taxonID	type	description	language	source
03DE5A0EFF999D42FF34FBD4FB2B2EC2.taxon	materials_examined	Type species: Quilessa lutea Fennah, 1942, by original designation.	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF999D42FF34FBD4FB2B2EC2.taxon	description	Amended description (modified after Fennah 1942). Small (mostly less than 3 mm), somewhat flattened species. Head with eyes about two-thirds width of pronotum. Eyes slightly longer than broad, emarginate above antennae. Vertex quadrate, usually slightly longer than broad, expanding posteriorly, posterior margin shallowly, and usually angularly, excavated; median and lateral carinae well developed; apical transverse carina absent. In lateral view, head smoothly rounded from posterior margin to frons. Frons and clypeus collectively roughly rhomboid (apex truncate); frons longer than wide at widest point (just above frontoclypeal suture, 1.2 to 1); median and lateral carinae distinct. Clypeus at base 3 / 4 th as wide as widest part of frons, tapering acutely to apex, median and lateral carinae present; clypeus and apical half of frons somewhat convex, basal part of frons sloping smoothly posteriorly into vertex. Frontoclypeal suture nearly straight, median ocellus absent, lateral ocelli anterior to antennae, below anteroventral margin of eye. Eyes emarginate ventrally above antennae. Genae somewhat tumid below antennae, sub-antennal process absent, antennae with scape very short, pedicel slightly longer than broad. Pronotum at midline approximately as long as vertex, anterior margin shallowly excavated behind eyes; posterior border broadly and shallowly emarginate, curving anteriorly near tegulae; median carina distinct, lateral carinae obsolete; lateral margin (from dorsal view) with a strong carina behind eye and tegula; paradiscal region broad with ventral margin truncate. Mesonotum feebly convex from lateral view with weak concavity at scutellum, in dorsal view bearing three distinct carinae, tip of scutellum acute. Hind tibiae without lateral spines, apical spinulation of hindleg: 7 - 6 - 5. Pygofer usually with a lateral process which can be positioned at the dorsal-, central-, or ventral-lateral side. Anal segment of male bifid, often asymmetrically; phallotheca with a ventral keel, often containing a complex sclerotised rod, and often with a process overhanging dorsally.	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF999D42FF34FBD4FB2B2EC2.taxon	diagnosis	Diagnosis. Median carina on vertex, frons and clypeus complete (incomplete in Apocathema Emeljanov, 2017, Eparmene Fowler, 1904, Dineparmene Fennah, 1945, Lomagenes Fennah, 1945 & Mauriciana Campodonico, 2018). Vertex ± as broad as long (longer than broad in Eparmenoides Fennah, 1945). Pronotum with only a median carina (pronotum with median and lateral carinae in Atopocixius Muir, 1926, Oreopenes Ramos, 1957 & Prosotropis Uhler, 1895). Forewings subhyaline or hyaline with apical venation regular and apex widely and evenly rounded (thickened with irregular venation and oblique forewing apex in Microissus Fennah, 1947). Forewing veins M 3 + 4 forked before apex or simple to apex (always simple to apex in Eparmenoides Fennah, 1945 & Oreopenes Ramos, 1957).	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF999D42FF34FBD4FB2B2EC2.taxon	etymology	Etymology. Dmitriev (2022) states that the generic name is derived from the Spanish word ‘ quilo ’ (kilo, thousand) plus the diminutive suffix ‘ – essa ’; however, the name also appears to allude to the type locality of the type species, the Quilesse forest reserve in St. Lucia. The genus name is feminine in gender.	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	description	(Figs 2 – 9)	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	materials_examined	Material examined. Holotype ♂: “ leg. R. H. Cobben / Simson Bay / Devils Swamp / 10 / 1 1957 / St. Maarten // RMNH Leiden / ex. Collectie / WAU 2010 // Holotypus / Quilessa martinensis sp. nov. ♂. ” In coll. RMNH. Paratypes: 2 ♂, 7 ♀ “ leg. R. H. Cobben / Simson Bay / Devils Swamp / 10 / 1 1957 / St. Maarten // RMNH Leiden / ex. Collectie / WAU 2010 // Paratypus / Quilessa martinensis sp. nov. ” In coll. RMNH. Additional material: same data as type-series, two specimens of unknown sex. Type locality. Devils Hole Swamp, Simpson Bay, St. Martin (Fig. 1).	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	description	Description. Measurements. Length: ♂ 2.28 – 2.36 (average 2.32) mm (n = 3); ♀ 2.40 – 2.56 (average 2.51) mm (n = 6). Colouration. Ground colour of head, thorax and abdomen dull-orange (Fig. 2). Genae ventrad of antennae brown. Rostrum stramineous, apex black. Colouration intensity variable. Legs stramineous. Forewings hyaline with indistinct yellowish tinge, veins stramineous (Figs 2 B – C). Hindwings hyaline, clear. Structure. Head (including eyes) 0.66 times width of pronotum. Vertex quadrate, about as long as broad, narrowest at anterior margin, posteriorly expanding, posterior margin evenly rounded, concave; median carinae of vertex complete, but hardly elevated; lateral carinae distinct; in lateral view sloping anteriorly, confluence with frons smooth, without apical transverse carina. Frons broadest in lower half, at broadest point (just above frontoclypeal suture) 1.15 x as broad as long. Frons and clypeus with complete median carina. Rostrum reaching hind-coxae. Pronotum with median carina slightly elevated, anterior margin of pronotum keeled, carina between eye and tegula distinct (forming lateral margin of pronotum from dorsal view); paradiscal region relatively broad, distinctly exceeding ventral margin of antenna, apex smoothly rounded. Tegulae without carinae. Mesonotum much longer at midlength than vertex and pronotum combined (approximately 1.6 x), bearing three distinct, slightly elevated, straight parallel carinae; mesonotum in lateral view convex with concavity at transition to scutellum. Forewings elongate-ovate (Fig. 3; costal margin arched, trailing margin straight), deflexed medially at nodal line, clavus reaching wing margin at nodal line, crossveins absent prior to nodal line; vein MP 1 + 2 usually simple to apex (rarely forked), branching pattern RA 1 - branched, RP 3 - branched, MP 2 - or 3 - branched, CuA 1 - branched; C 5 cell open at wing margin. Hind legs without lateral teeth on tibiae, apical spinulation: 7 (3 + 4) - 6 - 5. Male terminalia. Pygofer in lateral view slender, irregular in outline, narrowest in middle with the anterior margin convex and posterior margin concave, bearing an oblong, distally rounded lateroventral process, medially inflected, directed posterodorsally, pygofer appearing incised between process and ventral margin (medioventral process absent) (Fig. 4). Gonostyli in lateral view relatively slender, cupped, and upcurved, with lateral margin (near midpoint) bearing an elongated slender upcurved, hooked appendage; apices of gonostyli in lateral view flat and broadened into a relatively wide plate; in ventral view median margin bearing a broad, thick hook-shaped structure (also visible from lateral view) (Fig. 5). Phallotheca tubular, simple, weakly downcurved in lateral view, dorsal margin of base hooked, apex posteriorly directed; aedeagal shaft in lateral view of nearly equal width for about 3 / 5 length, apical 2 / 5 expanded into flattened spatulate structure with broadly rounded apex; left lateral lobe more elongated than right lateral lobe; in ventral view phallotheca proximally with wide lateral slightly asymmetrical flanges (collectively appearing nearly cordate, base distad) wide at base, medially fused, with the left lobe slender and long, right lobe shorter; aedeagal apex bulbous; in dorsal view phallotheca widest at base, compressed medially, apex with left lobe slender and long, right / medial lobe bulbous, extending somewhat to the right and covering the apical dorsal part; gonopore subapical on ventral margin (Figs 6 – 7). Connective tubular. Anal tube in lateral view long and slender, downcurved; in dorsal view deeply bifurcated forming elongated (forming horseshoe-shape), weakly asymmetrical, lateral appendages, both appendages curved slightly anteroventrally; apices of appendages distally broadening and with small apical denticle; epiproct short, apical margin straight; paraproct nearly three times as long as epiproct, with apex approximately straight (Fig. 8). External female terminalia. Sternite VIII (gonocoxae) in ventral view funnel-shaped, six-sided, base 4.86 x as broad as apex (Fig. 9 B). Sternite VIII in lateral view broad, covering most of distal terminalia (Fig. 9 A). Tergite IX in dorsal view short and arched. Anal tube in dorsal view as long as wide; epiproct short, apically rounded, paraproct relatively large, disc-shaped (Fig. 9 C).	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	diagnosis	Diagnosis. The new species can be separated from all other species in Quilessa by the absence of markings on the forewings, the pygofer with an oblong rounded ventrolateral process and the long, bifurcated anal tube. The most similar species may be Quilessa lutea from St. Lucia (Fennah 1942, figs 42 – 50), from which Q. martinensis sp. nov. differs most obviously in male terminalia where the pygofer bears an oblong, distally rounded lateroventral process (versus a narrow elongate process in Q. lutea, viz. Fennah 1942, fig. 45), Gonostyli (lateral view) bearing an elongated slender upcurved, hooked appendage (absent in Q. lutea), and the apex of the aedeagus spatulate (in Q. lutea bearing ‘ two large prongs’, viz. Fennah 1942, figs 43 – 44).	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	distribution	Distribution. Saint Martin (Fig. 1). Only known from the type location (Devils Hole Swamp) which is now a residential area.	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	biology_ecology	Biology. No specified information on the ecology of Quilessa martinensis sp. nov. is available. At the time the type series was collected in Devils Hole Swamp, the habitat was a tidal saline pool in coastal limestone, mud and Avicennia L. (Acanthaceae, presumably Avicennia germinans (L.) L., black mangrove; Hummelinck 1981, fig XIb).	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	etymology	Etymology. The specific epithet ‘ martinensis ’ is derived from Saint Martin with the termination ‘ - ensis’ (meaning ‘ of’ or ‘ from’ i. e., ‘ from Saint Martin’), and is intended as feminine in gender.	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
03DE5A0EFF9A9D44FF34FE13FBC828BF.taxon	discussion	Remarks. Two specimens of this species are damaged and are excluded from the type series, one with a missing abdomen and one with wings glued to the specimen card so the abdomen cannot be seen.	en	Haas, Matthé Cornelis De, Bartlett, Charles R. (2024): A new species of Quilessa Fennah from Saint Martin (Fulgoromorpha: Fulgoroidea: Kinnaridae: Prosotropini). Zootaxa 5537 (4): 551-560, DOI: 10.11646/zootaxa.5537.4.7, URL: http://dx.doi.org/10.11646/zootaxa.5537.4.7
