenly convex, simple; mesopleuron evenly convex, without transverse sulcus, anterior border lacking carina; metanotum 0.8 × scutellum in mid-line, anterior border deepened transversely, with medial septum, lateral margin depressed, disc evenly raised; metapleural carina distinguishable, crossing laterodorsal propodeal surface; propodeum with distinguishable lateral, dorsal, and posterior surfaces; dorsum short, shelf-like, posteriorly limited by transverse carina, with some distinct enclosures of irregular size, disc slightly depressed; posterolateral corner produced into a short process; posterior surface suddenly sloped, medially crossed by longitudinal median carina; tarsal claw simple; metabasitarsus without two rows of spines; dorsum of metafemur evenly convex, without anterior expansion; costal cell broad, first abscissa of M diverging slightly before 1cu-a, gently and smoothly curved submedially; Rs+M arising apically on M; medial cell indicated only by spectral veins; R1 as long as stigma; Rs long, smoothly curved, nearly reaching costal margin; wing membrane entirely setose; dorsal sclerites with distinct white bands, particularly on pronotum, scutum, and scutellum.</p><p>Metasoma: Four exposed terga, internalized segments partly exposed, composing a thin, delicate, metasomal tube; lateral terga sharp, without carinae delimiting the lateral fold; sterna convex and undivided medially.</p><p>Type material</p><p>Holotype female in amber piece DZUP Bur-564. The specimen is well preserved and intact. Syninclusion: a spider.</p><p>Etymology</p><p>From Latin meaning ‘female’. The word is a feminine noun and is treated here as a noun in apposition to the genus name.</p><p>Remarks</p><p>The new species undoubtedly belongs to † Miracorium based on several characters, particularly wing venation, simple tarsal claw, and structure of mesosomal sclerites. A few other distinctive characters prevent us from treating it as the opposite sex of † M. tetrafoveolatum . The new species and † M. tetrafoveolatum exhibit substantial differences in metanotal sculpturing (with four large and deeply impressed foveae in the male of † M. tetrafoveolatum, which are lacking in † M. amica), the general shape of the head (rather pyriform and frontally flattened in the female herein described), sculpturing of the propodeum, and several sexually dimorphic characters, as follows: scape (longer than 4 × its maximum width in † M. amica, and ~2.5–3.5 × longer than its maximum width in † M. tetrafoveolatum), malar sulcus (present in † M. amica, indistinct or absent in † M. tetrafoveolatum), sulcus along medial line of frons (present in † M. amica, absent in † M. tetrafoveolatum), mandible (apparently lacking subapical tooth in † M. amica, with two subapical teeth in † M. tetrafoveolatum), maxillary palpomeres (with four articles in † M. amica, five in † M. tetrafoveolatum), anteromedial portion of pronotum (crossed transversely by groove in † M. amica, lacking groove in † M. tetrafoveolatum), posterior propodeal angle (with acute process in † M. amica, short and bluntly angulate in † M. tetrafoveolatum). Thus, we believe these morphological differences between the sexes indicate that they do belong to distinct species.</p></div>	https://treatment.plazi.org/id/03D187CA7079B86EFF42F9B9FAB2609E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lucena, Daercio A. A.;Melo, Gabriel A. R.	Lucena, Daercio A. A., Melo, Gabriel A. R. (2025): Expanding the Amiseginae: remarkable new Cretaceous-age fossils shed light on the early diversification of a highly specialized clade of cuckoo wasps (Hymenoptera: Chrysididae). Zoological Journal of the Linnean Society 204 (2), DOI: 10.1093/zoolinnean/zlaf054, URL: https://doi.org/10.1093/zoolinnean/zlaf054
03D187CA707BB868FC16FDE4FA5563BB.text	03D187CA707BB868FC16FDE4FA5563BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protognathosega Lucena & Melo 2025	<div><p>Genus † Protognathosega gen. nov.</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org: act: 9BE4602A-53FC-4423-8832-F02885662B06</p><p>Type species: † Protognathosega depilis sp. nov.</p><p>Diagnosis</p><p>Female mandible straight, strongly tapering apically, with basal tooth on laminar edge; frons longitudinally crossed by medial groove inserted between parallel crests, extending from median ocellus towards scapal basin; pronotum short, slightly shorter than scutum along mid-line, posterior border deeply concave, crossed transversely by anterior and posterior grooves and longitudinally by a medial groove; scutellum crossed longitudinally by a medial groove.</p><p>Description</p><p>Head: Pyriform, with relatively long and bulging front; frons longitudinally crossed by medial groove inserted between parallel crests, extending from median ocellus towards scapal basin, fading gradually below on frons; eye bulging, teardrop-shaped, occupying much of lateral head, labial and maxillary palpomeres with three and five articles, respectively; malar space ~0.3 × eye height, crossed by longitudinal groove; pre-hypostomal foveae absent; ocelli deeply inserted, with well-marked small sulci around them, ocellar area slightly produced on vertex; occipital carina long, extending from dorsal head towards malar area; scapal basin shallowly excavated, disc smooth, borders smoothly round, lacking carina or sharp edges; scape length 3.2 × maximum width; flagellum cylindrical, not distinctly fusiform, progressively shortened from F4 to F10 and with flattened ventral areas on F4–F11; outer contour of malar area distinctly angulate; main axis of mandible straight, strongly tapering apically, with basal tooth on laminar edge.</p><p>Mesosoma: Dorsum not flattened; pronotum relatively short, subequal with scutum in mid-line, posterior border deeply concave, crossed transversely by anterior and posterior grooves and longitudinally by a medial groove (Fig. 8F); lateral depression slightly excavated, posterolateral lobe well separated from tegula, lacking posterolateral and posteromedial pits; notaulus deeply impressed, strongly converging posteriorly; parapsidal line well marked posteriorly; scutellum about half length of scutum in mid-line, with longitudinal medial groove anteromedially; mesopleuron anteriorly bordered by discrete carina, evenly convex, without transverse sulcus; metanotum about one-third of scutellum in mid-line, expanded and slightly depressed laterally, disc undifferentiated; propodeum box-like, with distinguishable lateral, dorsal, and posterior surfaces; dorsum of propodeum as long as high; dorsal and posterior surfaces medially crossed by contiguous longitudinal groove; dorsum and posterior surface delimited laterally by confluent grooves, metapostnotum vaguely distinguishable anteriorly on disc of dorsal propodeum; posterolateral corner round, lacking posterior propodeal projection, posterior surface gradually declivitous; tarsal claw with minute submedial tooth, nearly indistinguishable; dorsum of metafemur evenly convex, without anterior expansion; costal cell narrow, first abscissa of M diverging before 1cu-a, gently and smoothly curved submedially; Rs+M arising apically on M; medial cell indicated only by spectral nebulous veins; R1 distinguishable, apparently shorter than stigma; Rs short, smoothly curved, distal trace fading gradually towards distal margin.</p><p>Metasoma: Four exposed terga, internalized segments partly exposed, composing a very slender, relatively long, and delicate metasomal tube; sterna convex and undivided medially.</p><p>Female: Unknown.</p><p>Etymology</p><p>The name is drawn from Greek ‘proto’ meaning ‘first’, ‘gnathos’ meaning ‘jaw’, and ‘-sega’ from Amisega Cameron. The name is feminine.</p><p>† Protognathosega depilis sp. nov.</p><p>(Fig. 8)</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: E474 5760-8B00-4163-81D9-F2C2A011E1B5</p><p>Description</p><p>Holotype female (DZUP Bur-1154a). Body length, ~ 3.6 mm.</p><p>Head: Upper limit of eye surpassing dorsal margin of lateral ocellus, inner margins strongly convergent on upper frons, continuing parallel until posterior margin of head; LID very narrow, 0.4 × eye width; lower gena 0.3 × eye width, upper gena nearly indistinct; lateral ocelli nearly touching inner margin of eye, closer together than with median ocellus; inner margin of antennal sockets nearly touching; F1 length 2.8 × breadth, subequal to combined length of F2 + F3; F11 evenly round, not acuminate; subantennal distance ~1 × MOD, lower medial margin of clypeus straight; head finely and sparsely punctate, particularly on vertex and below inner eye margin; gena smooth, apparently impunctate.</p><p>Mesosoma: Lower posterior corner of propleuron evenly round; metabasitarsus with distinctive comb of thin, suberect setae, preceding two rows of thick, short spines; dorsum of mesosoma regularly punctate, becoming coarser on scutellum; interspaces slightly rugose, particularly on pronotum and scutum; metanotum and propodeum mostly smooth.</p><p>Metasoma: T1–T3 laterally bordered by confluent strong carinae delimiting the laterotergites; metasomal terga coarsely punctate, becoming denser marginally on T2–T3; interspaces apparently densely rugose.</p><p>Vestiture indistinct, apparently mostly glabrous, except wing membrane and legs. Integument coloration apparently dark metallic green.</p><p>Male: Unknown.</p><p>Type material</p><p>Holotype female in amber piece DZUP Bur-1154a. The specimen is intact and well preserved. The amber matrix contains dense reddish particles that partly obscure the specimen, although most structures can be observed perfectly under strong light.</p><p>Etymology</p><p>The species name is a Latin adjective meaning ‘hairless’. Remarks</p><p>The compact short body of † Protognathosega resembles the extant Isegama Krombein and, to a lesser extent, Perissosega Krombein. Notwithstanding, † Protognathosega is readily distinguished based on the following combination of unique characters, which are absent in Isegama and Perissosega: female mandible with basal tooth on the laminar edge; frons longitudinally crossed by medial groove inserted between parallel crests, extending from median ocellus towards scapal basin; pronotum crossed transversely by anterior and posterior grooves; and scutellum crossed longitudinally by medial groove. Additionally, other combinations of characters found in Isegama and Perissosega are lacking in † Protognathosega as follows: scapal basin densely cross-ridged; disc of metanotum with distinct triangular enclosure; borders of scapal basin margined above and on upper side by a ridge ( Perissosega only); mesopleuron with distinct scrobal sulcus ( Isegama only); posterolateral corner of propodeum produced into long, acute spine ( Perissosega only).</p></div>	https://treatment.plazi.org/id/03D187CA707BB868FC16FDE4FA5563BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lucena, Daercio A. A.;Melo, Gabriel A. R.	Lucena, Daercio A. A., Melo, Gabriel A. R. (2025): Expanding the Amiseginae: remarkable new Cretaceous-age fossils shed light on the early diversification of a highly specialized clade of cuckoo wasps (Hymenoptera: Chrysididae). Zoological Journal of the Linnean Society 204 (2), DOI: 10.1093/zoolinnean/zlaf054, URL: https://doi.org/10.1093/zoolinnean/zlaf054
03D187CA707DB86AFC74FD06FC2D6598.text	03D187CA707DB86AFC74FD06FC2D6598.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Telistosega Lucena & Melo 2025	<div><p>Genus † Telistosega gen. nov.</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: C9 E49B91-6966-4112-A0FC-A4E53DAC17C0</p><p>Type species: † Telistosega intermedia sp. nov.</p><p>Diagnosis</p><p>Head not pyriform, somewhat flattened frontally; maxillary palpomeres with four articles; malar space short, ~0.2 × eye height, without longitudinal sulcus; scape length 4.6 × maximum width; flagellum fusiform, with intermediate articles broader than long; mandible with distinctly expanded basal area on outer surface, becoming strongly thin and tapering apically, main axis straight, slightly bent submedially, lacking subapical teeth on laminar edge; pronotum strongly declivitous anteriorly, posterior margin nearly straight, distinctly swollen and without transverse posterior groove, posterolateral lobe reaching tegula, lacking posterolateral and posteromedial pits; mesopleuron evenly convex, without transverse scrobal sulcus, anterior border lacking carina; dorsum of propodeum very short, shelf-like, posteriorly delimited by transverse carina, disc raised; posterior propodeal projection short, bluntly angulate; tarsal claw with very short, perpendicular, submedial tooth; metasoma with four exposed terga, internalized segments partly exposed, composing a relatively long and robust distal tube (comparatively with other Amiseginae).</p><p>Description</p><p>Head: Not pyriform, somewhat flattened frontally, wider than high; frons lacking medial longitudinal groove; eye bulging, occupying much of lateral head, upper limit surpassing dorsal margin of lateral ocellus, inner margin gently converging at ocellar area; maxillary palpomeres with four countable articles; malar space ~0.2 × eye height, longitudinal groove indistinct, apparently absent; ocelli inserted smoothly, without well-marked small sulci around them; occipital carina short, extended only dorsally; scapal basin shallowly excavated, disc smooth, borders smoothly round, without carina or sharp edges; scape length 4.6 × maximum width; flagellum fusiform, with intermediate articles broader than long, F1 longer than combined length of F2 + F3; mandible with distinctly expanded basal area on outer surface, becoming strongly thin and tapering apically, main axis straight, slightly bent submedially, lacking subapical teeth on laminar edge.</p><p>Mesosoma: Evenly convex, not flattened dorsally, somewhat shortened anteroposteriorly; pronotum about as long as scutum in mid-line, strongly declivitous anteriorly, crossed transversely by anterior groove, posterior margin nearly straight, distinctly swollen and without transverse groove, posterolateral lobe reaching tegula, lacking posterolateral and posteromedial pits; notaulus deeply impressed, parallel-sided posteriorly, parapsidal line well marked posteriorly; scutellum slightly longer than half length of scutum in mid-line, evenly convex, simple; mesopleuron evenly convex, without transverse scrobal sulcus, anterior border lacking carina; metanotum shorter than one-third of scutellum in mid-line, expanded and slightly depressed laterally, disc slightly raised; metapleural carina distinguishable, extending across laterodorsal propodeal surface; propodeum with distinguishable lateral, dorsal, and posterior surfaces; dorsum short, shelf-like, posteriorly delimited by transverse carina, disc raised; posterior propodeal projection short, bluntly angulate; posterior declivity strongly declivitous, apparently crossed by longitudinal medial carina; tarsal claw with very short, perpendicular, submedial tooth; costal cell narrow, first abscissa of M diverging before 1cu-a, gently and smoothly curved submedially; Rs+M arising apically on M; medial cell indicated only by spectral veins; R1 shorter than stigma; Rs short, smoothly curved, distal trace fading gradually towards costal margin.</p><p>Metasoma: Four exposed terga, internalized segments partly exposed, composing a relatively long and robust distal tube (comparatively with other Amiseginae); sterna convex and undivided medially.</p><p>Male: Unknown.</p><p>Etymology</p><p>The name is derived from the Greek telistos, meaning ‘farthest’,</p><p>plus ‘-sega’ from Amisega Cameron. The name is feminine.</p><p>† Telistosega intermedia sp. nov.</p><p>(Fig. 9)</p><p>ZooBank registration: LSID: urn:lsid:zoobank.org:act: D3C96 A8D-A554-406C-8344-0B6FFF753598</p><p>Description</p><p>Holotype female (DZUP Bur-728a). Body length, ~ 2.8 mm.</p><p>Head: LID wide, subequal to eye width; gena narrow, slightly wider basally; lateral ocelli closer to inner eye margin and median ocellus than to each other; inner margin of antennal sockets touching; F4–F10 progressively shortened, F4–F11 with ventral flattened areas, F11 gently acuminate; clypeus with distinctly elongated, erect, thin, gold setation; eye glabrous; dorsum of head finely and densely punctate.</p><p>Estimated years given in millions of years and in accordance with the Paleobiology database (Uhen et al. 2023). Exclusively extinct genera are indicated by daggers.</p><p>aOriginally described in Hedychridium Abeille de Perrin ( Chrysidinae:Elampini).Transferred to Atoposega by Brazidec et al. (2024).</p><p>bDisputed in the cladistic analysis by Lucena and Almeida (2022).</p><p>Mesosoma: Lateral pronotal depression shallowly excavated; metabasitarsus without two rows of spines; dorsum of metafemur evenly convex, without anterior expansion; dorsal sclerites of mesosoma finely and densely punctate; short, sparse, suberect setation on dorsum, becoming longer and denser posterolaterally on propodeum; wing membrane entirely setose.</p><p>Metasoma: Lateral terga broadly round, without carinae delimiting the lateral fold.</p><p>Male: Unknown.</p><p>Etymology</p><p>The species name is a Latin adjective meaning ‘intermediate’.</p><p>Type material</p><p>Holotype female in amber piece DZUP Bur-728a. The specimen is intact and well preserved. Some cracks in the amber piece prevent the lateral left view of the specimen.</p><p>Remarks</p><p>† Telistosega most resembles † Miracorium (the first female of this genus is described above), based on the general shape and structure of pronotum, frons not bulging, dorsum of propodeum short, shelf-like, posterior propodeal projection short, bluntly angulate, metabasitarsus lacking two rows of spines, and tarsal claw with small, perpendicular, submedial tooth. The new genus is readily distinguished from † Miracorium based on the narrow costal cell (distinctly broad in † Miracorium; for comparison of this feature, see Fig. 7A, C; see also Lucena and Melo 2018: their fig.3; Lucena and Almeida 2022: their figs 15 and S22, S23); head wider than high, with broad face (pyriform in † Miracorium); occipital carina extended dorsally (lacking in † Miracorium); and short and simple malar space (moderately long and longitudinally crossed by sulcus in † Miracorium). Additionally, the modified pronotum, strongly declivitous anteriorly and with swollen posterior border, combined with the above-mentioned character states, readily distinguish † Telistosega from all other genera of Amiseginae s.l.</p></div>	https://treatment.plazi.org/id/03D187CA707DB86AFC74FD06FC2D6598	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lucena, Daercio A. A.;Melo, Gabriel A. R.	Lucena, Daercio A. A., Melo, Gabriel A. R. (2025): Expanding the Amiseginae: remarkable new Cretaceous-age fossils shed light on the early diversification of a highly specialized clade of cuckoo wasps (Hymenoptera: Chrysididae). Zoological Journal of the Linnean Society 204 (2), DOI: 10.1093/zoolinnean/zlaf054, URL: https://doi.org/10.1093/zoolinnean/zlaf054
