taxonID	type	description	language	source
03DEE569FFE9FFFE1932FDD25D66FA44.taxon	distribution	Distribution — Tropics and subtropics of the world. Most species of the genus Mucuna belong to this subgenus. Note — Only some specimens of M. mollissima have ± crenate leaflets with nerves ± ending in the margin.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE9FFFE1A7DFF015B3CFCAF.taxon	description	Macranthus Lour. (1790) 461, ‘ Marcanthus ’. — Type: Mucuna cochinchinensis Lour. Lianas or ± herbaceous climbers, very rarely treelets (M. stans). Leaflets ± crenate, secondary nerves ending in or anastomosing at the margin. All anthers glabrous. Ovary S-shaped. Pods straight to S-shaped, valves longitudinally ribbed or not. Seeds rather small, ± bean-shaped; hilum 4 – 7 mm long, 1 / 8 – 1 / 3 of the circumference, with rim aril. Distribution — Tropics and subtropics of the world. Mucuna pruriens var. utilis is widely cultivated. In Africa the subgenus is represented by: M. coriacea Baker, M. ferox Verdc., M. glabrialata (Hauman) Verdc., M. melanocarpa Hochst. ex A. Rich., M. poggei Taub., M. pruriens (L.) DC., M. stans Welw.; in Asia by: M. bracteata Roxb. ex Kurz, M. diabolica, M. papuana Adema, M. pruriens, M. sericophylla Perkins; in Australia by: M. diabolica subsp. kenneally Verdc., M. reptans Verdc.; and in the Americas by: M. pruriens.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE6FFF11932FF7A5DE8FCCE.taxon	distribution	Distribution — Malesia: Peninsular Malaysia; Singapore; Sumatra; Java; Lesser Sunda Islands: Bali, Flores. Habitat & Ecology — Primary and secondary forest, water- sides, shrubbery, hedges, edges of sugarcane fields. Altitude up to 1400 m. Flowering: January, February, July, August, December; fruiting: January, February, May to August. Note — The inflorescence is basically a pseudoraceme with (few) brachyblasts clustered at the apex of the rachis with 3 flowers at the apex of the brachyblasts. The whole looks like an umbel. The calyx is in bud more or less cylindrical or urnshaped, when the corolla starts to expand the calyx widens and gets pushed backwards, finally the calyx is campanulate and tucked up. Ripe pods are mostly glabrous except for the wings, the irritant hairs are present at the wings for a long time.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE6FFF11932FCCF5A01FE53.taxon	description	Twigs ferruginous tomentose. Petioles 4 – 8 cm long, rachis 1 – 3 cm long. Leaflets elliptic or ovate, 6.5 – 15 by 4 – 9.5 cm, above ferruginous sericeous to velutinous. Inflorescences pseudoracemes, 15 – 16 cm long. Calyx campanulate, 11 – 12 mm long, tube c. 5 mm long. Pods 18 by 3 cm, wings 12 – 25 mm wide, valves with irregular, transverse lamellae. Seeds 20 – 25 by 18 – 27 by 10 – 12.4 mm. — Type: ANU 2606 (Flenley) (holo L; iso LAE), W Highlands Prov., Wabag, Ecological site 11, near Yogonda, 1 / 4 mile W of R. Lai. Liana to 4 m long. Twigs terete, 3 – 5 mm diam, ferruginous sericeous to velutinous (see Note). Stipules narrowly ovate, c. 21 by 2.5 mm, outside sericeous, inside glabrous. Petioles 4 – 8 cm long, terete, ferruginous sericeous; rachis mostly as the petiole 1 – 3 cm long; pulvinus 8 – 13 mm long. Stipellae acicular, 4 – 6 by 0.2 – 0.3 mm, hirsute or ± sericeous. Leaflets: terminal elliptic or ovate, 6.5 – 15 by 4 – 9.5 cm, index 1.3 – 1.9, base acute to rounded, apex acuminate, acumen 6 – 11 mm long, above sericeous, below sericeous to velutinous, midrib and nerves slightly raised above, nerves 5 – 7 per side, 8 – 30 mm apart, anastomosing near the margin; lateral mostly as the terminal 7 – 13 by 4 – 6.5 cm; pulvinus 5 – 8 mm long. Inflorescences axillary, pseudoracemes, 15 – 16 cm long, peduncle 13.5 cm long, ferruginous sericeous to velutinous. Bracts to the brachyblasts ovate, 20 – 25 by 5 – 11 mm, outside sericeous, inside thinly sericeous. Brachyblasts in fruit 2.5 – 3 mm long. Bracts to the flowers narrowly ovate, 20 – 26 by 5 – 10 mm, outside sericeous, inside thinly sericeous. Pedicels c. 2 cm long (in fruit). Bracteoles narrowly ovate, 20 – 21 by 2.5 – 3 mm, both sides sericeous. Calyx campanulate, 11 – 12 mm long, tube c. 5 mm long; upper lip triangular, 3 mm long, lateral teeth triangular, 5 mm long, median tooth triangular, 6 – 7 mm long; outside pubescent and with scattered irritating hairs. Corolla (only immature ones seen). Pods flattened ellipsoid, 18 by 3 cm, sericeous and with irritating hairs, sutures winged, upper wing 15 – 25 mm wide, lower one 12 mm wide, with irregular transverse lamellae, lamellae interrupted, overlapping in the middle, 8 – 10 mm high. Seeds flattened ovoid or discoid, 20 – 25 by 18 – 27 by 10 – 12.4 mm; hilum 32 – 33 mm long, 0.45 of the circumference. Distribution — Malesia: Papua New Guinea: W Highlands Prov. Habitat & Ecology — Mountain rain forest, with good drain- age. Altitude 2750 – 2900 m. Flowering September; fruiting March. Uses — Seeds eaten. Specimens seen. PAPUA NEW GUINEA, W Highlands Prov., Wabag, Ecological site 11, near Yogonda, 1 / 4 mile W of R. Lai, alt. 2750 m, ANU 2606 (Flenley), 11 Mar. 1965; W Highlands Prov., near Sirunki, alt. 2900 m, ANU 909 (Walker), 19 Sept. 1962; E Highlands Prov., Kainantu Subprov., Aiyura, alt. 1500 m, NGF 19030 (Womersley), 8 Sept. 1963. Note — Hairs tend to be patent, longest ones 1 – 2.5 mm long. In vegetative characters M. aimun resembles M. mollissima, M. platyphylla, M. tomentosa and M. verdcourtii. From these species it differs in pod and seed characters.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE6FFF31A7DFE6F5D79FD2B.taxon	description	Liana up to 8 m high. Leaflets narrowly ovate, rarely elliptic, 5.7 – 11.6 by 1.3 – 3.9 cm, index 2.6 – 4.4. Inflorescences axillary, rarely terminal, pseudoracemes, 90 – 105 cm long, peduncle 75 – 84 cm long. Pedicels 30 – 35 mm long. Keel petals 44 – 46 mm long. Ovules 3 – 4. — Type: Ridsdale & Reynoso 1420 (holo L), Philippines, Luzon, Zambales, Santa Cruz, Acoje Mine concession area, ultrabasic, 23 May 1986. Liana up to 8 m. Twigs terete, striate, 1 – 3 mm diam, glabrous to very thinly sericeous. Stipules narrowly elliptic, 3 – 4 by 0.6 – 0.7 mm, outside with few appressed hairs, inside glabrous, caducous. Petioles 3 – 6 cm long, ± grooved, glabrous or with few scattered appressed hairs; rachis mostly as the petiole, 0.8 – 2 cm long; pulvinus 4 – 8 mm long. Stipellae acicular, 2.4 – 4.8 by 0.1 – 0.3 mm, glabrous or with few appressed hairs. Leaflets: terminal narrowly ovate, rarely elliptic, 5.8 – 11.6 by 1.3 – 3.9 cm, index 2.6 – 4.4, base cuneate or rounded, apex acuminate, acumen 3 – 15 mm long, above glabrous or with few appressed hairs, below glabrous or with few appressed hairs mainly at midrib, midrib and nerves raised above, nerves 4 – 7 per side, 3 – 31 mm apart, anastomosing near the margin; lateral mostly as the terminal, obliquely narrowly ovate, 5.7 – 11.6 by 1.4 – 4 cm; pulvinus 2 – 6 mm long. Inflorescences axillary, rarely terminal, pseudoracemes, 90 – 105 cm long, peduncle 75 – 84 cm long, peduncle mostly glabrous, upwards sericeous, flowering part densely sericeous. Bracts to the brachyblasts ovate, 32 – 37 by 15 – 20 mm, outside (thinly) sericeous, inside thinly sericeous, caducous. Brachyblasts 2 – 9 mm long, sericeous. Bracts to the flowers ovate or elliptic, 19 – 22 by 6 – 11 mm, both sides thinly sericeous, caducous. Pedicels 30 – 35 mm long, sericeous. Bracteoles elliptic or narrowly obovate, 17.5 – 22 by 2.5 – 6 mm, both sides thinly sericeous, caducous. Calyx 14 mm long, tube 7 – 10 mm long; upper lip triangular or semicircular, 2 – 14 by 14 – 34 mm, lateral teeth triangular, 2.5 – 5 by 4 – 6 mm, median tooth triangular, 5 – 7 by 6 – 7 mm; outside sericeous and with irritating hairs, inside sericeous. Corolla pale green. Standard: claw 3 – 4 mm long, glabrous; blade broadly ovate or broadly elliptic, 22 – 26 by 18 – 23 mm, bidentate, auricle 1 mm long, outside glabrous or with few appressed hairs in basal part, inside glabrous. Wings: claw 5 – 6 mm long, outside sericeous along both margins, ciliate along both margins, inside sericeous along both margins; blade elliptic, 35 – 38 by 8 – 13 mm, rounded, auricles 3 mm long, lateral pocket 6 – 8 mm long, outside sericeous between claw and auricle, at auricle and just above, ciliate along lower margin in lower 1 / 4 – 1 / 3, inside glabrous. Keel petals: claw 7 – 8 mm long, glabrous; blade ± boatshaped, 37 – 38 by 8 – 12 mm, acute, auricles 1 mm long, lateral pocket 7 – 8 mm long, hard part 7 mm long, both sides glabrous, short-ciliate along upper margin. Stamens 44 – 45 mm long, tube 30 – 37 mm long, free part of filaments below versatile anthers 8 – 10 mm long, below basifixed ones 7 – 8 mm long, glabrous; versatile anthers 1.5 by 0.5 mm, with few hairs at the base, basifixed anthers 2.6 by 0.4 mm, outside with some appressed hairs in basal part. Disc 0.8 – 0.9 mm high, glabrous. Ovary 6 – 8 mm long, sericeous, stipe c. 1 mm long, sericeous; ovules 3 – 4; style 42 – 43 mm long, sericeous at base, thinning upwards, apical part glabrous. Pods flattened ellipsoid, 13 by 3 cm, upper wing 5 mm wide, lower wing 3 mm wide, lamellae oblique, 2 mm high, valves ± puberulous and with irritating hairs. Seeds discoid, 16 by 15 by 8.1 mm; hilum 41 mm long, c. 3 / 4 of the circumference. Distribution — Malesia: Philippines: Luzon, Zambales Prov. Habitat & Ecology — Secondary forest. Soil: ultrabasic. Altitude up to 200 m. Flowering: May, November; fruiting May. Specimens seen. PHILIPPINES, Luzon, Zambales Prov., Mansinlok Mine, on road to Bontak, Argent et al. 99290; Luzon, Zambales Prov., Santa Cruz, Acoje Mine concession area, Ridsdale & Reynoso 1420, 1546. Note — In many aspects rather similar to M. curranii. The new species differs especially in the narrower leaflets (index: 2.6 – 4.4, in M. curranii 2.1 – 2.2); the inflorescence with a longer peduncle, longer pedicels and smaller flowers (44 – 66 mm long, in M. curranii 69 – 80 mm long), the indumentum of the corolla parts and the number of ovules (3 – 4, in M. curranii 4 – 7). According to the fields notes the colour of the corolla is pale green in the new species and whitish in M. curranii. The new species is recorded from ultrabasic at c. 200 m, while M. curranii is found on limestone at 1300 – 2200 m.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE4FFF31932FCA65D8EFABD.taxon	description	Mucuna aurea C. B. Rob. (1908) 183; Merr. (1910) 118; (1923) 307; Wilmot-Dear (1991 b) 229. — Type: Williams 1292 (holo NY; iso K, PNH, US), Philippines, Luzon, Prov. Benguet, Baguio. Distribution — Malesia: Philippines: Luzon. Habitat & Ecology — Thickets. Altitude c. 1650 m. Flowering: July, November, December; fruiting: March. Note — This species is remarkable in its golden yellowish calyx indumentum. H. C. Conklin & Buwaya (PNH 79648) give the flower colour as red, however, the duplicates of this specimen in K and L lack flowers. Mucuna aurea is very similar to M. platyphylla differing mainly in the somewhat smaller flowers and slightly larger pods. The flower colour of M. aurea is recorded as red, while M. platyphylla has white to green or yellowish flowers. Mucuna platyphylla is up to now not found in the Philippines. According to the original description the flowers occur singly or in pairs. However, the few specimens seen all have brachyblasts with three scars, so presumably there have been three flowers, as is normally found in Mucuna.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE4FFF31932FADC5A1CFE52.taxon	description	Mucuna bennettii F. Muell. (1876) 63; Verdc. (1979) 439; Wilmot-Dear (1990) 32. — Type: D’Albertis s. n. (A n. v.), Papua New Guinea, Fly river. Mucuna miniata Merr. (1917 a) 278; [Parrana miniata Rumph. (1747) 10]. — Type: Robinson Pl. Rumph. Ambon. 566 (A, K, L, NY), Moluccas, Ambon, between Paso and Roema Tiga. Mucuna elegans Merr. & L. M. Perry (1942) 406; Verdc. (1979) 442. — Type: Brass 2734 (BISH, L), Solomon Isl., San Cristoval Isl., Magona river. Mucuna warburgii auct. non K. Schum. & Lauterb.: Verdc. (1979) 457, p. p.; Wilmot-Dear (1992) 243. Distribution — Malesia: Celebes; Moluccas; New Guinea; Solomon Islands; New Hebrides: Vanuatu. Habitat & Ecology — Primary or secondary, or depleted forests, often along rivers, swamp forest, sago swamps, fresh water swamp behind the beach. Altitude up to 400 m. Soil: limestone, clay. Flowering: April to October, December; fruiting: March, April, October. Note — Brass 5724, a flowering specimen, was collected between January and March. Calyx in bud ± cylindrical, when the flower expand it becomes campanulate, but not tucked up. Three fruiting specimens have been included in the present description: Van Balgooy 6500 from Aru Islands, fruit collected from underneath a flowering specimen of M. bennettii; Prawiroadmodjo & Maskuri 1541 from Celebes, originally identified as M. elegans; Sands 1211 from PNG, W Sepik Prov. The fruits of these specimens are all rather similar differing only in length. The vegetative parts of the last two specimens agree very well with the description of M. bennettii. Jeswiet 145: Pedicels sometimes swollen, probably some kind of gall. Specimens from the Moluccas (originally M. miniata) and the Solomon Islands (originally M. elegans) have usually slightly smaller flowers. In other characters no differences were found. Mucuna warburgii was described with stipellae. Here we include the specimens identified as M. warburgii, but lacking stipellae.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE4FFF31A7DFE6C5BF8FB2E.taxon	distribution	Distribution — Malesia: Sumatra; Peninsular Malaysia; Borneo. Habitat & Ecology — Primary, secondary or disturbed forests, often along rivers, riverbanks, belukar. Soil: limestone, red soil, yellow sandy clay, loam, sand. Altitude up to 600 m. Flowering: January, February, May to August, October, December; fruiting: January, February, April, June to August, November, December. Uses — Exudate is used for healing cuts and wounds; in- ternally it is used for curing diarrhoea. Note — The horizontal part of the lamellae is 1 – 4 mm wide. Vegetatively resembling M. toppingii. However, M. biplicata differs from the latter in the inflorescences and fruits. Also closely resembling M. acuminata. From this species it differs in the obscure calyx teeth and the oblique bifurcate lamellae interrupted in the middle. Mucuna acuminata has developed calyx teeth and ± smooth pods. LÖrzing 12877: One of the pods with lamellae and wings broadly fimbriate, lamellae not bifurcate. The specimen SAN 81200 probably belongs here, however, the label gives the flower colour as ‘ pinkish’.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE4FFF31A7DFAAA5B99F881.taxon	description	Mucuna brachycarpa Rech. (1913) 562; Verdc. (1978 a) 459; (1979) 439, pl. A, f. 106 C; Wilmot-Dear (1990) 11. — Type: Rechinger 4807 (W n. v.), Bougainville, Kieta. Distribution — Malesia: Papua New Guinea: Bougainville; Solomon Islands; Fiji. Habitat & Ecology — Old secondary forest, along creeks, in swamp forest. Altitude up to 600 m. Flowering: April to November. Note — The original description was based on a single pod. According to Verdcourt (1979) the common yellow-flowered Mucuna of the Solomons most probably belongs to the same species. Merrill & Perry (1942: 405) associated the pod with a specimen collected by Brass on Bougainville (Brass 3514 n. v.). This specimen is the type of M. subumbellata. Wilmot-Dear (1990) notes that two specimens from Santa Ysabel (BSIP 2245) and Guadalcanal (R 55613) should be included here. Verdcourt (1978 a: 459) mentioned these specimens as ‘ Sp. C’. The description of the pods and seeds were taken from the description by Wilmot-Dear (1990).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFE4FFEC1A7DF83F5D6EFCE1.taxon	distribution	Distribution — India, Burma, China (Yunnan), Thailand, Laos, Vietnam; Malesia: Sumatra. Habitat & Ecology — Forest, thickets, open grasslands, along paths and streams. Altitude 600 – 2000 m. Flowering: January, July, November, December; fruiting: February, May, October. Note — Wilmot-Dear (1984) gives 2 flowers per brachyblast. However, the structure of the inflorescence is a bit more com- plicated. Often there are two or three flowers at the top of a brachyblast, but sometimes the brachyblasts are slightly longer with one flower at the base and two or three flowers at the top of a very short lateral branch. The whole inflorescence than looks a bit like a mix of a pseudoraceme and a pseudopanicle. According to Wilmot-Dear (1984) De Candolle is the author of the epithet of this species. However, De Candolle (1825: 406) did not describe or name the species. He only placed Roxburgh’s name for an Indian plant in Mucuna. Kurz (1873) was the first to describe this species attributing the species name to Roxburgh. Roxburgh (1814) cites as ‘ donor’ of the specimen J. R. (= James Roxburgh) and as place of collection Chittagong. Bracts and bracteoles are mostly caducous, however, some of the basal sterile bracts may be still present when fruiting. Young pods are ± S-shaped, mature pods are nearly straight.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFBFFEC1932FCE25D32FB0F.taxon	description	Mucuna canaliculata Verdc. (1978 a) 460; (1979) 440. — Type: NGF 35864 (Streimann) (BULOLO, CANB, K, L, LAE), Papua New Guinea, Morobe Prov., Wau Subprov., Upper Watut, Minnoa Creek. Distribution — Malesia: Papua New Guinea: Morobe, W Highlands, Western Prov. Habitat & Ecology — Foothill, montane or Castanopsis forest, swamp forest. Altitude (3 m, see Note) 1000 – 1600 m. Flowering: April, November; fruiting: June, September, October. Note — The indumentum of the pods mainly consists of irritating hairs. The specimen NGF 18443 from Western Prov. was collected at an altitude of 3 m much lower than all other specimens. NGF 22604 from Morobe Prov. lacks stipellae and differs slightly in the size of the flower parts.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFBFFEC1932FA8C5EB0F8D2.taxon	distribution	Distribution — Malesia: Philippines: Luzon. Habitat & Ecology — Soil: limestone. Altitude 1600 m. Flowering: January to March, December; fruiting: March to July. Altitude 1300 – 2200 m. According to Merrill (1923: 308) in ravines and thickets. Note — Closely resembling M. longipedunculata from which it differs in pod characters: flattened with wings along the sutures and oblique lamellae in M. curranii, almost cylindrical, without wings and lamellae in M. longipedunculata. The type specimen (Elmer 8442) is probably a fruiting specimen. To the Kew and Leiden duplicates some (dissected) flowers are added, may be from another specimen.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFBFFEC1932F8EE5A8FFD8C.taxon	distribution	Distribution — Malesia: Java; Celebes; Lesser Sunda Islands: Sumbawa, Flores, Lombok, Alor, Timor. Habitat & Ecology — Forest, monsoon forest, shrubbery, village margin. Soil: clay, loam. Altitude 150 – 1200 m. Flowering: March, May to July; fruiting: June, August. Note — Young pods are ± S-shaped, mature ones are nearly straight. When Backer described M. diabolica he was unaware of the earlier published name Stizolobium forbesii Piper. Later he probably found this name and in 1945 he made a new combination for the species as M. forbesii (Piper) Backer. However, he clearly missed the name M. forbesii Baker f. (see M. platyphylla) which was published in 1923. At the moment Backer transferred Stizolobium forbesii Piper to Mucuna there was already M. forbesii Baker f. blocking the use of Piper’s epithet in Mucuna and the species has to be called M. diabolica. In Backer’s original description of M. diabolica no specimens were cited. In L there is only one specimen collected by Backer on Java available. This specimen is selected as neotype.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFBFFEC1A7DFD0F5A07FB5F.taxon	description	Tussac. — Neotype (here designated): Merrill Sp. Blanc. 779 (holo L; iso K), Luzon, Rizal Prov., San Mateo. Mucuna monosperma auct. non Wight: Fern. - Vill. (1880) 63. Mucuna imbricata auct. non Baker: Merr. (1905 a) 38; (1906) 67. Mucuna nigricans auct. non (Lour.) Steud.: Merr. (1910) 116; (1918) 187. Distribution — Malesia: Philippines: Luzon, Polillo, Mindoro. Habitat & Ecology — Forest, riverside regrowth. Altitude up to 70 m. Flowering: January, December; fruiting: February. Note — According to Wilmot-Dear (1991 b) the patch of hairs at the outside of the standard reaches higher than the upper lip of the calyx. The L duplicate of Merrill Spec. Blanc. 779 has only some scattered hairs visible above the calyx rim. The horizontal part of the lamellae is 2 mm wide.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFBFFEC1A7DFB7A5CFBF9D1.taxon	description	Mucuna discolor Merr. & L. M. Perry (1942) 405; Verdc. (1979) 442. — Type: Brass 3901 (holo A?; iso BO, NY), Papua New Guinea, Dieni, Ononge road. Distribution — Malesia: Papua New Guinea: Central Prov. Habitat & Ecology — Secondary bushes, roadsides. Flowering: April. Note — Only known from the type collection. According to the label the leaflets are purple below. The flowering part of the inflorescences is rather short, in the shortest inflorescence ± ‘ pseudo-umbellate’.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFBFFEC1A7DF9F15B7CF7F9.taxon	description	Mucuna elmeri Merr. (1929) 108. — Type: Elmer 20416 (C, K, L, NY, P), Borneo, Sabah, near Tawao. Mucuna monosperma auct. non Wight: Miq. (1855) 214; Craib (1928) 44, p. p. Distribution — Malesia: Borneo. Habitat & Ecology — Primary or disturbed forest along rivers. Altitude up to 1000 m. Flowering: February, May, June, August, October; fruiting: January, February, June, September, October. Uses — The sap is used against mouth ulcers. Note — Distinct from most other Mucuna species by having only 1 ovule and 1 seed. In the upper part of the valves of the pods the lamellae are often inconspicuous or even absent. S 46210 (Yii & Othman) probably belongs here, however, the flower colour is given as yellow.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFAFFEE1932FF015E62FE22.taxon	description	Twigs tomentose. Petioles 23 – 45 mm long, tomentose, rachis 10 – 18 mm long, tomentose. Longest hairs on twigs 1.0 – 2.0 mm long, on petiole and rachis 2.5 mm long. Pods c. 9.5 by 2.5 cm, upper wing 15 – 20 mm wide, lower wing 14 – 17 mm wide, lamellae oblique c. 22 mm high. Seeds ± globular, 18 – 20 by 19 – 20 by 14.4 mm. — Type: LAE 60525 (Croft & Lelean) (holo L; iso K), Papua New Guinea, Central Prov., Port Moresby subprov., SW slope Lake Myola No. 1, 1900 m. Liana up to 3.5 m. Twigs terete, 2 – 3 mm diam, tomentose (see Note). Stipules ovate, c. 7 by 3 mm, outside sericeous, inside glabrous, caducous. Petioles 23 – 45 mm long, ± terete, tomentose (see Note); rachis mostly as the petiole, 10 – 18 mm long; pulvinus 4 – 8 mm long. Stipellae acicular, 6 – 8 by 0.3 – 0.4 mm, ± sericeous. Leaflets: terminal broadly elliptic, obovate or ± orbicular, 6 – 10 by 4.5 – 6.5 cm, index 1.2 – 1.4, base rounded, apex acuminate, acumen 3 – 6 mm long, above with scattered appressed hairs, midrib and nerves sericeous, below velutinous, midrib and nerves sericeous, sometimes with some very long hairs, midrib raised above, nerves flat or raised above, 3 – 5 per side, 9 – 25 mm apart, anastomosing close to the margin; lateral mostly as the terminal, obliquely ovate, 4.5 – 8.0 by 3.0 – 5.0 cm, index 1.5 – 1.7; pulvinus 4 – 5 mm long. Inflorescences axillary, pseudoracemes, 2 – 12 cm long, peduncle 0.5 – 2.5 cm long, tomentose. Brachyblasts in fruit c. 15 mm long. Bracts to the flowers ovate, c. 16 by 8 mm, outside sericeous and with some irritating hairs, inside with some hairs at the base. Pedicel in fruit c. 23 mm long. Bracteoles narrowly ovate, c. 14 by 5 mm, outside sericeous, inside glabrous. Corolla greenish white. Pods flattened ellipsoid, c. 9.5 by 2.5 cm, stipe 5 mm long, upper wing 15 – 20 mm wide, lower wing 14 – 17 mm wide, lamellae oblique, c. 22 mm high, valves puberulous and with many irritating hairs. Seeds ± globular, 18 – 20 by 19 – 20 by 14.4 mm; hilum 46 – 49 mm long, c. 4 / 5 of the circumference. Distribution — Malesia: Papua New Guinea: Central Prov. Habitat & Ecology — Secondary growth, edge of forest and grassland. Altitude 560 – 2100 m. Flowering: January, June; fruiting: October. Specimens seen. PAPUA NEW GUINEA, Central prov., Uniri river, c. 1500 ft, Carr 15251, 26 Jan. 1930; Central Prov., Port Moresby Subprov., SW slope Lake Myola No. 1, 1900 m, LAE 60525 (Croft & Lelean), 1 Oct. 1973; Central Prov., Myola, near airstrip, 1080 m, Hopkins & Hopkins 1017, 19 May 1989. Note — Longest hairs on twigs 1.0 – 2.0 mm long, on petiole and rachis 2.5 mm long.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF9FFEE1932FE5E5FBCFCAB.taxon	description	[Lobus litoralis Rumph. (1747) 10, t. 6.] See Merr. (1917 a) 277. Mucuna gigantea (Willd.) DC. subsp. plurisema Verdc. (1978 b) 126; (1979) 444; Wilmot-Dear (1991 b) 218. — Type: LAE 51631 (Streimann & Kairo) (holo LAE; iso A, BO, BRI, CANB, K, L, NSW, SING), Papua New Guinea, Central Prov., km 27 Port Moresby-Sogeri road, sec. forest by rim in savannah.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF9FFEE1932FC265E6DF82D.taxon	distribution	Distribution — Africa, Seychelles, India, Burma, China, Japan, Indochina; throughout Malesia; Australia, Solomon Islands, Hawaii, Tahiti, Niue Island, Cook Island, Mariana Islands, New Hebrides, New Caledonia. Habitat & Ecology — Beaches, sand dunes, coastal forest, primary and secondary forest, along rivers and roads. Soil: volcanic ash, sand, limestone. Altitude up to 700 m. Flowering and fruiting: throughout the year. Note — Pods often have a stipe up to 10 mm long. BNBFD 7393 (Keith) probably belongs here. According to the label the flower colour is pink. Several times M. gigantea was split into subspecies. However, the authors who proposed these splits use different characters, making it difficult to compare these subspecies. Verdcourt (1979) uses fruit characters (size of fruits and number of seeds), while Ohashi & Tateishi (1976) and Tateishi & Ohashi (1981) use flower characters (size of standard in relation to size of wings, length of the claw of wing petals in relation to the whole length of wing petals). In the Flora Malesiana region the subspecies described by Verdcourt are found: subsp. gigantea and subsp. plurisema. According to Verdcourt the differences between these subspecies are: subsp. gigantea: Flowers small. Fruits 10 – 18 by 3.5 – 6 cm with 1 – 4 seeds; subsp. plurisema: Flowers larger (up to 4 cm). Fruits 15.5 by 3.5 – 4 cm with 5 – 6 seeds. However, the flower-size in subsp. gigantea varies from 27 mm to 53 mm, so flower-size is not a good character. Pods of subsp. gigantea measure 8 – 18 by 3 – 6.5 cm. In this character there is no difference at all; the size of the pods of subsp. plurisema falls right in the range of subsp. gigantea. In specimens of subsp. gigantea pods may be found with 4 – 5 seeds. In the ovary up to six ovules may be found, so 6 seeds may be expected. To conclude: the differences as given by Verdcourt are insufficient to support subspecies. According to Ohashi & Tateishi (1976) and Tateishi & Ohashi (1981) the Malesian material belongs to subsp. gigantea. The other subspecies, subsp. tashiroi (Hayata) H. Ohashi & Tateishi, is a Taiwanese endemic.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF9FFEE1A7DFE885B1CFCBC.taxon	description	Negretia urens auct. non Tussac: Blanco (1837) 586, p. p.; (1845) 409, p. p.; (1879) 387, p. p. Distribution — Nepal, India, Bangladesh, Burma; Malesia: Philippines: Luzon, Guimaras Isl., Leyte, Mindanao. Habitat & Ecology — Along creeks. Altitude up to 100 m. Flowering: January, March, July, October; fruiting: November. Note — Also known as M. nigricans var. / subsp. nigricans. Mucuna hainanensis is one of the species formerly confused with M. nigricans. The latter name is now a rejected name, see Dubious and Excluded Species. For more detailed synonymy and a comprehensive discussion, see the papers by Wilmot-Dear (1991 a, b). The subsp. hainanensis is found in China, Vietnam and Thailand.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF9FFE91A7DFC215DA0FC72.taxon	description	Twigs ferruginous tomentose. Leaflets 6.5 – 10.5 by 4 – 6.5 cm, terminal elliptic or obovate, lateral obliquely ovate. Inflorescences pseudopanicles or pseudoracemes, 1.5 – 7 cm long. Calyx campanulate, 12 – 15 mm long, golden sericeous, tube 3 – 9 mm long. Blade of standard 20 – 23 mm long; blade of wing petals 3.3 – 4 cm long; blade of keel petals 3.2 – 4 cm long. Pods 8.5 by 3 cm, lamellae inconspicuous. — Type: Haviland s. n. (holo K), Sarawak, near Kuching Liana. Twigs terete, striate, 2 – 4 mm diam, ferruginous sericeous to pilose, glabrescent. Stipules caducous. Petioles 6.5 – 11 cm long, grooved, ferruginous puberulous, glabrescent; rachis mostly as the petiole, 2 – 3.5 cm long; pulvinus 7 – 10 mm long. Stipellae acicular, 1.9 – 3.6 by 0.1 – 0.5 mm, with few hairs to sericeous at both sides. Leaflets: terminal elliptic or obovate, 8.5 – 10.5 by 4.5 – 6.5 cm, index 1.4 – 1.7, base truncate or rounded, apex acuminate, acumen 2 – 10 mm long, above with scattered appressed hairs to thinly sericeous, below tomentose, midrib raised above, nerves slightly raised above, 4 – 6 per side, 13 – 26 mm apart, anastomosing near the margin; lateral mostly as the terminal, obliquely ovate, 6.5 – 10.5 by 4 – 6.5 cm; pulvinus 4 – 8 mm long. Inflorescences axillary or raminascent, pseudopanicles or pseudoracemes, 1.5 – 7 cm long, peduncle 0.2 – 0.7 cm long, ferruginous sericeous, branches 2.5 – 4 cm long. Bracts to the brachyblasts ovate, c. 7.5 by 5 mm, outside ± pilose, inside glabrous. Brachyblasts 1 – 5 mm long. Bracts to the flowers ovate or elliptic, 11 – 21 by 5 – 6 mm, outside sericeous and with irritating hairs, inside thinly sericeous, caducous. Pedicels 11 – 15 mm long, ferruginous tomentose. Bracteoles ± elliptic or narrowly ovate, 11 – 15 by 3 – 5 mm, outside sericeous, inside sericeous and with irritating hairs. Calyx in bud cup-shaped, in anthesis broadly campanulate, 12 – 15 mm long, tube 3 – 9 mm long; upper lip triangular, 1 – 5 by 10 – 15 mm, lateral teeth triangular, 3.5 – 6 by 4 – 8 mm, median tooth triangular 6 – 9 by 5 – 6 mm; outside golden sericeous and with irritating hairs, inside sericeous. Corolla purplish. Standard: claw 3 – 4 mm long; blade broadly ovate, 20 – 23 by 15 – 20 mm, auricles 2 mm long, both sides glabrous. Wings: claw 5 – 7 mm long, outside with some hairs between claw and auricle, ciliate along both margins in upper part, inside with some hairs at upper margin at base; blade elliptic, 3.3 – 4 by 0.4 – 1.2 cm, rounded, auricle 2.5 – 3.5 mm long, lateral pocket 5 – 6 mm long, outside sericeous at auricle and just above, ciliate at lower margin at base, inside glabrous. Keel petals: claw 6.5 – 8.5 mm long, with some scattered hairs to ciliate along upper margin; blade narrowly elliptic, ± falcate at apex, 3.2 – 4 by 0.5 – 0.7 cm, auricles 2 – 3.5 mm long, hard part 7 – 9 mm long, lateral pocket 5 – 7 mm long, both sides glabrous, short-ciliate along upper margin. Stamens 3.8 – 4.5 cm long, tube 2.8 – 3.6 cm long, free part of filaments below versatile anthers 6 – 8 mm long, below basifixed ones 7 – 9 mm long; versatile anthers 0.9 – 1.1 by 0.7 – 1.2 mm, bearded, basifixed anthers 1.8 – 2 by 0.6 – 0.7 mm, outside ± villous at base more appressed upwards. Disc l d 0.7 – 1.1 mm high. Ovary 5 mm long, stipe 0.9 – 1.0 mm long, sericeous and with irritating hairs; ovules 2; style 3.5 – 3.8 cm long, sericeous, thinning upwards, upper part glabrous. Pods flattened ellipsoid, 8.5 by 3 cm, upper wing c. 7 mm wide, lower wing 6 – 9 mm wide, lamellae inconspicuous, more distinct at lower margin in basal part, interrupted in the middle, up to 2 mm high, the lower part of the lamellae more developed than the upper part, valves with few hairs and some irritating hairs. Seeds orbicular, 1.5 by 1.0 cm; hilum> 1 / 2 the circumference. Distribution — Malesia: Borneo: Sarawak, Sabah. Habitat & Ecology — Lowland or secondary forest. Flowering: July, November. Note — This species is ± similar to M. acuminata but differs in the ferruginous indumentum of the lower surface of the leaflets, the less well developed calyx teeth, the two-seeded pods with at least some lamellae. Mucuna acuminata has ± smooth pods and usually more than 2 seeds. The type of M. havilandii (Haviland s. n.) is a flowering specimen to which an old pod (collected from the forest floor under or near the specimen?) is added. This pod is very different from the pods of other Bornean species of Mucuna in the inconspicuous, interrupted lamellae. One of the flowers of the specimen has a very young pod, which shows after partial removal of the indumentum obscure lamellae. This young fruit does not contradict the idea that the pod added to Haviland s. n. belongs to the present species.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFEFFE91932FC0E5EEFFB54.taxon	description	Mucuna hooglandii Verdc. (1978 a) 459, f. 1; (1979) 444, f. 102,106 E. — Type: Hoogland 4328 (CANB, K, L, LAE), Papua New Guinea, Milne Bay Prov., Cape Vogel Peninsula, some km inland of Tapio. Distribution — Malesia: Papua New Guinea: Milne Bay,? Central Prov. Habitat & Ecology — Secondary forest. Altitude up to 150 m. Flowering: March, July; fruiting: July.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFEFFE91932FB775BD5FC1D.taxon	description	Twigs thinly sericeous. Stipellae absent. Leaflets with 2 – 3 nerves per side. Inflorescences pseudopanicles, 10.5 cm long. Corolla green with purple throat. — Type: McDonald & Ismail 4003 (holo L), Celebes, Kabaena, Gunung Sabampolulu, 1 km S-SE of Tangkeno. Altitude 700 – 900 m, 3 July 1993. Liana. Twigs terete, 3 mm diam, thinly sericeous. Stipules caducous. Petioles 5 – 6.3 cm long, grooved, striate, thinly sericeous; rachis mostly as the petiole, 2.5 – 3.7 cm long, pulvinus 5 – 6 mm long. Stipellae absent. Leaflets: terminal narrowly elliptic or narrowly obovate, 9 – 9.6 by 4 – 4.2 cm, index 2.0 – 2.4, base rounded, apex acuminate, acumen 5 – 8 mm long, above glabrous or with few appressed hairs, below very thinly sericeous, midrib and nerves more densely so, midrib raised above, nerves flat or slightly raised above, 2 – 3 per side, 23 – 36 mm apart, anastomosing near the margin; lateral mostly as the terminal, obliquely, narrowly ovate, 8.7 – 8.8 by 4.2 cm; pulvinus 5 – 6 mm long. Inflorescences axillary, pseudopanicles, 10.5 cm long, peduncle 2.5 cm long, thinly sericeous, becoming denser upwards, branches 8 – 9 cm long. Bracts to the brachyblasts caducous. Brachyblasts 1 – 2 mm long. Bracts to the flowers caducous. Pedicels c. 26 mm long, sericeous. Bracteoles caducous. Calyx 11 mm long, tube 8 – 9 mm long; upper lip ± semicircular, 1 by 14 mm, lateral teeth triangular, 1 by 4 mm, median tooth triangular, 2.5 by 5 mm; outside sericeous and with some irritating hairs, inside sericeous. Corolla light green with dark purple throat. Standard: claw 3 mm long, glabrous; blade ± orbicular, 20 by 19 mm, emarginate, auricles 1 mm long, both sides glabrous. Wings: claw 5 mm long, outside sericeous along both margins, ciliate along both margins, inside sericeous along lower margin; blade ± elliptic, 25 by 7 mm, rounded, auricles 2 mm long, lateral pocket 10 mm long, outside sericeous at auricle and just above up to top of lateral pocket, ciliate along lower margin in lower part, inside sericeous along lower margin in lower part. Keel petals: claw 8 mm long, glabrous, ciliate along upper margin in upper part; blade ± elliptic, bent in apical part, 23 by 5 mm, acute, auricles 1.5 mm long, lateral pocket 8 mm long, hard part 5 mm long, glabrous, short-ciliate along upper margin. Stamens 36 mm long, tube 23 – 29 mm long, free part of filaments below versatile anthers 6 mm long, below basifixed ones 6 mm long; versatile anthers 0.7 by 0.6 mm, bearded, basifixed anthers 2.2 by 0.4 mm, outside with some appressed hairs at base. Disc 0.6 mm high, glabrous. Ovary 7 mm long, sericeous; ovules 5; style 28 mm long, sericeous thinning upwards apical part glabrous. Distribution — Malesia: Celebes: Kabaena. Habitat & Ecology — W slopes of Gunung Sabampolulu. Mixed ecotone of grassland and short forest. Soil: serpentine. Altitude 700 – 900 m. Flowering: July. Note — Only known from the type. The irritating hairs on the calyx seem to be softer than in other species. Vegetatively similar to M. gigantea, but different in stipellae: absent in the new species, present in M. gigantea and the inflorescences: pseudopanicles in the new species, umbel-like pseudoracemes in M. gigantea. Because of the absence of stipellae similar to M. bennettii, which has much larger, red flowers and M. kawakabuti of Sumba, which differs slightly in the size of the leaflets, in the ratio l / w of the leaflets 2.5 – 3.0 in M. kawakabuti, 2.0 – 2.4 in the new species and the number of nerves 6 – 7 in M. kawakabuti, 2 – 3 in the new species.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFEFFE91A7DFBBE5A84FA55.taxon	description	Mucuna kawakabuti Wiriad. in Wiriad. & H. Ohashi (1990) 97. — Type: Iboet 395 (BO, L), Lesser Sunda Islands, Sumba, Maumarru. Distribution — Malesia: Lesser Sunda Islands: Sumba. Habitat & Ecology — Forest. Fruiting: May. Note — Only known from the type, a fruiting specimen consisting of vegetative shoots and loose pods. The pods are very different from those of other Malesian species with pods without wings and lamellae. They look similar to those of M. macrocarpa Wall., which are, however, usually longer with more seeds and less hairy (to almost glabrous). Mucuna macrocarpa is found from India to China, Japan, Laos, Vietnam and Thailand.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFEFFE91A7DFA785A5DF8F8.taxon	distribution	Distribution — Malesia: Moluccas: Key Islands. Note — The only specimens present in L are all collected in the Bogor Botanical Garden. The original material was collected in the Key Islands. Mucuna keyensis is very similar to M. platyphylla from which it differs in indumentum, shape and size of bracteoles, size of median (lower) calyx tooth, length of claw of the keel petals and the length of the ovary.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFEFFE91A7DF89F5A1CF7B9.taxon	description	Mucuna kostermansii Wiriad. in Wiriad. & H. Ohashi (1990) 99, f. 3. — Type: Kostermans & Wirawan 775 (holo BO; iso AAU, K, L), Lesser Sunda Islands, Flores, along road Bea Laing-Rana Mese. Distribution — Malesia: Lesser Sunda Islands: Flores. Habitat & Ecology — Altitude 1000 m. Flowering and fruiting: May. Note — Only known from the type specimen.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1932FF015E9EFDD6.taxon	description	Mucuna lamii Verdc. (1978 a) 463, f. 2; (1979) 446, f. 103. — Type: BW 5523 (Van der Sijde) (holo L; iso Djajapura, L), Papua, Cycloop Mountains. Distribution — Malesia: New Guinea: Papua Barat; Papua New Guinea: Chimbu, W Sepik, E Sepik Prov. Habitat & Ecology — Primary or secondary forest. Soil: sandy. Altitude 100 – 800 m. Flowering: March, May, September, October; fruiting: February, May, September, October. Note — Several labels give the colour of the lower surface of the leaflets as purplish. Some inflorescences may look ‘ pseudo-umbellate’ especially when young. The blade of the wings is ± constricted above the claw.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1932FDF75E06FB00.taxon	description	Mucuna longipedunculata Merr. (1905 b) 18; (1910) 117; (1923) 308; Wilmot-Dear (1991 b) 223, f. 3 A – H, 4. — Type: Elmer 6233 (K, NY, PNH †), Luzon, Prov. Benguet, Sablan.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1932FDF75E06FB00.taxon	distribution	Distribution — Malesia: Philippines: Luzon, Polilo, Catanduanes, Surigao, Biliran, Mindanao. Habitat & Ecology — Forest along stream, logged forest. Altitude 200 – 300 m. Flowering: March to June; fruiting: May, June. Note — The peduncle of the inflorescence is extremely long, slender and wiry, quite elastic and strong. The branches of the pseudopanicles become shorter upwards, in the apical part they are reduced to brachyblasts. The seeds of the two fruiting specimens seen by us are misshapen (see also Wilmot-Dear 1991 b: 223: “ seeds …. very misshapen in dry state (? soft in living state) ”). From Mindanao M. macmillanii was described which only differs in some measurements most clearly in the smaller bracteoles. Here we unite M. macmillanii with M. longipedunculata.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1932FA855A1BFD66.taxon	description	Mucuna macrophylla Miq. (1855) 213; Burck (1893) 190; Backer & Bakh. f. (1964) 630. — Type: Zollinger 1143? (BO?, P?, fragm. U), Java, Tjikoja (see Note).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1932FA855A1BFD66.taxon	description	Mucuna gigantea auct. non (Willd.) DC.: Benth. (1852) 237. Distribution — Malesia: Sumatra (once collected, see Note); Java; Lesser Sunda Islands: Flores. Habitat & Ecology — Forest, open place in forest. Soil: limestone. Altitude up to 1350 m. Flowering: January to April, October, December; fruiting: March to May, August. Note — The only specimen mentioned by Miquel for M. macrophylla is: “ Java, tusschen struiken bij Tjikoja ”. According to the first author this is the specimen Zollinger 1147. The Utrecht herbarium (now at L) holds a specimen labelled as Typus fragment. The only other information is on an older label saying in Miquel’s handwriting: “ Mucuna macrophylla M, Java ”. This fragment consist of a lateral leaflet of 19.5 by 13.5 cm, much larger than usually in M. macrophylla. Prain (1897 b: 407) mentions the name M. junghuhniana in observations sub M. imbricata. It is unclear whether or not he meant to make a new combination. Burck (1893) mentions several localities for his species M. blumei: Java, Koeripan, prope Buitenzorg et Oengarang, Forbes in herb. Lugd. Bat. The first locality refers to Blume 771. The second and third localities to a Junghuhn collection (Junghuhn 193); the L duplicate of that collection consist of a piece of a branch and a leaf, the sheet bears three labels: one attached to fragment ‘ Jul montis Rembang’ and two that refer to ‘ Oengarang’, one with the months Mei, Jun., the other with the months April-Jun. The fourth locality probably refers to Forbes 1417, Lampongs, Kota-Djawa, the type of M. ovalis Baker f. This specimen was collected in Sumatra. For M. blumei the Blume specimen was selected as the lectotype. The twigs of older specimens tend to have a more appressed indumentum. Forbes 1417 (the type of M. ovalis) has both pseudopanicles and very short pseudoracemes with only 1 – 2 brachyblasts. A Teysmann specimen (4492 HB), collected in ‘ Lampongs’, probably also belongs here. The flower colour of Schmutz 4292 is given as blue (‘ blaublühend’).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1A7CFD625AB1FBB8.taxon	description	Mucuna macropoda Baker f. (1923) 11; Verdc. (1979) 448; Hopkins & Hopkins (1993) 297, f. 1 – 4. — Type: Forbes 289 (BM, seen photo at K), PNG, Sogeri. Distribution — Malesia: Papua New Guinea: Central Prov. Habitat & Ecology — Forests. Altitude 800 – 1100 m. Flowering: June, July, October; fruiting: October. Note — Hopkins & Hopkins (1993) observed visits by small bats at night. Also several species of ants were seen crawling over the flowers and small beetles were collected from an inflorescence. The dissected flowers were all ± damaged by insects. Some parts were not complete. Several measurements were taken from the publication by Hopkins & Hopkins (1993).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEA1A7DFBDD5AAFFA56.taxon	description	Mucuna mindorensis Merr. (1908) 231; (1923) 309; Wilmot-Dear (1991 b) 221. — Mucuna acuminata Merr. (1906) 196, nom. illeg. — Type: Merrill 4069 (K, L, NY, PNH, US), Mindoro, Baco R. Distribution — Malesia: Philippines: Mindoro. Habitat & Ecology — On beaches. Fruiting: March, May. Note — Irritating hairs seem to be lacking. The cells of the reticulum of the pod are often quite conspicuous, 3 – 8 by 1 – 5 mm, although borders between cells may be inconspicuous.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFDFFEB1A7DFA775EBFFE24.taxon	description	Mucuna clemensiae Merr. & L. M. Perry (1948) 156. — Type: Clemens 6573 (holo A; iso K), NE New Guinea, Tobou. Mucuna urens DC. var. papuana F. M. Bailey (1910) 20. — Syntypes: Le Hunte s. n. (n. v.), Schenkler s. n. (K) (see Note). Distribution — Malesia: Moluccas; New Guinea: Solomon Islands. Habitat & Ecology — Primary and secondary forest, disturbed forest, Sago swamps, forest edge, along paths and roads. Soil: black volcanic soil, sand, clay, rocky clay, gravel. Altitude up to 1500 m. Flowering and fruiting: throughout the year. Note — Wilmot-Dear (1990) gives Schlenker s. n. as the type of M. urens var. papuana. However, Bailey based his variety on sterile specimens collected by Le Hunte and fruits collected by Schlenker, stating: “ All the above is from some specimens collected by Sir G. R. Le Hunte. ” That description is followed by a description of the pods and seeds collected by Schlenker. Clearly a lectotype should be chosen from the specimens of Le Hunte. The calyx in bud is ± cylindrical. When the corolla expands the calyx widens and becomes campanulate, however, it is not pushed downwards and in the end it is not tucked up. Longest hairs 0.4 – 1.5 mm long.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFCFFEB1932FDA75E02F903.taxon	description	Mucuna novo-guineensis Scheff. (1876) 18; Verdc. (1979) 450, f. 104, 107 A; Wilmot-Dear (1990) 32. — Neotype (here designated by Adema): Van Royen & Sleumer 6300 (L), Netherlands New Guinea, distr. Hollandia, Cycloop Mountains, road Sentani to Bozai village, 110 m, 26 July 1961. Mucuna kraetkei Warb. (1891) 329. — Type: Warburg s. n.? (n. v.), Papua New Guinea, Hatzfeldhaven. Distribution — Malesia: Moluccas: Halmahera; New Guinea. Habitat & Ecology — Primary, secondary or swamp forest, usually along rivers, at river or stream banks. Altitude up to 2000 m. Soil: stoney clay, limestone, granite, volcanic sedi- ments. Flowering: February to November; fruiting: March, July, December. Uses — Stems are used for lashing and bridge construction. The sap is used for dying stringbags. Men in Haus Tambura (E Sepik prov.) drink the sap during ceremonial occasions. Note — Sap is watery, colourless or milky at first slowly turn- ing to red and later to black. Mucuna novo-guineensis is in New Guinea a rather common liana. From other red / orange-flowered Mucuna species (M. bennettii, warburgii) it can easily be distin- guished by its very short calyx teeth. Mucuna novo-guineensis is a rather variable species, especially in the length of inflorescences and peduncles and in the indumentum of several parts. Shrimps are attracted by flowers that drop into the water (pers. comm. Wanda Ave, Wim Vink). Mucuna novo-guineensis was described by Scheffer (1876) on fruiting specimens. The description of calyx and fruits fit nicely with that of specimens usually called M. novo-guineensis. He based his description on three specimens collected by Teijsmann in New Guinea: near Doré, near Andaj and in the Humboldt baai. As far as known to us no Teijsmann material from these localities exist nowadays. In his manuscript for a Flora Malesiana treatment Wiriadinata named Teijsmann 7465 as ‘ type’. The BO and L duplicates of this specimen consist of leafy twigs only and are in a bad shape. The L duplicate seems to lack stipellae and probably belongs to M. bennettii. As Scheffer used three specimens in his description Wiriadinata should have selected a lectotype. However, as there are no original Teijsmann specimens available, a neotype is needed. The specimen Teysmann 7465 proposed as type is of uncertain provenance and not identifiable. We decide to select as neotype a specimen from the same general area (W Irian, ‘ Netherlands New Guinea’) were Teijsmann collected his original specimens.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFCFFEB1932F8BF5BABFE59.taxon	distribution	Distribution — Malesia: Philippines: Mindanao. Notes — E. P. Parreno described this species as part of his revision of Mucuna of the Philippines for a MSc thesis at the University of Kentucky. This manuscript was never published. According to Wilmot-Dear (1991 b) Harry Wiriadinata was about to publish the description of Parreno’s new species in the Journal of Japanese Botany, however, also that never oc- curred. So Wilmot-Dear was the first to published the name of M. pachycarpa, however, at that time a latin description was needed. So the name was not valid published. Here we validate the name by reference to the description by Wilmot-Dear. Mucuna pachycarpa is in several aspects rather similar to M. longipedunculata which differs in: the narrower leaflets (4.5 – 10 cm wide), longer inflorescences (1 – 7 m long), sutures of the pods which are not thickened.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFCFFEB1A7DFE7A5B8BFB8E.taxon	description	Mucuna pruriens (L.) DC. subsp. novo-guineensis Verdc. (1978 a) 462; (1979) 453. — Type: Hartley 10172 (CANB, K, L, LAE), Papua New Guinea, Morobe Prov., Burep River, NE of Lae. Distribution — Malesia: Irian Jaya: Ayawasi; Papua New Guinea: Morobe, W Sepik. Habitat & Ecology — Primary and secondary forest, regrowth, roadsides, bushes, streamside in grassland, savannah, dry gully. Altitude up to 1100 m. Flowering: April to June, September, October, December; fruiting: April, June, September, November. Note — Most specimens are collected in Morobe Prov., PNG. One collection from Irian Jaya (Ridsdale 2252, Ayawasi) is included here. Probably also a collection from W Sepik Prov., PNG (Hoover 439) belongs to the present species. However, the last specimen is collected at a much higher altitude (600 – 1100 m). Verdcourt (1979) cites several specimens seen by him in Lae and remarks that the specimens cited by Schumann & Lauterbach (1901: 365) as M. pruriens may be referable to the present species. However, no duplicates of these specimens (Hollrung 147, Lauterbach 15, 461, 2092, 2279) were seen by Verdcourt, Wiriadinata or Adema.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFFCFFEB1A7DFB0D5B6FF7AE.taxon	description	Mucuna schmutzii Wiriad. in Wiriad. & H. Ohashi (1990) 102. — Type: Schmutz 342 (holo L), Flores, Dentjang. Distribution — Malesia: Christmas Isl.: Indian Ocean, introduced?; Celebes: Gorontalo; Lesser Sunda Islands: Flores: Moluccas: Ternate, Morotai; New Guinea; Solomon Islands; New Caledonia. Habitat & Ecology — Primary and secondary forests, disturbed forests, open woodland, Sago swamp, regrowth border- ing road, along rivers. Altitude up to 800 m. Flowering: January, April to August, December; fruiting: July. Notes — Sometimes two pseudopanicles per axil. Longest hairs 0.6 – 1.5 mm long. Kostermans 1366 (Moluccas, Morotai) probably belongs here, however, on the label the flower colour is given as red. Mitchel 46 (Christmas Isl., Indian Ocean) probably belongs here, however, the specimen differs slightly in the measurements of the flower parts. Four New Guinean collections (Hoogland 3683, NGF 1595, 16324, Van Royen & Sleumer 6175) differ slightly from the majority of the material, mainly in the wider terminal leaflets, the smaller bracts, bracteoles and flowers, however, overlaps in many measurements occur. No matching material with fruits and seeds has been found. For now these specimens are included in M. platyphylla.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF3FFE51932F82A5D13FF26.taxon	description	Mucunaplatyplekta Quisumb. & Merr. (1928) 152; Wilmot-Dear (1991 b) 244, f. 12, map 3. — Type: BS 47232 (Ramos & Edaño) (holo PNH †; iso NY, UC), Luzon, Isabela prov., San Mariano. Distribution — Malesia: Philippines: Luzon. Habitat & Ecology — Dry open forest, along streams. Altitude low. Fruiting: February to April. Note — This species is only known in fruit. The fruits are similar to those of M. biplicata and M. diplax, which differ in the indumentum of twigs and inflorescence axes (appressed in M. biplicata and M. diplax, glabrous to thinly puberulous in M. platyplekta), the indumentum of the lower surface of the leaflets (thinly strigose in M biplicata, very thinly sericeous in M. diplax, sericeous in M. platyphylla) and the width of the horizontal part of the lamellae (1 – 4 mm in M. biplicata, 2 mm in M. diplax, 5 – 15 mm in M. platyplekta).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF2FFE51932FEA25AB2FE39.taxon	description	[Cacara pilosa Rumph. (1747) 392.] See Merr. (1917 a) 279. [Cacara pruritus Rumph. (1747) 393, t. 142.] See Merr. (1917 a) 277. Marcanthus cochinchinensis Lour. (1790) 461. — Mucuna cochinchinensis (Lour.) A. Chev. (1919) 91. — Mucuna pruriens (L.) DC. forma cochinchinensis (Lour.) Backer in Backer & Bakh. f. (1964) 629. — Type: Loureiro s. n. (BM), Cochinchina. Carpopogon niveus Roxb. [(1814) 54, nom. nud.]; (1832) 285. — Mucuna nivea (Roxb.) Wight & Arn. (1834) 255. — Lectotype (Wilmot-Dear 1984): Roxburgh drawing 1601 (K). Carpopogon capitatus Roxb. (1832) 284. — Lectotype (Wilmot-Dear 1984): Roxburgh drawing 285 (K) (‘ Soorootoo’).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF2FFE51932FEA25AB2FE39.taxon	distribution	Distribution — Tropical Africa, Madagascar, Mascarenes, S Asia, Malesia: Sumatra; Java; Philippines: Luzon, Mindanao; Celebes; Lesser Sunda Islands: Bali, Lombok, Soemba, Flores, Timor; Moluccas: Ceram, Halmahera; New Guinea. Also cultivated in the tropics of the old and new world. Habitat & Ecology — Primary and secondary forests, monsoon forest, grasslands, along hedges and fields, along rivers, at the sea coast, along roads. Soil: sand, loam, limestone, volcanic soil. Altitude up to 1300 m. Flowering: January, April to July, September, October, December; fruiting: January, April to December. Note — A very variable species. At several levels, from species to formae, entities have been described, many of the names concern cultivated forms. Wilmot-Dear (1984) who revised Mucuna for several areas in Asia is consistent in the use of varieties for the most distinguishable forms. Here we follow her lead and distinguish several varieties of which two occur in Malesia (var. pruriens and var. utilis). All cultivated forms are combined in var. utilis. Backer (1916 and in Backer & Bakhuizen van den Brink 1964) mentions the form / variety hirsuta for E Java. No specimen of this variety from Java was seen by the present authors. M. pruriens var. hirsuta is found in Continental Asia.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF2FFE71A7DFBCF5D23FE1E.taxon	description	Carpopogon niveus Roxb. [(1814) 54, nom. nud.]; (1832) 285. — Mucuna nivea (Roxb.) Wight & Arn. (1834) 255. — Lectotype (Wilmot-Dear 1984): Roxburgh drawing 1601 (K). Carpopogon capitatus Roxb. (1832) 284. — Lectotype (Wilmot-Dear 1984): Roxburgh drawing 285 (K) (‘ Soorootoo’).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF2FFE71A7DFBCF5D23FE1E.taxon	distribution	Distribution — Cultivated in the tropics and subtropics, also in Malesia. Sometimes escaped and naturalized. Habitat & Ecology — Cultivated fields and gardens; as an escape in secondary and disturbed forests, roadsides. Altitude up to 300 m. Flowering: April, June, September, October, December; fruiting: April to July, October, December. Note — Mainly cultivated for the presence of L-DOPA (see Hegnauer & Hegnauer 2001: 343 – 345). Also cultivated as food for humans and animals. Var. utilis is at present seen as a cultivar-group (Cv. - group Utilis) see Westphal (1974) 121 and Wulijarni-Soetjipto & Maligalig (1997) 199.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF0FFE71932FDB95FAEFB8D.taxon	distribution	Distribution — Malesia: Philippines; Celebes. Habitat & Ecology — Primary or secondary forest. Soil: (coral) limestone. Altitude up to 900 m. Flowering: May to August; fruiting: March, June, August, October. Uses — The seed is reported as a medicine for diarrhoea (Koorders 1898). Note — The species is remarkable for the reticulate lamellae on the pods. In vegetative characters M. reticulata resembles M. longipedunculata, which, however, has rather different pods. Several specimens have a small leaf at the top of the peduncle of the inflorescence. The indumentum of the lamellae tends to be patent, that of the pod valves is more appressed. Afriastini 1992 (Celebes, Dusun Rea, Desa Sondoong) probably belongs here, however, the label gives the flower colour as glaucous green.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF0FFE71932FB0B5B52FC97.taxon	description	Twigs glabrous, (thinly) puberulous or very thinly sericeous. Petioles 4 – 11 cm long. Leaflets ovate, 5.5 – 12.2 by 3 – 7 cm, acumen 5 – 11 mm long, nerves 3 – 6 per side. Inflorescences axillary, pseudoracemes, 8 – 11.5 cm long. Calyx 10 – 11 mm long. Corolla pale green. Ovules 2. — Type: De Haan 1773 (holo BO; iso A, K, L), Halmahera, Distr. Weda, Pajahi road, 26.11.1950. Liana. Twigs terete, striate, diam 1 – 6 mm, glabrous, (thinly) puberulous or very thinly sericeous. Stipules caducous. Petioles 4 – 11 cm long, striate, puberulous or with scattered appressed hairs; rachis mostly as the petiole, 1.5 – 3.2 cm long; pulvinus 3 – 6 mm long. Stipellae acicular, 2.4 – 3.8 by 0.1 mm, glabrous or with some hairs. Leaflets: terminal ovate, 6 – 12.2 by 3 – 7 cm, index 1.6 – 2.1, base rounded or truncate, apex acuminate, acumen 5 – 11 mm long, above with scattered appressed hairs, below with scattered appressed hairs or thinly sericeous, midrib and nerves slightly raised above, nerves 3 – 6 per side, 5 – 46 mm apart, anastomosing near the margin; lateral mostly as the terminal, 5.5 – 12 by 3 – 7 cm; pulvinus 3 – 7 mm long. Inflorescences axillary, pseudoracemes, 8 – 11.5 cm long, peduncle 7 – 10 cm long, at base glabrous to sericeous at apex or thinly puberulous at base becoming denser upwards. Bracts to the brachyblasts ovate, 6 – 7 by 2.5 mm, outside sericeous, inside glabrous. Brachyblasts 1 – 3 mm long. Bracts to the flowers ovate, c. 7.5 by 3.0 mm, outside sericeous, inside glabrous. Pedicels 10 – 16 mm long, ferruginous puberulous. Bracteoles caducous. Calyx in bud cup-shaped, at anthesis broadly campanulate, 10 – 11 mm long, tube c. 6 mm long; upper lip semicircular, 4 by 14 mm, bidentate, lateral teeth triangular, 2 – 4 by 5 mm, median tooth triangular, 4 – 5 by 6 mm; both sides (ferruginous) sericeous. Corolla pale green. Standard: claw 2 – 3 mm long, with some hairs at apex; blade ± orbicular, 2 – 2.1 by 1 – 2 cm, outside with some hairs at base, inside glabrous. Wings: claw 5 mm long, ciliate along lower margin; blade elliptic-falcate, 2.7 – 3 by 0.5 – 0.7 cm, auricle 3 mm long, outside sericeous at auricle and just above, ciliate along lower margin at base, inside sericeous along lower margin. Keel petals: claw 5 – 7 mm long; blade falcate, 2.5 – 2.6 by 0.4 cm, auricle inconspicuous, hard part c. 5 mm long, lateral pocket inconspicuous, outside with some hairs between claw and auricle. Stamens 2.5 cm long, tube 2 cm long, glabrous; anthers of longer filaments 0.1 mm long, bearded, of shorter filaments 0.2 mm long, with some hairs. Disc c. 1.0 mm high. Ovary 4.5 – 5 mm long, sericeous; ovules 2; style 22 mm long, sericeous, apical part glabrous. Pods flattened ellipsoid, 6.0 – 7.5 by 2.5 – 4.5 cm, reddish brown irritating hairs, upper wing 4 – 7 mm wide, lower wing 4 – 8 mm wide, with transverse, 6 – 8 mm high lamellae, valves with few hairs and irritating hairs. Seeds discoid, 13 – 16 by 13 – 15 by 3 – 3.6 mm; hilum 33 mm long, 4 / 5 of the circumference. Distribution — Malesia: Moluccas: Halmahera. Habitat & Ecology — Primary forest. Soil: volcanic tuff. Altitude up to 20 m. Flowering: November; fruiting: September, November. Note — The flowers of the only flowering specimen available to Adema at L are insect-damaged. Some of the characters could not be fully described.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF0FFE71A7DFC335BC3FB7D.taxon	description	Mucuna samarensis Merr. (1922) 390; (1923) 309; Wilmot-Dear (1991 b) 237, f. 9 b, c, 10. — Type: BS 24341 (Ramos) (PNH †; iso K, NY, US). Distribution — Malesia: Philippines: Luzon, Samar, Mindanao. Habitat & Ecology — Primary or disturbed forest, along rivers. Soil: limestone or ultrabasic. Altitude up to 50 m. Flowering: April; fruiting: February, April, May.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF0FFE71A7DFB1D5AA0F7A9.taxon	distribution	Distribution — Malesia: New Guinea: Irian Jaya; Papua New Guinea: Morobe, W Sepik, Central, Northern, Milne Bay Prov. Habitat & Ecology — Primary and secondary forest, riverside. Altitude up to 1760 m. Soil: ultrabasic (once recorded). Flowering: January, June to September, November; fruiting: July to September. Note — Flowering parts of inflorescences often quite long, densely set with short, thick brachyblasts, giving the top part a knobby outlook or less dense and brachyblasts more slen- der. Brachyblasts usually 1.5 – 2 mm thick, more slender ones 0.5 – 1.1 mm thick. Apex of wing petals ± hardened like that of keel petals. Pods show no visible ornamentation on the valves. Seeds look ± mature, however, in drying they seem to have shrunk and become irregular (‘ misshapen’, see also M. longipedunculata). Seed coat ± shiny black, pitted. Verdcourt (1980) discusses the differences between M. lane-poolei and M. schlechteri citing the close similarity of these species. He decided to keep the species separate. Comparing the available material of M. lane-poolei (Darbyshire 346, Eyma 4681) with the description of M. schlechteri gives only individual differences, except for the length of the pedicels (7 mm in both specimens) and the indumentum of the disc-lobes (glabrous or with a few hairs in the two specimens). We think that these differences are too small to keep the species separate and have united the two, making M. lane-poolei a synonym of M. schlechteri.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF7FFE01932FF015EBBFDA1.taxon	distribution	Distribution — Malesia: Philippines: Leyte, Luzon, Mindanao, Jolo. Habitat & Ecology — Primary or secondary forests. Soil: clayey loam. Altitude up to 500 m. Flowering: January, February, May, November. Note — The label of Bernhardt s. n. gives the flower colour as black, according to Merrill in the description of M. luzoniensis the flower colour is black purple.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF7FFE01932FD215DE8F988.taxon	distribution	Distribution — Malesia: Papua New Guinea: Morobe, E Highlands, New Britain, Central, Milne Bay Prov. Habitat & Ecology — Primary and secondary forest, logging area, roadsides. Soil: limestone. Altitude up to 2100 m. Flowering: April to November; fruiting: September to November. Notes — Twigs, petiole and rachis, and inflorescence axes with patent hairs of various length. Longest hairs at twigs, petiole and rachis 2.0 – 4.2 mm long, at inflorescence axes 2.8 – 3.5 mm long. In several aspects rather similar to M. platyphylla. Mucuna stanleyi differs in the conspicuous indumentum with the longest patent hairs 1.8 – 4.2 mm long (in M. platyphylla 1 – 1.5 mm long), the much longer stipellae, larger bracts and brachyblasts, larger calyx, the presence of hairs at the stamen tube and the smaller seeds. Brass 5327 according to the label with ‘ panicles very stiff; petals pale green’ probably belongs here. The L sheet is rather incomplete consisting of a twig with an old inflorescence without flowers or fruits to which a leaf and an old flower with a young fruit are added. It is not certain that all parts belong together. Also NGF 24252 with, according to the label, pale green flowers, could belong to M. stanleyi from which it differs mainly in the seemingly glabrous stamen tube and glabrous longer anthers. It also resembles M. platyphylla from which it differs in the length of the longest hairs. Veldkamp & Stevens 5924 has been included here, this specimen differs from M. stanleyi in its smaller leaflets, on average shorter ‘ longest’ hairs and longer pedicels. According to the label of Hopkins & Hopkins 1016 the ‘ Flowers [are] visited by striped possum at night’; Hopkins & Hopkins 1018 states: ‘ Seeds attacked by moth and fly larvae. ’.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF7FFE01932F9085D4BF7BA.taxon	distribution	Distribution — Malesia: Peninsular Thailand; Peninsular Malaysia. Habitat & Ecology — Primary and secondary forests. Altitude up to 150 m. Flowering: February; fruiting: February to April. Note — The pods of this species resemble those of the species of the M. biplicata group. However, the lamellae of the pods of M. stenoplax are not bifurcate, while M. biplicata has bifurcate lamellae. The long brachyblasts of M. stenoplax easily distinguish it from similar looking species (M. hainanensis, M. interrupta Gagnep., M. revoluta Wilmot-Dear).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF7FFE01A7DFF015C7CFC6B.taxon	description	Mucuna ‘ spec. E’, Verdc. (1979) 410. Distribution — Malesia: Papua New Guinea: Bougainville; Solomon Islands: Guadalcanal. Habitat & Ecology — Rainforest, along track. Altitude c. 900 m. Flowering: January; fruiting: April. Note — Known from just a few specimens. The specimens in L are not complete, only one flower was available. Several characters could not be described in full detail. However, the description has been corrected by comparison with the description of Wilmot-Dear (1990). Merrill & Perry (1942) associated the species M. brachycarpa Rech. with Brass 3514 (green-flowered), the type of the present species. In Merrill’s time the common yellow-flowered species from Bougainville was not known. Verdcourt (1979) in his comments to the unnamed species Mucuna spec. E argued that Rechinger’s species, described on a single pod, was probably identical with the yellow-flowered species of Bougainville. Wilmot-Dear (1990) has taken the same view and described the green-flowered species as a new one: M. subumbellata, with Brass 3415 as the type. Vegetatively, M. subumbellata is very similar to M. gigantea.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF7FFE01A7DFC665B0BFA4D.taxon	description	Mucuna sumbawaensis Wiriad. in Wiriad. & H. Ohashi (1990) 104, f. 5, 6. — Type: Kuswata 148 (A, BO, K, L), W Sumbawa, Setongkat Atas. Distribution — Malesia: Lesser Sunda Islands: Sumbawa. Habitat & Ecology — Secondary forest, riverside, savannah forest. Soil: andesite breccia. Altitude 200 – 650 m. Flowering: May, December; fruiting: May. Note — Close to M. gigantea which differs mainly in the pods: M. sumbawaensis pods wrinkled and with thick wings, M. gigantea pods smooth with thin wings. Mucuna sumbawaensis is also rather similar to M. longipedunculata of the Philippines from which it differs in the shorter inflorescences that are pseudoracemes, not pseudopanicles, the short pedicels, smaller calyces, slightly smaller corollas and the winged pods. The material seen at K and L by Adema has flowerbuds and young and ripe pods.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF7FFE01A7DFA4C5A96F809.taxon	description	Mucuna tomentosa K. Schum. in K. Schum. & Lauterb. (1905) 277; Verdc. (1979) 457. — Neotype (here designated): NGF 27849 (Streimann & Kairo) (holo BO; iso A, BISH, BRI, CANB, K, L, SING), Papua New Guinea, Morobe Prov., Wau Subprov., Bulolo, Taun Creek, BGD lease. Distribution — Malesia: New Guinea: Irian Jaya: Manokwari Prov.; Papua New Guinea: E Highlands, Morobe, Oro, Central Prov. Habitat & Ecology — Primary or secondary forests, aban- doned garden, roadsides. Altitude 800 – 2100 m. Flowering: May to August; fruiting: June. Note — Schumann based his description of M. tomentosa on Nyman 663 (Kaiser Wilhelmsland, Sattelberg, 700 m ü. M.). According to Verdcourt (1979) this material was destroyed. A neotype has been chosen. The indumentum of axial parts consists of hairs of different lengths, longest hairs 1.0 – 1.5 mm long. The keel petals resemble some kind of hockey stick.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF6FFE11A7DFB1B5AB1FA46.taxon	description	The lectotype is sterile and could belong to a number of species. The history of the use of the name is rather confusing. The name is now a rejected name (McNeil et al. 2006: 472, 476). See for a more detailed discussion Wilmot-Dear (1991 c, 1992).	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF6FFE11932FF015DC3FD88.taxon	description	Mucuna toppingii Merr. (1917 b) 85. — Type: Clemens 10085 (K), Borneo, Mt Kinabalu, Kiau. Distribution — Malesia: Borneo, Sabah: Mt Kinabalu, Tam- bunan. Habitat & Ecology — Primary forest, along trail. Altitude 500 – 1700 m. Flowering: January, February, October to December. Note — Vegetatively rather similar to M. biplicata differing in the inflorescence and pods. Kokawa & Hotta 5257 has been included here; it differs mainly in the indumentum of the axial parts: more hirsute with short patent hairs. Also SAN 44348 may belong here; this specimen differs in the more hirsute indumentum of the axial parts and in some small differences in sizes of flower parts and in the indumentum of the petals.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF6FFE11932FD085EC8F8E8.taxon	description	Twigs brown tomentose. Leaflets broadly ovate to orbicular, 8 – 16 by 5 – 13 cm, above thinly tomentose or sericeous, below brown tomentose, nerves 5 – 6 per side. Inflorescences pseudoracemes, up to 42 cm long. Pods 25 – 28 by 3 – 5 cm. — Type: NGF 4896 (Womersley) (holo L), Papua New Guinea, E Highlands Prov., A 1 valley near Nondugl. Liana. Twigs terete, 3 – 6 mm diam, brown tomentose. Stipules caducous. Petioles 5.5 – 15.5 cm long, ± grooved above, brown tomentose; rachis mostly as the petiole 2 – 5 cm long; pulvinus 8 – 10 mm long. Stipellae acicular, 7 by 0.3 mm, ± hirsute. Leaflets: terminal, broadly obovate to orbicular, 8 – 15.5 by 5.6 – 13 cm, index 1.2 – 1.6, base truncate to cuneate, apex acuminate, acumen 2 – 8 mm long, above thinly tomentose or sericeous, below brown tomentose, ± shiny, midrib and nerves slightly raised above, nerves 5 – 6 per side, 17 – 30 mm apart, anastomosing near the margin; lateral mostly as the terminal, obliquely ovate, 8 – 16 by 5 – 12 cm; pulvinus 5 – 10 mm long. Inflorescences axillary, pseudoracemes, up to 42 cm long, peduncle up to 30 cm long, brown tomentose. Bracts to the brachyblasts obovate, 50 by 25 mm, inside thinly sericeous, outside sericeous. Brachyblasts c. 3 mm long. Bracts to the flowers narrowly ovate, 35 by 9 mm, inside thinly sericeous, outside sericeous. Flowers not known. Pods flattened, broadly strap-shaped, 25 – 28 by 3 – 5 cm, upper wing 8 – 15 mm wide, lower wing 5 – 15 mm wide, lamellae oblique, 11 – 12 mm high, valves villous and puberulous with abundant irritating hairs. Seeds undeveloped. Distribution — Malesia: Papua New Guinea: Madang, E Highlands Prov. Note — Verdcourt (1979) described the species (B) without giving it a name. According to him M. ‘ B’ (= M. verdcourtii) is rather similar to M. albertsii or M. stanleyi, but very different in the very large bracts and pods. The specimen ANU 909 (Walker) mentioned by Verdcourt belongs to M. aimun. According to Womersley on the label of NGF 4696 the flowers are reported to be red. This is, according to Verdcourt, a mis- take due to confusion with other species. The longest hairs of M. verdcourtii are 1.3 – 3 mm long.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF6FFE11932F8E85A58FDBF.taxon	description	Mucuna warburgii K. Schum. & Lauterb. (1901) 365; Verdc. (1979) 457; Wilmot-Dear (1992) 243. — Syntypes: Lauterbach 856, 953, 1162, 3205 (n. v.), Papua New Guinea, ‘ Kaiser Wilhelmsland’.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF6FFE11932F8E85A58FDBF.taxon	distribution	Distribution — Malesia: New Guinea. Habitat & Ecology — Primary forest, swamp forest, along rivers. Altitude up to 50 m. Note — The original description of M. warburgii mentions stipellae: ‘ stipellis minutis filiformibus’; ‘ Die Stipellen sind kaum 5 mm lang. ’. This species forms with M. bennettii and M. novo-guineensis the group of Mucuna with large, red to orange flowers. The first species lacks stipellae and has large calyx teeth, the second species has stipellae and short calyx teeth, M. warburgii has stipellae and large calyx teeth, it also has more ovules than the two other species. In the original description of M. warburgii four Lauterbach specimens are mentioned. Up to now I have not seen any of these, also Verdcourt (1979) and Wilmot-Dear (1992) mention none of these specimens. For the moment I refrain from selecting a lecto- or a neotype.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFF6FFE11A7DFA455B3EF873.taxon	description	In Kew I have seen a photograph annotated as: Mucuna cf wertheimii Burck, cult. Hort. Bog. XVIII. D. 10, fl. 8. XI. 1910. Fl. auran- tiacum rubri (orange red). According to Verdcourt not Mucuna bennettii, warburgii, elegans, etc. The flowers of Mucuna? wertheimii are not known. The flowers of Dioclea hexandra are white, pinkish or purplish.	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
03DEE569FFC9FFDE1B07FE925AA6FD58.taxon	description	subsp. gigantea 16 a var. pruriens 35 a subsp. plurisema Verdc. 16 var. sericophylla (Perkins) Wilmot-Dear 40 subsp. tashiroi (Hayata) Ohashi & Tateishi var. utilis (Wall. ex Wight) Burck 35, 35 b	en	Wiriadinata, H., Ohashi, H., Adema, F. (2016): Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61 (2): 90-124, DOI: 10.3767/000651916X692799, URL: https://doi.org/10.3767/000651916x692799
