identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E787B8FFA7B709FF7134AFFC459923.text	03E787B8FFA7B709FF7134AFFC459923.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afrixalus dorsalis (Peters 1875)	<div><p>Afrixalus aff. dorsalis (Peters, 1875) complex</p><p>Figures 6A–C</p><p>Material. ANGOLA – Cabinda Province • Tando Zinze; –5.3241, 12.5073; 29 m a.s.l.; P1.138, FKH 0586; GenBank: PQ455620–21. • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; P2.110; GenBank: PQ455622. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0950−51, P2.070−71; GenBank: PQ455623–26. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.127; GenBank: PQ455627.</p><p>Identification. A small reed frog (SVL = 28–30 mm), with diurnal colouration showing a dark brown dorsum and light cream colouration between the eyes that extends as two broad dorsolateral bands towards the anterior insertion of the hindlimbs (Figure 6A), and at night a mostly green-yellow dorsum with slightly darker greenish bands. Some individuals may present a light cream vertebral stripe (Figure 6B), and rarely complete absence of dark brown coloration on the dorsum (Figure 6C). The specimens reported here are genetically identical. However, material from Cabinda Province differs by a minimum of 5% (16S p -distance) from any published material, including material from Mount Kupe, Cameroon (6.5% 16S p -distance; GenBank: KX671711), ∼ 100 km north of the type locality (Limbe, Cameroon). Therefore, these results suggest that A. dorsalis represents a species complex that needs further investigation. Due to the large genetic distance between our samples and with other published material and the sub structuring exhibited by this group, we regard the material from Cabinda Province as Afrixalus aff. dorsalis until more information becomes available.</p><p>Biology and distribution. The Afrixalus dorsalis complex is widely distributed in West Africa and the lowland forests of Central Africa, from Guinea Bissau to northern Angola (Channing and Rödel 2019). In Cabinda Province, the species was found on vegetation near water on the edge of gallery forest, often in disturbed habitat in the Littoral Peneplain and Lower Mayombe, but was not recorded from the Coastal Drier Belt and Upper Mayombe. This suggests some ecological segregation between A. aff. dorsalis and the other two Afrixalus species recorded in Cabinda Province ( A. osorioi and A. paradorsalis).</p></div>	https://treatment.plazi.org/id/03E787B8FFA7B709FF7134AFFC459923	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA9B709FF71336FFCD79FDD.text	03E787B8FFA9B709FF71336FFCD79FDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afrixalus osorioi Ferreira 1906	<div><p>Afrixalus osorioi Ferreira, 1906</p><p>Figure 6D</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1253; GenBank: PQ455628 .</p><p>Identification. A medium-sized reed frog (SVL = 31–35 mm), with shorter body than the other two congeners occurring in Cabinda Province (see Afrixalus aff. dorsalis and A. paradorsalis species accounts). Dorsum with light golden-brown color with a dark brown central blotch (Channing and Rödel 2019). Originally described from Angola, the single specimen of A. osorioi recorded from Cabinda Province is almost identical (0.2% 16S p -distance) to other material from neighbouring Mayongongo, Republic of the Congo (GenBank: KY079930). Additionally, it differs ≥3% from material from northern DRC (GenBank: KY079929), Kenya (GenBank: MK509538) and Uganda (GenBank: MH378405), which suggests cryptic diversification. Furthermore, Baptista (2024) reported structuring between populations north and south of the Congo River that needs further investigation.</p><p>Biology and distribution. Afrixalus osorioi is widely distributed from northwestern Angola northwards to Ogooué River in Gabon and eastwards through the northern rim of the Congo River, to southern Uganda (Channing and Rödel 2019). In Cabinda Province we only collected one specimen, a male found in the rainy season, calling from vegetation near a slow-flowing forest stream in Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FFA9B709FF71336FFCD79FDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA9B70AFF713438FC769ABB.text	03E787B8FFA9B70AFF713438FC769ABB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Afrixalus paradorsalis Perret 1960	<div><p>Afrixalus paradorsalis Perret, 1960 complex</p><p>Figures 6E–F</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Mbongo Zimune; – 4.7442, 12.6914; 306 m a.s.l.; FKH 1244; GenBank: PQ455631. • Mayombe NP, Sanga Wanda; –4.6270, 12.4527; 215 m a.s.l.; P3.520; GenBank: PQ455629. • Mayombe NP, Entrance Sanga Wanda; –4.6304, 12.4714; 219 m a.s.l.; P3.530; GenBank: PQ455630. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0945−46; GenBank: PQ455632–33.</p><p>Identification. Afrixalus paradorsalis represents a species complex of medium-sized (SVL = 28–30 mm) reed frogs (Charles et al. 2018). However, we found low genetic distances within our material (0.6–1.6% 16S p -distance; GenBank: MK509656, MK509588) compared to material from Tsinguidi, Republic of the Congo and Monts de Cristal National Park, Gabon, respectively, and ascribed by Portik et al. (2019) to Afrixalus paradorsalis paradorsalis . Therefore, we ascribe our material to be conspecific with this taxon until more information becomes available.</p><p>Biology and distribution. The Afrixalus paradorsalis complex is widely distributed in the lowland forests of the Gulf of Guinea (Charles et al. 2018). This material constitutes the first records of the species for Cabinda Province and Angola (Baptista 2024) and the southernmost records for this species complex. This species has a unique call, unlike most other Afrixalus, consisting of short sequences of clicks rather than a long series. Individuals from Cabinda Province were found calling from low vegetation near small temporary pools, but we often found them perched far from permanent water.</p></div>	https://treatment.plazi.org/id/03E787B8FFA9B70AFF713438FC769ABB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF99B739FF7133BBFC0B9ED9.text	03E787B8FF99B739FF7133BBFC0B9ED9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agama congica (Peters 1877)	<div><p>Agama congica (Peters, 1877)</p><p>Figure 8D</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Inhuca; –4.8590, 12.4440; 81 m a.s.l.; P2.103; GenBank: PQ455963. • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3804&amp;materialsCitation.latitude=-5.1021" title="Search Plazi for locations around (long 12.3804/lat -5.1021)">between Dinge and Chindende</a>; -5.1021 12.3804, 113 m a.s.l.; sight record. • Nganzi; –5.5292, 12.3264, 89 m a.s.l.; sight record. • Lagoa Massabi; –5.0603, 12.0488, 10 m a.s.l.; sight record .</p><p>Identification. Agama congica was described as a variety of A. colonorum ( Agama colonorum var. congica) by Peters (1877) based on different coloration of juveniles as follows (translation from German): “the pattern of the young specimens (which disappears in the adults) is very different and consists of two rows of black rings on each side and faint lines and spots along the middle of the back on an olive-coloured background”. However, these two morphs were synonymised by Loveridge (1936) and this action was followed by several subsequent authors. Nevertheless, Wagner et al. (2009) considered A. congica to be a valid taxon. A recent phylogenetic analysis (Butler 2024) that includes material from Cabinda Province and the nearby Republic of Congo recovered an independent clade ascribed to this species. However, this genetic material remains unpublished for comparison. Our samples differ by 1.2% (16S p -distance) from samples of A. finchi from Kenya (GenBank: GU128452), 4.3% (16S p -distance) from A. picticauda from Mali (GenBank: GU128441) and 4.1% (16S p -distance) from the neotype of A. agama agama from Margui-Wandala Province, northern Cameroon (GenBank: GU133323).</p><p>Biology and distribution. An arboreal agamid.The species is only known from Cabinda Province, Zaire Province in northern Angola (Marques et al. 2018) and the neighbouring areas of the Republic of the Congo.In Cabinda Province,the species has been observed on the Littoral Peneplain and Coastal Drier Belt but extending at least to Lower Mayombe, being frequently recorded in urban environments, near villages, plantations and mangroves.</p></div>	https://treatment.plazi.org/id/03E787B8FF99B739FF7133BBFC0B9ED9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFBAB71CFF713417FD619F48.text	03E787B8FFBAB71CFF713417FD619F48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthroleptis adelphus (Perret 1966)	<div><p>Arthroleptis adelphus (Perret, 1966)</p><p>Figure 4A</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1255; GenBank: PQ455662 .</p><p>Identification. A medium-sized Arthroleptis (SVL = 30−33.6 mm, Zimkus and Blackburn 2008). The individual could not be clearly assigned to any species by morphology alone. Therefore, the species was identified genetically, being identical or almost identical (0.6% 16S p -distance) to material from Minkébé, Gabon (GenBank: KX289618) and Lekoumou, Republic of the Congo (GenBank: KY080052). Deichmann et al. (2017) found two different lineages that include material from Cameroon (near the type locality), suggesting that A. adelphus might represent a species complex which needs further investigation.</p><p>(Continued)</p><p>Biology and distribution. The species is widely distributed in the Gulf of Guinea, from Nigeria to the Republic of the Congo (Nneji et al. 2019; Sánchez-Vialas et al. 2020). This represents the species’ southernmost record, and the first for Angola (Baptista 2024). As with the other members of the genus, A. adelphus is usually found moving among leaf litter (Zimkus and Blackburn 2008). The only specimen recovered was found foraging among leaf litter at night in the dry season, and in sympatry with the much more common and closely-related species, A. aff. poecilonotus .</p></div>	https://treatment.plazi.org/id/03E787B8FFBAB71CFF713417FD619F48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFBCB71EFF713545FDB59AD8.text	03E787B8FFBCB71EFF713545FDB59AD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthroleptis carquejai Ferreira 1906	<div><p>Arthroleptis carquejai Ferreira, 1906</p><p>Figure 4B</p><p>Material. ANGOLA – Cabinda Province • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; FKH 0978; GenBank: PQ455663. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1232; GenBank: PQ455665. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.083; GenBank: PQ455664. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.6914&amp;materialsCitation.latitude=-4.7442" title="Search Plazi for locations around (long 12.6914/lat -4.7442)">Mbongo Zimune</a>; –4.7442, 12.6914; 324 m a.s.l.; P3.147; GenBank: PQ455666. • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.158; GenBank: PQ455667.</p><p>Identification. Arthroleptis carquejai is a bulky medium-sized Arthroleptis (SVL = 20– 29.7 mm). The specimens from Cabinda Province differ by ∼0.6% (16S p -distance) with material from near topotypic material from Golungo Alto, Kwanza Norte, Angola (&gt; 10 km from the type locality at Cambondo). Our material is identical to material from Loufika, Republic of the Congo and differ by 0.2% (16S p -distance) from material from Tchikoulou, Republic of the Congo (Jongsma unpublished data). In addition, the specimens differ by ∼2.2% (16S p -distance) from type material of A. palava (GenBank: HM238187) and by 3.2% from A. kroskosua (GenBank: EU350211). Arthroleptis carquejai differs morphologically from A. palava in having well-defined black spots on the lateral surface of the head (absent in A. palava) (Blackburn et al. 2010), but similar to specimens of A. variabilis from Gabon (Larson and Zimkus 2018).</p><p>Biology and distribution. Arthroleptis carquejai is known to occur in northern Angola, recorded in disturbed forest habitats and along logging roads (Ernst et al. 2020). However, the species was reported from Gabon by Dewynter and Frétey (2019), although the authors have not provided commentary on their material. Here we report the first records of the species in Cabinda Province (Baptista 2024) and Republic of the Congo (Jongsma unpublished data). In Cabinda Province, the species was recorded from the Littoral Peneplain to higher elevations in Mayombe NP. Specimens were found moving in the leaf litter and on dirt roads at night.</p></div>	https://treatment.plazi.org/id/03E787B8FFBCB71EFF713545FDB59AD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFBEB71EFF713335FBDE9D15.text	03E787B8FFBEB71EFF713335FBDE9D15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthroleptis poecilonotus Peters 1863	<div><p>Arthroleptis aff. poecilonotus Peters, 1863 complex</p><p>Figure 4C</p><p>Material. ANGOLA – Cabinda Province • Nganzi; –5.5304, 12.3236; 78 m. a.s.l.; EI 1129; GenBank: PQ455668 • Cacata; –5.3242, 12.4980; 25 m. a.s.l.; P1.294, FKH-0697; GenBank: PQ455669–70 • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3667&amp;materialsCitation.latitude=-5.1208" title="Search Plazi for locations around (long 12.3667/lat -5.1208)">between Dinge and Chindende</a>; –5.1208, 12.3667; 15 m a.s.l.; P9.050, FKH 0155–56; GenBank: PQ455671–73. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0959, FKH 0969; GenBank: PQ455674–75. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.128, P3.217, P4.099−100, P3.539−40; GenBank: PQ455676–81. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.119−20, FKH 1262; GenBank: PQ455682–84. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.087−89; GenBank: PQ455685–87. • Mayombe NP, Miconge; – 4.4860, 12.8780; 377 m a.s.l.; FKH 1040−44, FKH 1051, P2.221, P2.230; GenBank: PQ455693–700. • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.156; GenBank: PQ455701. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0921 −24; GenBank: PQ455688–91. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; –4.6403, 12.935; 269 m a.s.l.; P3.160; GenBank: PQ455692.</p><p>Identification. Arthroleptis poecilonotus represents a species complex. Material from Cabinda Province is identical to material reported from Mount Kupe, Cameroon (GenBank: KX671725), Mayongongo, Republic of the Congo (GenBank: KY080058) and other material from Angola (Baptista 2024). However, this material differs by 3.2% (16S p -distance) from topotypic material of the species in Ghana (Rockney et al. 2015; GenBank: KU166807). Therefore, it is better to regard material from Cabinda Province and northern Angola as Arthroleptis aff. poecilonotus .</p><p>Biology and distribution. Members of the A. poecilonotus complex are widely distributed from Guinea Bissau eastwards to Uganda and southwards to Angola (Sánchez-Vialas et al. 2020; Baptista 2024). In Angola, the species is widely distributed in the northwestern forest regions (Ernst et al. 2020; Baptista 2024). In Cabinda Province, the taxon was recorded in all the four main ecological zones, from the Coastal Drier Belt to Upper Mayombe. It proved to be one of the most common amphibians in Mayombe NP, often found moving among the leaf litter or crossing dirt roads mostly at night, but sometimes during the day.</p></div>	https://treatment.plazi.org/id/03E787B8FFBEB71EFF713335FBDE9D15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFBEB71FFF713700FC8D996D.text	03E787B8FFBEB71FFF713700FC8D996D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthroleptis sylvaticus Laurent 1954	<div><p>Arthroleptis sylvaticus Laurent, 1954 complex</p><p>Figure 4D</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4407&amp;materialsCitation.latitude=-4.6661" title="Search Plazi for locations around (long 12.4407/lat -4.6661)">Sanga Mongo</a>; –4.6661, 12.4407; 146 m a.s.l.; FKH 0970; GenBank: PQ455703. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.085; GenBank: PQ455702. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1035−37, FKH 1045, FKH 1048; GenBank: PQ455704–08.</p><p>Identification. Arthroleptis sylvaticus also represents a species complex with three distinct mitochondrial lineages (Larson and Zimkus 2018; Deichmann et al. 2017). Material from Cabinda Province is identical to material from the Republic of the Congo (GenBank: KY080080), which represent the closest genetic samples available from the type locality of Buta, DRC (Deichmann et al. 2017), still 1,300 km from the type locality. Also, it is genetically identical to material from Uíge Province, Angola (Baptista 2024). Therefore, material from Cabinda Province and northern Angola may represent true Arthroleptis sylvaticus . However, due to the large distance from the type locality we suggest caution and recommend further taxonomic investigation.</p><p>Biology and distribution. Members of the A. sylvaticus complex are distributed in the Congo Basin. The specimens from Cabinda Province were found in dense rainforest among leaf litter near streams in Lower and Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FFBEB71FFF713700FC8D996D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA4B704FF713335FEF29F6E.text	03E787B8FFA4B704FF713335FEF29F6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Astylosternus robustus (Boulenger 1903)	<div><p>Astylosternus robustus (Boulenger, 1903)</p><p>Figure 4P</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1023; GenBank: PQ455957 .</p><p>Identification. A large species (SVL = 90–130 mm) in a monotypic genus, with reproductive males developing hair-like appendages on the lower parts of the hind limbs (Channing and Rödel 2019). Only one specimen was recorded during this study, and it differs by 1.2% (16S p -distance) from other material from Serra do Pingano, Uíge Province, Angola (GenBank: HG940465) and by 2.6% from material from Cameroon (GenBank: KF991265).</p><p>Biology and distribution. It has a wide distribution in lowland forests and submontane streams from Nigeria southwards to northwestern Angola (Channing and Rödel 2019; Ernst et al. 2020). The specimen reported here was an adult male found perched on a rock in the middle of a fast-flowing river close to a waterfall in Upper Mayombe.</p><p>Bufonidae</p></div>	https://treatment.plazi.org/id/03E787B8FFA4B704FF713335FEF29F6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF82B722FF7132EFFE829EC8.text	03E787B8FF82B722FF7132EFFE829EC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atheris squamigera (Hallowell 1854)	<div><p>Atheris squamigera (Hallowell, 1854)</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3273&amp;materialsCitation.latitude=-5.5279" title="Search Plazi for locations around (long 12.3273/lat -5.5279)">Nganzi</a>; –5.5279, 12.3273; 109 m a.s.l.; photographic record .</p><p>Identification. Atheris squamigera is the only known representative of the genus to occur in Cabinda Province, although no molecular identification has been provided for this species in the territory. Rochebrune (1885) described two species of Atheris ( Atheris lucani and Atheris proximus) from Landana, Cabinda Province, which were subsequently considered as junior synonyms of Atheris squamigera by Loveridge (1957). The specimen reported here was photographed in the forest canopy and we provisionally regard it as Atheris squamigera . The species has very variable colouration.</p><p>Biology and distribution. An arboreal viper widely distributed in the Congo Basin, found in all types of forest, including woodland and savannahs (Spawls et al. 2018). In Angola, it is known to occur in the north (Branch 2018) and has recently been recorded in gallery forest along the northern escarpment (PVP pers. obs.) in Cuanza Norte Province.</p></div>	https://treatment.plazi.org/id/03E787B8FF82B722FF7132EFFE829EC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9EB73FFF7134E8FC4C9A70.text	03E787B8FF9EB73FFF7134E8FC4C9A70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atractaspis congica Peters 1877	<div><p>Atractaspis congica Peters, 1877</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3092&amp;materialsCitation.latitude=-5.0922" title="Search Plazi for locations around (long 12.3092/lat -5.0922)">Dinge</a> and Chindende; –5.0922, 12.3092; 31 m a.s.l.; EI 811 (roadkill); GenBank: PQ455964 .</p><p>Identification. A stiletto snake (max. SVL = 481 mm) with 19–21 midbody scale rows and divided cloacal scales (Peters 1877). The species was originally described from Chinchoxo, Cabinda Province, but there has never been topotypical material for genetical comparisons. Therefore, the obtained sample represents the genetic material closest to the type locality. This material differs by 4.1% (16S p -distance) from material collected from Hillwood Farm, Zambia (GenBank: MK464459), and by&gt;8% (16S p -distance) from material collected from Soyo, Angola (GenBank: MK621461) and DRC (GenBank: MK621462). This suggests some intraspecific diversification south of the Congo River, which is currently under investigation (Lobón-Rovira et al. in prep).</p><p>Biology and distribution. Widely distributed in Central Africa, from Cameroon to Zambia (Chippaux and Jackson 2019). Very little is known about its biology.</p></div>	https://treatment.plazi.org/id/03E787B8FF9EB73FFF7134E8FC4C9A70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF80B721FF71348DFD949A35.text	03E787B8FF80B721FF71348DFD949A35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Boaedon virgatus (Boie 1827)	<div><p>Boaedon virgatus (Boie, 1827)</p><p>Material. ANGOLA – Cabinda Province • Nganzi; –5.1028, 12.3709; 107 m a.s.l.; (shedding). • Cacata; –5.3200, 12.5022; 32 m a.s.l.; P1.296 (roadkill); GenBank: PQ455965.</p><p>Identification. The species is characterised by having a low number of ventral scales (&lt;198), &lt;25 dorsal scale rows at midbody,and a venter with a central broad greyish or yellowish stripe with darker lateral edges (Hallermann et al. 2020). The individual reported here from Cacata is almost identical (&lt;1% 16S p -distance) to a sample from Rabi, Gabon (GenBank: AY611825). In Angola, first Branch (2018) and later Hallermann et al. (2020) assigned three specimens from Cabinda Province reported by Peters (1877) to this species based on morphology. This represents the first genetically corroborated record for the species from Angola.</p><p>Biology and distribution. The species is widely distributed in West Africa, north of the Congo River, from Guinea Bissau to Cabinda Province and the Congo River in DRC. Very little is known about its biology.</p></div>	https://treatment.plazi.org/id/03E787B8FF80B721FF71348DFD949A35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFBFB71FFF7133A8FD309F75.text	03E787B8FFBFB71FFF7133A8FD309F75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cardioglossa leucomystax (Boulenger 1903)	<div><p>Cardioglossa leucomystax (Boulenger, 1903)</p><p>Figures 4E–F</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.189; GenBank: PQ455717. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.8937&amp;materialsCitation.latitude=-4.6323" title="Search Plazi for locations around (long 12.8937/lat -4.6323)">Miconge</a>; –4.4860, 12.8780; 377 m a.s.l.; FKH 1032−34, FKH 1052, FKH 1054; GenBank: PQ455712–16. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0916−18; GenBank: PQ455709–11.</p><p>Identification. Material from Cabinda Province is almost identical (0.2% 16S p -distance) to material ascribed to C. leucomystax 1 (sensu Blackburn et al. 2021) from the Bandjoko, Republic of the Congo and Ivindo, Gabon (GenBank: MW624175, MW624173),and identical to other material from Lekoumou, Republic of the Congo (GenBank: KY080101).Our records represent the southernmost records of the species, and the first for Angola (Baptista 2024).</p><p>Biology and distribution. The species appeared to be more common at higher elevations in Upper Mayombe, and all specimens were found near fast-flowing forest streams, either at the water edge or on sandbanks.</p></div>	https://treatment.plazi.org/id/03E787B8FFBFB71FFF7133A8FD309F75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF98B739FF7134E8FC169923.text	03E787B8FF98B739FF7134E8FC169923.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamaeleo dilepis Leach 1819	<div><p>Chamaeleo dilepis Leach, 1819</p><p>Figure 8C</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.6914&amp;materialsCitation.latitude=-4.7442" title="Search Plazi for locations around (long 12.6914/lat -4.7442)">Nganzi</a>; –5.5025, 12.3679; 107 m a.s.l.; EI 817; GenBank: PQ455967. • Mayombe NP, Mbongo Zimune; –4.7442, 12.6914; 324 m a.s.l.; P3.146; GenBank: PQ455966 .</p><p>Identification. The material reported in this work differed by 2.5% (16S p -distance) from material collected from Gabon (GenBank: AY927270) and by 3% from material from other specimens from Huila, Angola (GenBank: HF570444).Nevertheless,phylogenetic analyses have suggested cryptic speciation within the C.dilepis complex (Main et al.2022).Given that the type locality of C.dilepis is coastal Gabon,these specimens may represent true C.dilepis,so no further taxonomic actions are recommended for C.dilepis from Cabinda Province.It must be noted that Rio Cuilo in coastal Gabon is also the type locality of C. dilepis quilensis, but genetics analysis found this taxon to be conspecific with C. d. dilepis, leading to regarding C. d. quilensis to be regard as a synonym of C. d. dilepis (Main et al. 2018, 2022)</p><p>Biology and distribution. Chamaeleo dilepis is widely distributed in sub-Saharan Africa. In Angola,the species is also widely distributed,spanning a wide range of habitats and ecoregions (Main et al. 2022). In Cabinda Province, the species is frequently found in the Littoral Peneplain and the Coastal Drier Belt.However, one specimen was found in Lower Mayombe on a riverine area next to a village, among extensive plantations of introduced bamboo.</p></div>	https://treatment.plazi.org/id/03E787B8FF98B739FF7134E8FC169923	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF96B736FF713217FD8D9DA2.text	03E787B8FF96B736FF713217FD8D9DA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chiromantis rufescens (Gunther 1869)	<div><p>Chiromantis rufescens (Günther, 1869)</p><p>Figures 5C and 7T</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.8937&amp;materialsCitation.latitude=-4.6323" title="Search Plazi for locations around (long 12.8937/lat -4.6323)">Chimbaunde</a>; –5.5582, 12.4479; 139 m a.s.l.; FKH 0933; GenBank: PQ455718. • between Dinge and Chindende; –5.1208, 12.3667; 15 m a.s.l.; FKH 0175–76; GenBank: PQ455719–20. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.116, FKH 1260 (Tadpoles), P3.521–22; GenBank: PQ455721–24. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; P2.053; GenBank: PQ455725 .</p><p>Identification. Chiromantis rufescens is the only representative of the genus in the area and can easily be distinguished from other tree frogs by having rough skin, long fingers and toes with terminal discs and extensive webbing between fingers and toes (Channing and Rödel 2019). Chiromantis rufescens has very little genetic variation across the entire distribution range, except for the western Guinean populations, which have been suggested to represent a new taxon (Leaché et al. 2019). Material from Cabinda Province is almost identical (≤ 1% 16S p -distance) to material from Médouneu, Gabon (GenBank: MK789356), Tsinguidi, Republic of the Congo (GenBank: MK789354, KY080102) and Dzanga-Sangha NP, Central African Republic (GenBank: MK789330).</p><p>Biology and distribution. A foam-nest tree frog widely distributed in the northern rim of the Congo Basin and extending southwards to northern Angola. The species proved to be very common in the province and was found at most sites surveyed, from primary moist forest to transitional zones between forest and savannah and even highly transformed habitats. Specimens were usually found perched on branches or leaves near the ground to a few metres high, and never far from water. In Cabinda Province the species was recorded from all four ecological zones considered, from the Coastal Drier Belt to Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF96B736FF713217FD8D9DA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF80B720FF713335FD8298C3.text	03E787B8FF80B720FF713335FD8298C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dendroaspis jamesoni (Traill 1843)	<div><p>Dendroaspis jamesoni (Traill, 1843)</p><p>Figure 8U</p><p>Material. ANGOLA – Cabinda Province • near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.498538&amp;materialsCitation.latitude=-5.306833" title="Search Plazi for locations around (long 12.498538/lat -5.306833)">Nganzi</a>; –5.306833, 12.498538; 32 m a.s.l.; photographic record .</p><p>Identification. Dendroaspis jamesoni can be distinguished from D. viridis by having a larger number of dorsal scale rows at midbody (Chippaux and Jackson 2019), their unique coloration with yellow tail in adults, and its geographic distribution (see below).</p><p>Biology and distribution. A Central African forest specialist, widely distributed from Togo to Kenya in the east and southwards to Angola (Chippaux and Jackson 2019). In Angola, the species is widely distributed in the moister habitats of the escarpment and in the northeast (Branch 2018). In Cabinda Province, the species has been recorded in the Littoral Peneplain (this work) and the Coastal Drier Belt (Marques et al. 2018), but it is expected to be present throughout.</p></div>	https://treatment.plazi.org/id/03E787B8FF80B720FF713335FD8298C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9FB73FFF7130D7FC689F2B.text	03E787B8FF9FB73FFF7130D7FC689F2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipsadoboa weileri (Lindholm 1905)	<div><p>Dipsadoboa weileri (Lindholm, 1905)</p><p>Figure 8R</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4981&amp;materialsCitation.latitude=-4.6875" title="Search Plazi for locations around (long 12.4981/lat -4.6875)">Mbundu</a>; –4.6875, 12.4981; 127 m a.s.l.; FKH 1246; GenBank: PQ455968. • Mayombe NP, Sanga Wanda; -4.6270, 12.4527; 215 m a.s.l.; P3.511; GenBank: PQ455969.</p><p>Identification. The genus was morphologically revised by Rasmussen (1993). Specimens from Cabinda Province agree morphologically with D. weileri by having single subcaudals, dorsum dark grey to black and ventrum with uniform yellow colour with the exception of the subcaudal region that becomes black on the tip. The specimens reported here were genetically identified, differing by 2.8% (16S p -distance) from other conspecific material from Cross River National Park, Nigeria (GenBank: KX660288).</p><p>Biology and distribution. The species is known to occur from Guinea Bissau eastwards to South Sudan and southwards to the Republic of the Congo, being widely distributed in the Congo Basin (Rasmussen 1993). These represent the southernmost records of the species and the first for Angola (Branch 2018). Both individuals were found at night in rainforest in Lower Mayombe, one perched on a branch overhanging a small pond, and the other moving among leaf litter in a cleared area of the forest.</p></div>	https://treatment.plazi.org/id/03E787B8FF9FB73FFF7130D7FC689F2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9CB73CFF7135A0FB579D15.text	03E787B8FF9CB73CFF7135A0FB579D15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Feylinia grandisquamis Muller 1910	<div><p>Feylinia grandisquamis Müller, 1910</p><p>Figure 8K</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3233&amp;materialsCitation.latitude=-5.5213" title="Search Plazi for locations around (long 12.3233/lat -5.5213)">Nganzi</a>; –5.5213, 12.3233; 98 m a.s.l.; Photographic record .</p><p>Identification. A small legless skink (SVL = 150–200 mm) with 18–20 longitudinal scale rows across the body and the third supralabial scale in contact with the eye (Chirio and LeBreton 2007). The species was identified based on the previously mentioned morphological traits; however, it was originally described as a subspecies of F. currori (Müller 1910) . Therefore, the genetic validation of this species remains unclear.</p><p>Biology and distribution. A fossorial skink found in forest areas. The biology and geographic distribution of this species remain poorly known and needs further investigation.</p></div>	https://treatment.plazi.org/id/03E787B8FF9CB73CFF7135A0FB579D15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF99B73AFF713700FBDE9940.text	03E787B8FF99B73AFF713700FBDE9940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gerrhosaurus nigrolineatus Hallowell 1857	<div><p>Gerrhosaurus nigrolineatus Hallowell, 1857</p><p>Figure 8E</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.498538&amp;materialsCitation.latitude=-5.306833" title="Search Plazi for locations around (long 12.498538/lat -5.306833)">Tando Zinze</a>; –5.306833, 12.498538; 32 m a.s.l. ; P1.143; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3804&amp;materialsCitation.latitude=-5.1021" title="Search Plazi for locations around (long 12.3804/lat -5.1021)">GenBank</a>: PQ455970. • between Dinge and Chindende; –5.1021, 12.3804, 113 m a.s.l.; sight record. • Fazenda Mandarim; –5.0401, 12.0508; 14 m a.s.l.; P4.078, P4.078; GenBank: PQ455972.•Mayombe NP,Miconge; -4.4860, 12.8780; 377 m a.s.l.;P2.228; GenBank:PQ455971.</p><p>Identification. A large lizard (SVL = 140–175 mm) with stout body and head. Four supraciliary scales, 22–24 longitudinal dorsal scale rows (Bates et al. 2013). Dorsum light brown with scattered white lateral scales, and white and black lateral stripes from occipitals to tail, and usually light cream broken vertebral stripe. Specimens collected in Cabinda Province differ by 2% (16S p -distance) from material collected from Loulema, Republic of the Congo (GenBank: KF717399).</p><p>Biology and distribution. A widely distributed species in western and central Africa (Piettersen et al., 2021). An elusive and shy lizard that can be frequently found in bushveld and savannahs. In Cabinda Province, it is commonly encountered in the Littoral Peneplain area and Coastal Drier Belt, moving rapidly between grassland and bushes. However, the species was also recorded in patches of secondary forest at Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF99B73AFF713700FBDE9940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9FB73FFF713567FCF99D15.text	03E787B8FF9FB73FFF713567FCF99D15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grayia ornata (Bocage 1866)	<div><p>Grayia ornata (Bocage, 1866)</p><p>Figure 8S</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Chimbete; –4.6631, 12.5447; 182 m a.s.l.; P3.145; GenBank: PQ455973 .</p><p>Identification. A large snake (max. SVL = 1190 mm) with dorsum brown or grey with numerous black transverse bands (Chaney et al. 2024). Material from Cabinda Province is almost identical (&lt;0.5% 16S p -distance) to that of G. ornata sensu stricto (fide Chaney et al. 2024) from Ogooue, Gabon (GenBank: AY611866).</p><p>Biology and distribution. Grayia ornata is an aquatic snake widely distributed in Central Africa from Gabon to the westernmost region of the DRC and northern Angola (Chaney et al. 2024). The species is typically found in a variety of aquatic habitats (Chippaux and Jackson 2019). The specimen reported here was found at night in a deep pool at the edge of a slow-moving forest stream in Lower Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF9FB73FFF713567FCF99D15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9FB720FF713700FD3C9A9D.text	03E787B8FF9FB720FF713700FD3C9A9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hapsidophrys smaragdinus (Schlegel 1837)	<div><p>Hapsidophrys smaragdinus (Schlegel, 1837)</p><p>Figure 8T</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.498538&amp;materialsCitation.latitude=-5.306833" title="Search Plazi for locations around (long 12.498538/lat -5.306833)">Tando Zinze</a>; –5.306833, 12.498538; 32 m a.s.l.; P1.144; GenBank: PQ455974 .</p><p>Identification. The species can be distinguished from its congeners by having divided subcaudals and fewer than 175 ventral scales (Chippaux and Jackson 2019). The specimen reported here differs by ∼1% (16S p -distance) from other specimens from Mount Kupe, Cameroon (GenBank: KX671734) and Rabi, Gabon (GenBank: AY611875).</p><p>Biology and distribution. The species is widely distributed in West, Central and East Africa from Senegal to Kenya in the east and the Congo River mouth in the DRC and south to northern Angola (Chippaux and Jackson 2019). In Cabinda Province the specimen was found in degraded lowland riparian forest.</p></div>	https://treatment.plazi.org/id/03E787B8FF9FB720FF713700FD3C9A9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9AB73BFF713472FB579A9D.text	03E787B8FF9AB73BFF713472FB579A9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus mabouia (Moreau de Jonnes 1818)	<div><p>Hemidactylus mabouia (Moreau de Jonnès, 1818)</p><p>Figure 8G</p><p>Material. ANGOLA – Cabinda Province • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; P2.105; GenBank: PQ455975. • Fazenda Mandarim; –5.0401, 12.0508; 14 m a.s.l.; P4.068; GenBank: PQ455978. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P4.106; GenBank: PQ455979. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; P2.232– 33; GenBank: PQ455976–77.</p><p>Identification. A medium-sized Hemidactylus (SVL &lt;68 mm) that can be distinguished from other Angolan Hemidactylus based on the presence of enlarged subcaudal scales, absent in other close relatives. The specimens collected at Mayombe NP are identical (16S p -distance) to other individuals from Angola, which represent H. mabouia sensu stricto (fide Agarwal et al. 2021). However, the specimens from Caio Cacongo differ by 1.3% (16S p -distance) from other relatives from Cabinda Province and other regions of Angola, which may suggest a posterior colonisation event of this invasive species.</p><p>Biology and distribution. Hemidactylus mabouia is a nocturnal gecko widely distributed across Africa, Madagascar and the Neotropics (Agarwal et al. 2021). Frequently associated with human infrastructure, the species can also be found in trees at night near villages or construction. All material from Cabinda Province was collected while active at night in or next to human infrastructure, such as walls, fences and planted trees, in villages and farms.</p></div>	https://treatment.plazi.org/id/03E787B8FF9AB73BFF713472FB579A9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9BB73BFF7130F8FD969D9B.text	03E787B8FF9BB73BFF7130F8FD969D9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemidactylus pfindaensis Lobon-Rovira, Conradie, Buckley, Ernst, Verissimo, Baptista and Vaz Pinto 2021	<div><p>Hemidactylus pfindaensis Lobón-Rovira, Conradie, Buckley, Ernst, Veríssimo, Baptista and Vaz Pinto, 2021</p><p>Figures 8H–I</p><p>Material. ANGOLA – Cabinda Province • Nganzi; –5.5213, 12.3233; 98 m a.s.l.; EI 1135; GenBank: PQ455980. • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; FKH 0980–82; GenBank: PQ455981–83. • Fazenda Mandarim; –5.0401, 12.0508; 14 m a.s.l.; P4.067, P4.107−08; GenBank: PQ455985–87. • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3667&amp;materialsCitation.latitude=-5.1208" title="Search Plazi for locations around (long 12.3667/lat -5.1208)">between Dinge and Chindende</a>; –5.1208, 12.3667; 15 m a.s.l.; FKH 0178–79; GenBank: PQ455988–89. • Lagoa Massabi; –5.0603, 12.0488; 10 m a.s.l.; FKH 0984–86, P2.120; GenBank: PQ455990–92, PQ455984. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; P2.106; GenBank: PQ455993. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.131; GenBank: PQ455994. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.105–06, P3.194, P3.510; GenBank: PQ455995–98. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1061; GenBank: PQ455999. • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.152; GenBank: PQ456000. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0975, FKH 0983; GenBank: PQ456001–02.</p><p>Identification. A medium-sized Hemidactylus (max. SVL = 49 mm) with a moderately long snout (Lobón-Rovira et al. 2021). The species usually has a characteristic black spot in the occipital region; however, some specimens from the Coastal Drier Belt do not have this black spot. The specimens reported here are segregated into four different lineages based on 16S, which differ among themselves by &lt;2.8%. Each lineage corresponds to the four ecological zones considered here (Upper Mayombe, Lower Mayombe, Littoral Peneplain and Coastal Drier Belt), with the lineage from Upper Mayombe being the most divergent, differing by a maximum of 2.8% from conspecific material. Material from Cabinda Province is topotypic and differs by 5.5−6% (16S p -distance) from material collected from Macocola, Uíge Province (FKH 004, see Lobón-Rovira et al. 2021).</p><p>Biology and distribution. A nocturnal, arboreal gecko frequently found on tree branches or large leaves. We recorded material from all four ecological zones considered in this work. However, at Fazenda Mandarim, where the species was found in syntopy with H. mabouia, individuals were actively hunting on the floor among leaf litter, perhaps a consequence of ecological competition with H. mabouia . The species is known to occur in Cabinda Province and Uíge Province, Angola (Lobón-Rovira et al. 2021). It is the most abundant gecko in Cabinda Province, having been collected or observed in most surveyed sites. It is expected to occur in the neighbouring areas of the Republic of the Congo and DRC.</p></div>	https://treatment.plazi.org/id/03E787B8FF9BB73BFF7130F8FD969D9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9AB73AFF713387FD259FDD.text	03E787B8FF9AB73AFF713387FD259FDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holaspis guentheri Gray 1863	<div><p>Holaspis guentheri Gray, 1863</p><p>Figure 8F</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; Photographic record .</p><p>Identification. A relatively small lizard (SVL = 45–50 mm) with flattened head and body. Head elongated with pointed snout. Dorsum black, bisected by two lateral bright bluishcoloured lines that become creamy-yellow on the head, belly and sides of the tail. This species can be distinguished from its sister taxon ( H. laevis) by having four yellowbeige lateral stripes (versus six in H. laevis) and thus we refer our material to H. guentheri .</p><p>Biology and distribution. Holaspis guentheri is an arboreal rainforest dweller found actively moving during the day on tree trunks (Spawls et al. 2018). The species is widely distributed in forests of West and Central Africa, with some scattered records in Uganda, Tanzania and Angola (Marques et al. 2018; Spawls et al. 2018). In this study, we observed and photographed several individuals moving up and down the trunks of large trees, and once on a very large fallen log in a clearing in mature rainforest which was subject to logging activities in Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF9AB73AFF713387FD259FDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA7B707FF71334DFE109E18.text	03E787B8FFA7B707FF71334DFE109E18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplobatrachus occipitalis (Gunther 1858)	<div><p>Hoplobatrachus occipitalis (Günther, 1858)</p><p>Figure 4W</p><p>Material. ANGOLA – Cabinda Province • Fazenda Mandarim; –5.0401, 12.0508; 14 m a.s.l.; P4.069; GenBank: PQ455727. • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3667&amp;materialsCitation.latitude=-5.1208" title="Search Plazi for locations around (long 12.3667/lat -5.1208)">between Dinge and Chindende</a>; –5.1208, 12.3667; 15 m a.s.l.; FKH 0177; GenBank: PQ455726. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; –4.6403, 12.935; 269 m a.s.l.; sight record.</p><p>Identification. A large frog (104–160 mm) and sole representative of the genus in Africa, with a characteristic transverse groove behind the eyes, visible supratympanic folds and eyes positioned on the top of the head. Specimens from Cabinda Province are genetically identical to material from Malanje Province, Angola (GenBank: MK036439), Tanzania (GenBank: AB272600) and Nigeria (GenBank: MH708922).</p><p>Biology and distribution. Hoplobatrachus occipitalis is widely distributed in the sub-Saharan tropical savannahs from Senegal to Ethiopia, Kenya and Tanzania in the east, and northern and western Angola in the south (Channing and Rödel 2019). This species is likely common throughout Cabinda Province, and across most habitats excluding the interior of intact primary forests. This material was collected during the rainy season in temporary pools in highly disturbed areas, in farmland and urban environments, in both the Coastal Drier Belt and Littoral Peneplain. However, on a different occasion we also observed and recorded a species chorus in a temporary pool in partially cleared areas in Lower Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FFA7B707FF71334DFE109E18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFAEB70EFF7131C8FD84983D.text	03E787B8FFAEB70EFF7131C8FD84983D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hylambates leonardi Boulenger 1906	<div><p>Hylambates leonardi Boulenger, 1906</p><p>Figure 6T</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0967; GenBank: PQ455728. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1230; GenBank: PQ455729. • Sanga Wanda entrance; –4.6304, 12.4714; 203 m a.s.l.; P3.528; GenBank: PQ455730.</p><p>Identification. A large hyperoliid (SVL = 54 mm) that can be distinguished from other members in the family by its characteristic black and yellow-orange stripes on the hind limbs and brownish dorsum. Specimens from the study area are identical (16S p -distance) to material from Gamba, Gabon (GenBank: KY080267), ∼ 300 km from N’Djolè, where a lectotype was restricted (Frétey et al. 2014).</p><p>Biology and distribution. A nocturnal tree frog that is known to occur in Equatorial Guinea, Gabon, Republic of the Congo and DRC (Channing and Rödel 2019). This material represents the first records of this genus for Angola and Cabinda Province (Baptista 2024). On one occasion, specimens in this study were observed jumping to the ground from 4 m high in the forest during a rainstorm, and breeding in a deep rain pool at the edge of the primary rainforest, in Lower Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FFAEB70EFF7131C8FD84983D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF95B735FF7130BFFF359D15.text	03E787B8FF95B735FF7130BFFF359D15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hylarana (Amnirana) albolabris (Hallowell 1856)	<div><p>Hylarana (Amnirana) albolabris (Hallowell, 1856)</p><p>Figures 5E and 7R</p><p>Material. ANGOLA – Cabinda Province • Fazenda Mandarim; –5.0553, 12.0610; 14 m a.s.l.; P4.074, P4.076; GenBank: PQ455636–37. • Tando Zinze; –5.3241, 12.5073; 29 m a.s.l.; P1.140; GenBank: PQ455634. • Mayombe NP, Chimbete; –4.6631, 12.5447; 182 m a.s.l.; P3.133; GenBank: PQ455635. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.212 (Tadpoles), FKH 1267 (Tadpoles), P3.222, P4.082 (Tadpoles); GenBank: PQ455638–41. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.115, P3.188, P3.195 (Eggs); GenBank: PQ455642–44. • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.154; GenBank: PQ455645. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1024, FKH 1026−28, FKH 1057−58, P2.211, FKH 1254, P3.187; GenBank: PQ455646–54.</p><p>Identification. A medium to large (SVL = 57–74 mm) white-lipped frog. The species can be distinguished from other Hylarana species present on Cabinda Province ( H. lepus) by its smaller size, males having large, elongated glands on the forelimbs (versus small and oval in H. lepus) and the presence of narrow and continuous glandular dorsolateral ridges from the posterior part of the eye to midbody (versus absent in H. lepus). Jongsma et al. (2018) reported potential cryptic diversification within this taxon, with one distinctive species in West Africa (which represents an undescribed taxon) and two well-defined lineages in the nominotypical lineage from north and south of the Congo River, respectively. Our material clusters (16S p -distance) with sequences from north of the Congo River (e.g., Republic of the Congo [GenBank: KY080037], Gabon [GenBank: KX289626], Cameroon [GenBank: MG 552470] and Nigeria [GenBank: MG 552474]), some collected within 50 km of the type locality (Jongsma et al. 2018). Therefore, we regard our material to be conspecific with H. albolabris sensu stricto .</p><p>Biology and distribution. The nominotypical lineage of H. albolabris is known to occur across the lowlands and mid-elevation forest along the northern rim of the Congo Basin, including Nigeria, Cameroon, Gabon, Equatorial Guinea, Republic of the Congo, northwestern DRC and Cabinda Province (Jongsma et al. 2018). It represents a forest-dwelling species frequently found near streams and ponds. Specimens from Cabinda Province were found at night in gallery, primary and secondary moist forests, which were often degraded and across the study area from the Coastal Drier Belt to Upper Mayombe. Individuals were typically found perched in branches near water ∼ 1 m above the ground or water.</p></div>	https://treatment.plazi.org/id/03E787B8FF95B735FF7130BFFF359D15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF95B736FF713700FE7699A8.text	03E787B8FF95B736FF713700FE7699A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hylarana (Amnirana) lepus (Andersson 1903)	<div><p>Hylarana (Amnirana) aff. lepus (Andersson, 1903)</p><p>Figure 7S</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1025, FKH 1029, FKH 1056; GenBank: PQ455655−57. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0935−37; GenBank: PQ455658−00. • Mayombe NP, Lombe River, Vaku; -4.6403, 12.935; 269 m a.s.l.; P3.159; GenBank: PQ455661 .</p><p>Identification. A large (SVL = 70–100 mm) white-lipped frog with small oval glands on the forelimbs in males and granular skin across the entire body. The material from Cabinda Province is identical (16S p -distance) to material from Lekoumou, Republic of the Congo (GenBank: KY080044, KY080047) and DRC (GenBank: MG552243), which was ascribed by Jongsma et al. (2018) to an undescribed species ( Hylarana “lepus 1”). Thus, we consider the newly collected material to be conspecific with the material of Jongsma et al. (2018), here referred to as H. aff. lepus .</p><p>Biology and distribution. Hylarana lepus sensu lato is distributed across the Congo Basin and Gulf of Guinea from Cameroon to Angola (Channing and Rödel 2019), with two separate lineages from either side of the Congo River (Jongsma et al. 2018). In Cabinda Province, the species was only found in primary moist forest at Upper Mayombe between 260–400 m a.s.l. perched low on branches over water in fast-flowing forest streams.</p></div>	https://treatment.plazi.org/id/03E787B8FF95B736FF713700FE7699A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF92B732FF7133E5FB8A9FDD.text	03E787B8FF92B732FF7133E5FB8A9FDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hymenochirus feae (Boulenger 1906)	<div><p>Hymenochirus feae (Boulenger, 1906)</p><p>Figure 7M</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Chilito, Mbundo; –4.7196, 12.5110; 336 m a.s.l.; P3.544; GenBank: PQ455731 .</p><p>Identification. A medium-sized (SVL = 42–46 mm) flat aquatic frog with characteristic fingers and toes characteristically fully webbed to the tips (Boulenger 1906). Large heads, dark brown to black dorsal coloration, different wart sizes along the body,and two distinctive black claws on the feet. The specimen reported here represents the first genetic sample of the species,which differs by 3.4–3.6% from H. boettgeri from Mbemba and Simombondo, Republic of the Congo (GenBank: OR360735 and KY080144, respectively).</p><p>Biology and distribution. Previously only known from Gabon. However, Nagy et al. (2013) reported Hymenochirus sp. aff. feae from Luki, Bas-Congo, DRC, which may represent the same species. This record represents a new genus for Angola (Baptista 2024). The specimen reported here was found in a small pond under permanent shade in a deep valley, surrounded by degraded and secondary forest in Lower Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF92B732FF7133E5FB8A9FDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFAAB70AFF7130D7FF259FB0.text	03E787B8FFAAB70AFF7130D7FF259FB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius adspersus Peters 1877	<div><p>Hyperolius adspersus Peters, 1877</p><p>Figure 6G</p><p>Material. ANGOLA – Cabinda Province • Tando Zinze; –5.3241, 12.5073; 29 m a.s.l.; FKH 0583–84, FKH 0693; GenBank: PQ455732–34.</p><p>Identification. Hyperolius adspersus is part of a larger H. nasutus species complex of small (SVL = 19−23 mm) green reed frogs with highly conserved morphology and basal to many Hyperolius species (Channing et al. 2013). In Angola, the group consisting of ‘little green frogs’ remains unresolved, comprising four highly divergent clades, two of which can be confidently ascribed to H. nasutus and H. benguellensis as they include topotypic material (Baptista 2024). Quite a lot of structure is present within Angolan H. nasutus, but with genetic distance between different lineages below 2% (16S p-distance) (Baptista 2024). Hyperolius adspersus was originally described from Chinchoxo, Cabinda Province (Peters, 1877). The material reported here clusters with material of H. adspersus from Plain of Vera (15 km SE Gamba), Gabon (GenBank: JQ863695), from which it differs by 1.2% (16S p -distance). However, it also differs from the topotypic material of H. nasutus from Calandula (GenBank: JQ863641) by 1.9%, thus raising doubts about the taxonomic status of H. adspersus .</p><p>Biology and distribution. Hyperolius adspersus is known to occur in the western lowlands of Central Africa, north of the Congo River (Channing et al. 2013). The only specimen collected in this study was found on vegetation near water in highly impacted farmland habitat.</p></div>	https://treatment.plazi.org/id/03E787B8FFAAB70AFF7130D7FF259FB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFAAB70BFF7135DDFE469878.text	03E787B8FFAAB70BFF7135DDFE469878.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius ocellatus subsp. purpurescens Gunther 1858	<div><p>Hyperolius ocellatus purpurescens Günther, 1858</p><p>Figures 5B and 6H–I</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.8937&amp;materialsCitation.latitude=-4.6323" title="Search Plazi for locations around (long 12.8937/lat -4.6323)">Dinge</a> and Chindende; –5.1208, 12.3667; 15 m a.s.l.; FKH 0166; GenBank: PQ455735. • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; P2.108; GenBank: PQ455736. • Mayombe NP, Sanga Mongo; -4.6661, 12.4407; 146 m a.s.l.; FKH 0953, FKH 0957−58, FKH 0971−73; GenBank: PQ455737–42. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; FKH 1234, FKH 1266 (Tadpoles), FKH 1268−69, P3.225, P4.081 (Tadpoles); GenBank: PQ455743–48. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.192; GenBank: PQ455749. • Mayombe NP, Entrance Sanga Wanda; –4.6304, 12.4714; 219 m a.s.l.; P3.534−5; GenBank: PQ455750−51. • Mayombe NP, Chimbete; -4.6631, 12.5447; 182 m a.s.l.; FKH 1240, FKH 1242; GenBank: PQ455752–53. • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.153; GenBank: PQ455754. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0947; GenBank: PQ455755 .</p><p>Identification. Hyperolius ocellatus is a medium-sized reed frog (SVL = 27−34 mm) with pronounced sexual dimorphism in size and colour pattern, displaying several documented female morphotypes, and with two subspecies currently recognised (Bell et al. 2017). Males are usually green with a white triangular mask between the top of the eyes and the snout, and two white dorsolateral bands from the eyes to the anterior insertion of the hind limb (Figure 6I). Our female specimens correspond to the morphotype in which the females have a bright yellow ventrum, and the dorsal and limb colouration is greyish to white with characteristic black “ocelli” (Figure 6H). The species was described from Bioko Island, Equatorial Guinea, and Angola (Laurent, 1943). However, Perret (1975) restricted the type locality to Bioko Island, Equatorial Guinea, to bring taxonomic stability to the species. Material from Cabinda Province differs from topotypic material of H. ocellatus (GenBank: MK509614) by 3.2% (16S p -distance). It clusters with a clade ascribed to Hyperolius ocellatus purpurescens from Ivindo, Gabon and Cuvette, Republic of the Congo (Bell et al. 2017), from which it differs by ∼2.5% and ∼1.5% (16S p -distance), respectively (GenBank: MF376543, MF376630). Consequently, we regard the material from Cabinda Province to be conspecific with H. o. purpurescens.</p><p>Biology and distribution. This species complex is widely distributed in the Gulf of Guinea and the Congo Basin, including Bioko Island. In Cabinda Province, the species appears to be relatively common and widely distributed in forested habitats, and only absent from the Coastal Drier Belt. It was typically found calling from vegetation near water in primary and secondary forest, and often in sympatry with other Hyperolius species reported in this work.</p></div>	https://treatment.plazi.org/id/03E787B8FFAAB70BFF7135DDFE469878	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFABB70CFF713295FCCE9905.text	03E787B8FFABB70CFF713295FCCE9905.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius olivaceus Peters 1876	<div><p>Hyperolius olivaceus Peters, 1876</p><p>Figures 6J–M</p><p>Material. ANGOLA – Cabinda Province • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; P2.109; GenBank: PQ455758. • Lagoa Massabi; –5.0603, 12.0488; 10 m a.s.l.; P2.121; GenBank: PQ455759. • Fazenda Mandarim; –5.0553, 12.0610; 14 m a.s.l.; P4.079−80; GenBank: PQ455756–57. • Tando Zinze; –5.3241, 12.5073; 29 m a.s.l.; P1.139, FKH 0585, FKH 0694; GenBank: PQ455760–62. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0952, FKH 0964−65, P2.087−88, P2.092−93; GenBank: PQ455763–69. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; FKH 1235, P4.097; GenBank: PQ455770, PQ455774. • Mayombe NP, Chilito Mbundu; –4.7280, 12.50681; 85 m a.s.l.; P3.541; GenBank: PQ455771. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1263; GenBank: PQ455772. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.094; GenBank: PQ455773. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1055, FKH 1059−60, P3.173; GenBank: PQ455775–78. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0925−26, FKH 0944; GenBank: PQ455779–81. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; –4.6403, 12.935; 269 m a.s.l.; P3.161; GenBank: PQ455782.</p><p>Identification. A medium-sized reed frog (SVL = 28−33 mm) which can display polymorphic colour patterns. Most specimens from Cabinda Province are rather large, do not display sexual dimorphism, and are typically bright green-yellow at night (Figure 6J), turning darker green during the day, with bright red inner thighs (Figure 6K). Some male specimens may be orangey to dark brown (Figure 6L), and rarely some morphs are mottled displaying a dorsal hourglass pattern resembling that of H. platyceps (Figure 6M). Material recovered from the study area is genetically identical (16S p -distance) to material from Lekoumou and Kouilou, Republic of the Congo (GenBank: KY080172, KY080174), while only differing by 1.5% (16S p -distance) from material from Sahoue, Gabon (Genbank: KY080177), ∼ 150 km from the type locality.</p><p>Biology and distribution. The species is distributed from Equatorial Guinea to northern Angola (Channing and Rödel 2019; Ernst et al. 2020). In Cabinda Province, H. olivaceus is the most abundant species of Hyperolius and was recorded in almost all sites surveyed, and across all four ecological zones considered. The species was found in many different habitats, from mangroves (sea level) to highly disturbed savannah and farmland, to moist rainforest at higher elevations (above 300 m a.s.l.). It was also recorded in sympatry with all the other Hyperolius reported in this work.</p></div>	https://treatment.plazi.org/id/03E787B8FFABB70CFF713295FCCE9905	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFACB70CFF713310FD679FDD.text	03E787B8FFACB70CFF713310FD679FDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius parallelus Gunther 1858	<div><p>Hyperolius parallelus Günther, 1858</p><p>Figure 6N</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3667&amp;materialsCitation.latitude=-5.1208" title="Search Plazi for locations around (long 12.3667/lat -5.1208)">Dinge</a> and Chindende; –5.1208, 12.3667; 15 m a.s.l.; FKH 0696; GenBank: PQ455783 .</p><p>Identification. Part of the H. viridiflavus complex, that in Angola includes material previously assigned to H. angolensis, now ascribed to H. parallelus (Channing 2022) . While juveniles and some males display a drab colour pattern (Phase J), two very different morphotypes are reported in adult females and some males (Phase F), namely a colourful red-marbled dorsum on the Angolan plateau, or with two broad dark dorsal stripes in populations on the coastal regions and up to the western escarpment forests. The material reported here differs by a minimum of 1.8% (16S p -distance) from other material in Angola (Channing 2022). The single specimen sampled in this work displays a “lined form” (Figure 6N), in agreement with specimens reported from the coastal region of Angola (Channing 2022).</p><p>Biology and distribution. Widely distributed in the Congo Basin and the savannahs south of the Congo River, including DRC, Zambia, Angola, and western Tanzania (Channing 2022). In Angola, it is widely distributed across the entire territory (Marques et al. 2018; Baptista 2024). In Cabinda Province, the species has only been recorded in the Littoral Peneplain (this work), in addition to some historical records on the Coastal Drier Belt at Landana (Marques et al. 2018). The specimen reported here was found on vegetation surrounding a pond in a highly disturbed area.</p></div>	https://treatment.plazi.org/id/03E787B8FFACB70CFF713310FD679FDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFACB70DFF713438FDCE983D.text	03E787B8FFACB70DFF713438FDCE983D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius pardalis Laurent 1948	<div><p>Hyperolius pardalis Laurent, 1948</p><p>Figures 6O–Q</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4597&amp;materialsCitation.latitude=-4.5981" title="Search Plazi for locations around (long 12.4597/lat -4.5981)">Dinge</a> and Chindende; –5.1208, 12.3667; 15 m a.s.l.; FKH 0161–62, FKH 0164, P9.063, FKH 0168; GenBank: PQ455784–88. • Mayombe NP, Chimbete; –4.6631, 12.5447; 182 m a.s.l.; FKH 1241; GenBank: PQ455789. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0962−63; GenBank: PQ455790–91. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.093; GenBank: PQ455792 .</p><p>Identification. Hyperolius pardalis is a medium-sized reed frog (SVL = 26−33 mm) within the H. steindachneri complex (sensu Baptista 2024), and is highly polymorphic. We recorded two different morphotypes in both males and females. Based on the strikingly different female patterns we can tentatively recognise two subpopulations, a western subpopulation found only in gallery forest in the Littoral Peneplain, and an eastern subpopulation found in several rainforest sites across Lower Mayombe. Males are characterised by a dark canthal line, white transversal line defining the base of the gular sac, reddish toe and finger discs and usually orange-brown dorsum, sometimes with a yellowish tinge, covered with darker spots (Figure 6O). In the eastern subpopulation females had the same black ventrum with white spots, but the dorsal parts displayed a goldenblack marbled pattern (Figure 6P). In the western subpopulation, one collected male was yellowish-green with white dorsolateral stripes, while all females were black with bright yellowish to orange spots on the dorsum, and white spots on the ventrum (Fig. 6Q). All material collected in Cabinda Province showed little genetic divergence (&lt;0.8% 16S p -distance) despite striking colour differences. In addition, specimens reported here were assigned to H. pardalis because they only differ by 1.2% (16S p -distance) from material collected from Pool, Republic of the Congo (GenBank: KY080201) and by 1.5% (16S p -distance) from material collected from Mambele, Cameroon (GenBank: MK509657), ∼ 340 km east of the type locality (Bitye, Cameroon).</p><p>Biology and distribution. Widely distributed in West-Central Africa, from southern Cameroon and the Central African Republic to northern Angola (Channing and Rödel 2019). All specimens collected in this work were found in thick vegetation near deep water pools in mature forest habitat. The geographical segregation between the two morphotypes, albeit apparently not supported genetically, suggests some phenotypic plasticity and/or ecological segregation that needs further investigation. The species was not recorded in Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FFACB70DFF713438FDCE983D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFADB70EFF713258FD2B9BCD.text	03E787B8FFADB70EFF713258FD2B9BCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyperolius platyceps (Boulenger 1900)	<div><p>Hyperolius platyceps (Boulenger, 1900)</p><p>Figures 6R–S</p><p>Material. ANGOLA – Cabinda Province • Fazenda Mandarim; –5.0553, 12.0610; 14 m a.s.l.; P4.077; GenBank: PQ455793. • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; FKH 0977; GenBank: PQ455796. • Tando Zinze; –5.3241, 12.5073; 29 m a.s.l.; FKH 0582; GenBank: PQ455797. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4407&amp;materialsCitation.latitude=-4.6661" title="Search Plazi for locations around (long 12.4407/lat -4.6661)">Sanga Mongo</a>; –4.6661, 12.4407; 146 m a.s.l.; FKH 0949, FKH 0956, FKH 0974; GenBank: PQ455798–00. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4527&amp;materialsCitation.latitude=-4.627" title="Search Plazi for locations around (long 12.4527/lat -4.627)">Sanga Wanda</a>; –4.6270, 12.4527; 210 m a.s.l.; P3.519; GenBank: PQ455801. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4597&amp;materialsCitation.latitude=-4.5981" title="Search Plazi for locations around (long 12.4597/lat -4.5981)">Bata Lunhuca</a>; –4.5981, 12.4597; 215 m a.s.l.; P4.095−96; GenBank: PQ455794–95. • Mayombe NP, Miconge; –4860, 12.8780; 377 m a.s.l.; P3.174; GenBank: PQ455802.</p><p>Identification. A medium-sized (SVL = 33−40 mm) reed frog with two very different morphotypes. They either have dark dorsal blotches against a light background forming an hourglass pattern with a transverse band on the lower back (Figure 6R), or a dark green to brown dorsum with two broad dorsolateral white bands from snout to anterior insertion of the hind limbs, and yellow underparts (Figure 6S). Specimens from Cabinda Province differ from material collected from Talangai, Republic of the Congo (GenBank: MK509613), Ndombam, Cameroon (GenBank: MK509701) and Iguéla, Gabon (GenBank: MK509680) by&gt;3% (16S p -distance), and from unpublished material from the remainder of Angola by a minimum of 2.8% (16S p -distance) (Baptista 2024). The large genetic divergence may suggest some cryptic diversification. However, genetic material from the type locality is not available, therefore no taxonomic action can be taken at this point.</p><p>Biology and distribution. Widely distributed in West-Central Africa, from southern Cameroon to northern Angola (Channing and Rödel 2019). The species is reportedly associated with degraded forest and swampy forest areas (Channing and Rödel 2019), and in Cabinda Province it has been found in all four ecological zones considered, from sea level to around 400 m a.s.l., including primary moist forest and lowland degraded forest. Frequently found in syntopy with H. olivaceus .</p></div>	https://treatment.plazi.org/id/03E787B8FFADB70EFF713258FD2B9BCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF98B738FF7131CAFBE0998C.text	03E787B8FF98B738FF7131CAFBE0998C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kinixys erosa (Schweigger 1812)	<div><p>Kinixys erosa (Schweigger, 1812)</p><p>Figure 8A</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.444&amp;materialsCitation.latitude=-4.859" title="Search Plazi for locations around (long 12.444/lat -4.859)">Inhuca</a>; –4.8590, 12.4440; 81 m a.s.l.; P2.208; GenBank: PQ456003. • Chilito, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.5068&amp;materialsCitation.latitude=-4.7279" title="Search Plazi for locations around (long 12.5068/lat -4.7279)">Mbundo</a>; –4.7279, 12.5068; 83 m a.s.l.; P3.543; GenBank: PQ456004.</p><p>Identification. A large tortoise with spiny and upturned marginal scutes (Branch 2008). Specimens collected in Cabinda Province differ by&gt;2% (16S p -distance) from material collected from Equateur, DRC (GenBank: LR697076) and by 3.5% from its sister species Kinixys homeana (GenBank: DQ 424975).</p><p>Biology and distribution. A rainforest dweller widely distributed across West and Central Africa, from Guinea and Sierra Leone to Uganda and southwards to northern Angola (Sánchez-Vialas et al. 2022). We sampled two individuals caught by local people in primary rainforest and for human consumption, afterwards confiscated by rangers and released in Mayombe NP.</p></div>	https://treatment.plazi.org/id/03E787B8FF98B738FF7131CAFBE0998C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFBFB700FF7135A0FED99D7C.text	03E787B8FFBFB700FF7135A0FED99D7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis aubryi (Dumeril 1856)	<div><p>Leptopelis aubryi (Dumeril, 1856)</p><p>Figures 4G and 5A</p><p>Material. ANGOLA – Cabinda Province • between Dinge and Chindende; –5.1208, 12.3667; 15 m a.s.l.; FKH 0171–73; GenBank: PQ455803–05. • Fazenda Mandarim; – 5.0553, 12.0610; 14 m a.s.l.; P4.072; GenBank: PQ455845. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0954−55, FKH 0961, FKH 0968, P2.091; GenBank: PQ455808–12. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.214, P3.223, P4.098, P4.103–04; GenBank: PQ455813–14, PQ455847–49. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1225−28, FKH 1261; GenBank: PQ455822– 26. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.086; GenBank: PQ455846. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1030, P2.231, P3.172; GenBank: PQ455834–36. • Mayombe NP, Vaku; –.6323, 12.8937; 319 m a.s.l.; FKH 0930, FKH 0939; GenBank: PQ455837–38. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; – 4.6403, 12.935; 269 m a.s.l.; FKH 1247, FKH 1249−50, P3.166, P3.169, FKH 1252, GenBank: PQ455839–44.</p><p>Identification. A small Leptopelis (SVL = 42–59 mm) with an orange upper margin to the eyes, often with a dark triangle between the eyes (Channing and Rödel 2019). The specimens here reported are identical (16S p -distance) to material from Lekoumou, Republic of the Congo (GenBank: KY080240) and almost identical to material from Tchimbélé, Gabon (0.4% 16S p -distances; GenBank: KF888326).</p><p>Biology and distribution. The species is widely distributed in the Gulf of Guinea from Nigeria to northern Angola (Marques et al. 2018; Channing and Rödel 2019; Ernst et al. 2020; Baptista 2024). The species can be found in branches near swampy areas, ponds and riverine areas in primary forest, but also in secondary forest and degraded forest edges (Channing and Rödel 2019). In Cabinda Province, it is the most abundant Leptopelis species, present in all four main ecological zones. Often found low in bushes, leaves or branches, and in sympatry with all the other Leptopelis species recorded in the province.</p></div>	https://treatment.plazi.org/id/03E787B8FFBFB700FF7135A0FED99D7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA1B701FF7131B0FC5699D5.text	03E787B8FFA1B701FF7131B0FC5699D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis aubryioides (Andersson 1907)	<div><p>Leptopelis cf. aubryioides (Andersson, 1907)</p><p>Figure 4H</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.514; GenBank: PQ455850 .</p><p>Identification. This is the smallest Leptopelis from Cabinda Province (SVL = 35−45 mm). It is slenderer than L. aubryi, lacks orange colouration on the eyes and has white spurs on the heels. The specimen reported from Cabinda Province differs by 2.2% (16S p -distance) from material collected at Lekoumou, Republic of the Congo (GenBank: KY080248), and by 4% from material collected at Mount Kupe, Cameroon (GenBank: KX671754). The large genetic distance may suggest cryptic diversification in the group; however, further analyses are needed to take taxonomic action. Consequently, we list the material from Cabinda Province as Leptopelis cf. aubryioides, pending further investigation.</p><p>Biology and distribution. An arboreal Leptopelis widely distributed in dense forests across the Gulf of Guinea (Channing and Rödel 2019). This material represents the first record for Cabinda Province, and for Angola (Baptista 2024). This specimen was found in a primary rainforest on a leaf about 4 m above ground in Lower Mayombe, and in sympatry with L. aubryi, L. boulengeri, L. cf. calcaratus and L. ocellatus .</p></div>	https://treatment.plazi.org/id/03E787B8FFA1B701FF7131B0FC5699D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA1B701FF7133C0FC0B9FFB.text	03E787B8FFA1B701FF7133C0FC0B9FFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis boulengeri (Werner 1898)	<div><p>Leptopelis boulengeri (Werner, 1898)</p><p>Figure 4I</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.203, P3.523; GenBank: PQ455851–52 .</p><p>Identification. A large-sized Leptopelis (SVL = 48–81 mm), with characteristic angular region between the nostrils and eyes, tympanum barely visible and white spot below the eyes (Channing and Rödel 2019). The specimens from this work are identical (16S p -distance) to material from Minkébé, Gabon (GenBank: KX260263). However, they differ by ∼1.6% (16S p -distance) from material from Mount Kupe, Cameroon (GenBank: KX671756), &lt;90 km from the type locality ( Victoria [now Limbé], Cameroon).</p><p>Biology and distribution. An arboreal species, widely distributed in the Gulf of Guinea, including Bioko Island (Channing and Rödel 2019). These records represent a range extension south from the previously reported southernmost record of Lekoumou, Republic of Congo, and represent the first country records (Baptista 2024). Specimens were collected from leaves between 2 and 3 m above ground in the primary rainforest.</p></div>	https://treatment.plazi.org/id/03E787B8FFA1B701FF7133C0FC0B9FFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA1B702FF713417FBDE9A08.text	03E787B8FFA1B702FF713417FBDE9A08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis calcaratus (Boulenger 1906)	<div><p>Leptopelis cf. calcaratus (Boulenger, 1906)</p><p>Figure 4J</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6270, 12.4527; 215 m a.s.l.; P3.513, P3.515; GenBank: PQ455853–54. • Mayombe NP, Bata Lunhuca; – 4.5981, 12.4597; 215 m a.s.l.; P4.084; GenBank: PQ455855.</p><p>Identification. A medium-sized Leptopelis (SVL = 42–57 mm) with characteristic white spurs on the heels. The specimens from Cabinda Province differ by 3.8–4% (16S p -distance) from material collected from Sapia, Cameroon (GenBank: KT967090, KT967091), ∼ 100 km south of the type locality at Efoulan, Cameroon. The large genetic distance suggests cryptic diversification within L. calcaratus . We therefore listed the material from Cabinda Province as Leptopelis cf. calcaratus, pending further investigation.</p><p>Biology and distribution. An arboreal Leptopelis widely distributed in the Gulf of Guinea and the Congo Basin,from Nigeria eastwards to DRC (Channing and Rödel 2019). The species had recently been reported from northwestern Angola (Ernst et al.2020; Baptista 2024), but these specimens constitute the first records of the species for Cabinda (Baptista 2024).</p></div>	https://treatment.plazi.org/id/03E787B8FFA1B702FF713417FBDE9A08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA2B702FF713005FCE89F48.text	03E787B8FFA2B702FF713005FCE89F48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis jordani (Parker 1936)	<div><p>Leptopelis cf. jordani (Parker, 1936)</p><p>Figure 4K</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P4.105; GenBank: PQ455858. • Mayombe NP, Mbongo Zimune; –4.7442, 12.6914; 306 m a.s.l.; FKH 1245; GenBank: PQ455856. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; –4.6403, 12.935; 269 m a.s.l.; FKH 1248; GenBank: PQ455857.</p><p>Identification. A medium-sized and robust Leptopelis (max. SVL = 62 mm). The species was originally described from Congulo in the central Angolan escarpment. Genetically, L. jordani belongs to the L. bocagii complex (Baptista 2024), from which it differs by only 1.3% 16S p -distance. It can be distinguished from the latter species by having arboreal habits (versus ground-dwelling in L. bocagii sensu stricto), well-developed discs on fingers and toes (versus absent in L. bocagii), and usually shows characteristic white leg and arm margins and spots in knees and elbows (versus absence of white markings on L. bocagii). Therefore, we ascribe specimens from Cabinda Province to L. cf. jordani based on the similar morphology to topotypic material (presence of discs and white markings) and behaviour (arboreal). However, the low genetic divergence on the 16S gene makes it impossible to resolve the taxonomy of this complex until other taxonomic tools can be applied.</p><p>Biology and distribution. The real distribution of this species is still poorly understood, but specimens ascribed to this forest-obligate, arboreal form have been recorded along the western escarpment of Angola (Baptista 2024) and now also in Cabinda Province. The specimens from Cabinda Province were found in Lower and Upper Mayombe, on leaves more than 2 m high in primary or secondary forest.</p></div>	https://treatment.plazi.org/id/03E787B8FFA2B702FF713005FCE89F48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA2B702FF713545FD7C9D7D.text	03E787B8FFA2B702FF713545FD7C9D7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis millsoni (Boulenger 1895)	<div><p>Leptopelis millsoni (Boulenger, 1895)</p><p>Figure 4L</p><p>Material. ANGOLA – Cabinda Province • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; P2.111; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3667&amp;materialsCitation.latitude=-5.1208" title="Search Plazi for locations around (long 12.3667/lat -5.1208)">GenBank</a>: PQ455859. • between Dinge and Chindende; -5.1208, 12.3667; 15 m a.s.l.; FKH 0174, EI 808; GenBank: MZ408748 –PQ455861.</p><p>Identification. A large brown Leptopelis (SVL = 49–87 mm) with irregular darker transverse bands on the dorsum. Material from Cabinda Province was included in the revision of Jaynes et al. (2022), where they report sub-structuring within the species either north and south of the Congo River. As expected, material from Cabinda Province clusters with material from Gabon, Cameroon and Nigeria (Jaynes et al. 2022).</p><p>Biology and distribution. An arboreal species widely distributed in the Gulf of Guinea (excluding the islands) and the Congo Basin south to northwestern Angola (Jaynes et al. 2022). The species is associated with large waterbodies in the lowland rainforest and secondary forests. In Cabinda Province it was collected inside gallery forest along slow-moving streams in the Littoral Peneplain.</p></div>	https://treatment.plazi.org/id/03E787B8FFA2B702FF713545FD7C9D7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA3B703FF7131B0FE50983D.text	03E787B8FFA3B703FF7131B0FE50983D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis ocellatus (Mocquard 1902)	<div><p>Leptopelis ocellatus (Mocquard, 1902)</p><p>Figure 4M</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4597&amp;materialsCitation.latitude=-4.5981" title="Search Plazi for locations around (long 12.4597/lat -4.5981)">Dinge</a> and Chindende; –5.1208, 12.3667; 15 m a.s.l.; FKH 0170; GenBank: PQ455863. • Mayombe NP, Sanga Wanda; -4.6253, 12.4583; 221 m a.s.l.; FKH 1229, P3.204, P3.517; GenBank: PQ455864–65, PQ455862. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.090; GenBank: PQ455866 .</p><p>Identification. A medium-sized Leptopelis (SVL = 46–58 mm) that can be distinguished from other Leptopelis occurring in Cabinda Province by having large dark markings on the flanks, hidden parts of the limbs bright yellow with dark marbling, and coppercoloured eyes. The material from this study is almost identical (0.8% 16S p -distance) to material from Lekoumou, Republic of the Congo (GenBank: KY080253), but differs by ∼2.3% (16S p -distance) from material collected from Birougou, Gabon (∼ 215 km from the type locality) (GenBank: KX260264).</p><p>Biology and distribution. The species is distributed from Cameroon to Cabinda Province and western DRC (Channing and Rödel 2019). Leptopelis ocellatus can be found in a wide panoply of ecological zones from lowland forest to more disturbed areas. In Cabinda Province, this species has been found in gallery, primary and secondary forests, in the Littoral Peneplain and Lower Mayombe, in sympatry with L. aubryi, L. cf. aubryioides, L. boulengeri and L. cf. calcaratus .</p></div>	https://treatment.plazi.org/id/03E787B8FFA3B703FF7131B0FE50983D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA3B703FF713258FBE49E63.text	03E787B8FFA3B703FF713258FBE49E63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptopelis rufus Reichenow 1874	<div><p>Leptopelis rufus Reichenow, 1874</p><p>Figure 4N</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1049, FKH 1053; GenBank: PQ455867–68. • Mayombe NP, Vaku; – 4.6323, 12.8937; 319 m a.s.l.; FKH 0931−32; GenBank: PQ455869–70 .</p><p>Identification. A large brown Leptopelis (SVL = 55–87 mm) sister to L. millsoni and L. macrotis, from which it can be difficult to differentiate morphologically other than by having a comparatively smaller tympanum (Jaynes et al. 2022). Material from Cabinda Province is identical (16S p -distance) to material from south of Ogooué River, Gabon (GenBank: MZ408848).</p><p>Biology and distribution. An arboreal Leptopelis widely distributed in the Gulf of Guinea lowlands forests, from southern Nigeria to Cabinda Province, including Bioko Island (Channing and Rödel 2019). The species is associated with primary moist forest. The specimens were collected from leaves near fast-flowing forest streams at Upper Mayombe between 300–400 m a.s.l. and represent the first country records (Baptista 2024).</p></div>	https://treatment.plazi.org/id/03E787B8FFA3B703FF713258FBE49E63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF81B721FF713060FEBE99D5.text	03E787B8FF81B721FF713060FEBE99D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Limaformosa savorgnani (Mocquard 1887)	<div><p>Limaformosa savorgnani (Mocquard, 1887)</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.498538&amp;materialsCitation.latitude=-5.306833" title="Search Plazi for locations around (long 12.498538/lat -5.306833)">Tando Zinze</a>; –5.306833, 12.498538; 32 m a.s.l; photographic record; roadkill .</p><p>Identification. Limaformosa savorgnani can be distinguished from congeners by having two rows of strongly developed secondary keels on the vertebral scales. The species is reported from neighbouring areas in the Republic of the Congo and DRC (Broadley et al. 2018).</p><p>Biology and distribution. Widely distributed in Central Africa from Cameroon eastwards to Ethiopia and southwards to Cabinda Province. The specimen from Cabinda Province was found in the Littoral Peneplain and represents the first confirmed record of the species in Angola.</p></div>	https://treatment.plazi.org/id/03E787B8FF81B721FF713060FEBE99D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9CB73CFF7131B0FB579940.text	03E787B8FF9CB73CFF7131B0FB579940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lygodactylus lobeke Roll, Vaz Pinto and Lobon-Rovira 2024	<div><p>Lygodactylus lobeke Röll, Vaz Pinto and Lobón-Rovira, 2024</p><p>Figure 8J</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1259; GenBank: PQ456005 .</p><p>Identification. This individual is part of the type series of a recently described taxon within the L. scheffleri group, L. lobeke . It differs by 3.5% (16S p -distance) from the only other known specimen from Cameroon (GenBank: GU593484), which is referred to as Lygodactylus sp. B (Röll et al. 2010) and Lygodactylus sp. 6 in Gippner et al. (2021). A recent phylogenetic and taxonomic revision of this group have considered this material from Cabinda and Cameroon to be conspecific and described it as a new species (Röll et al., in press).</p><p>Biology and distribution. The collected specimen was found in primary rainforest in Lower Mayombe. It was found during the day among leaf litter, while the terrain was being prepared for a camp site. Presumably, it may have fallen from the tree canopy above.</p></div>	https://treatment.plazi.org/id/03E787B8FF9CB73CFF7131B0FB579940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF81B721FF7133C0FE309FDD.text	03E787B8FF81B721FF7133C0FE309FDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mehelya poensis (Smith 1849)	<div><p>Mehelya poensis (Smith, 1849)</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3832&amp;materialsCitation.latitude=-5.473" title="Search Plazi for locations around (long 12.3832/lat -5.473)">Nganzi</a>; –5.4730, 12.3832; 129 m a.s.l.; EI 813 (roadkill); GenBank: PQ456006 .</p><p>Identification. A file snake with weak to smoothly developed secondary keels on the dorsal scales, 15 scale rows across the midbody and&gt;84 subcaudal scales (Chippaux and Jackson 2019). The Cabinda Province material is identical (16S p -distance) to material from DRC (GenBank: MF680168) and Gabon (GenBank: AY611863) and differs very little across its distribution range (&lt;1% 16S p -distance) (Trape et al. 2018).</p><p>Biology and distribution. This species is widely distributed from Sierra Leone to Uganda in the east and Angola in the south (Chippaux and Jackson 2019), being frequently found in forest patches (Spawls et al. 2018). In Angola, the species in known to occur in the northern half of the country (Branch 2018). The specimen from Cabinda Province was found killed on the road in the Littoral Peneplain, and additional information about its biology cannot be provided.</p></div>	https://treatment.plazi.org/id/03E787B8FF81B721FF7133C0FE309FDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF80B720FF7132CFFBEE9E45.text	03E787B8FF80B720FF7132CFFBEE9E45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Naja melanoleuca (Hallowell 1857)	<div><p>Naja melanoleuca (Hallowell, 1857)</p><p>Figure 8V</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.332535&amp;materialsCitation.latitude=-5.107126" title="Search Plazi for locations around (long 12.332535/lat -5.107126)">Nganzi</a>; –5.107126, 12.332535; 79 m a.s.l.; Photographic record .</p><p>Identification. Dorsum black, with 1–3 semi-divided yellow crossbands on the neck. Venter yellow with 4–6 black bands on the first half of the body and thereafter uniform black until the tail tip (Wüster et al. 2018).</p><p>Biology and distribution. A widely distributed species in the Congo Basin, extending northwards to Nigeria and southwards to Angola (Wüster et al. 2018). In Angola, the species is present along the escarpment and in the northern half of the country (Branch 2018).</p></div>	https://treatment.plazi.org/id/03E787B8FF80B720FF7132CFFBEE9E45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA4B704FF7135BCFC109DA4.text	03E787B8FFA4B704FF7135BCFC109DA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nectophryne afra Buchholz and Peters 1875	<div><p>Nectophryne afra Buchholz and Peters, 1875</p><p>Figure 4Q</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0948 (froglet); GenBank: PQ455871 .</p><p>Identification. A small toad (SVL = 19–25 mm) with slender body and webbed fingers and toes. Snout short and pointed (Channing and Rödel 2019). Two distinct clades have been recognised within N. afra, which might represent two valid taxa (Liedtke et al. 2021). The specimen reported here differs by ∼1% (16S p -distance) from material collected from Nyanga, Gabon (GenBank: MT724587), which clusters with other material from Republic of the Congo and southern Cameroon (Liedtke et al. 2021).</p><p>Biology and distribution. Nectophryne afra is an arboreal toad widely distributed in primary moist forest of the Gulf of Guinea and Congo Basin, including Bioko Island (Channing and Rödel 2019; Sanchez-Vialas et al. 2020). The material reported here represents a southern range extension and the first country record (Baptista 2024). The specimen was a metamorph found perched on a leaf ∼ 1.5 m high at Upper Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FFA4B704FF7135BCFC109DA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA5B705FF7131B0FC24983D.text	03E787B8FFA5B705FF7131B0FC24983D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nectophryne batesii Boulenger 1913	<div><p>Nectophryne aff. batesii</p><p>Figure 4R</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; FKH 0960, P2.081; GenBank: PQ455872−73 • Mayombe NP, Sanga Wanda; – 4.6253, 12.4583; 221 m a.s.l.; FKH 1265; GenBank: PQ455874 • Mayombe NP, Mbundu; – 4.6875, 12.4981; 127 m a.s.l.; FKH 1271; GenBank: PQ455875. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.109; GenBank: PQ455876.</p><p>Identification. A small toad (SVL = 23–25 mm) with compact body, webbed fingers and toes, and short, rounded and truncated snout (Channing and Rödel 2019). Liedtke et al. (2021) recognised three different clades within N. batesii that might justify specific status. The material from Cabinda Province represents an additional distinctive clade which differs by a minimum of ∼3.8% (16S p -distance) when compared with material of the southern clade from Liedtke et al. (2021), which includes material from Ivindo, Gabon (GenBank: KF665313) and Bioko Island (GenBank: MT724579).</p><p>Biology and distribution. An arboreal toad widely distributed in primary lowland forest of the Gulf of Guinea and Congo Basin, including Bioko Island (Channing and Rödel 2019; Sanchez-Vialas et al. 2020). The specimens reported here represent the southernmost records of the species, and taken together with the previous species, they add a new genus to the Angolan list. The collected individuals were recorded in Lower Mayombe between 100–250 m a.s.l. and found on top of tree or bush leaves between 1 and 3 m above the ground.</p></div>	https://treatment.plazi.org/id/03E787B8FFA5B705FF7131B0FC24983D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF98B738FF71321BFEA19EBA.text	03E787B8FF98B738FF71321BFEA19EBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Osteolaemus tetraspis Cope 1861	<div><p>Osteolaemus tetraspis Cope, 1861</p><p>Figure 8B</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; (P2.064) • Dinge; –4.6323, 12.8937; 319 m a.s.l.; EI 810; EI 812; GenBank: PQ456007–08 (street market) .</p><p>Identification. A small crocodile (max. SVL = ∼ 2 m) with black dorsum and yellowish patches on tail and jaw (Waitkuwait 1986), three transverse series of nuchal scales, and snout as long as wide at the base. Genetically, it differs by ∼3.6% from material collected from Ogooue Basin, Gabon (GenBank: MN885919).</p><p>Biology and distribution. Osteolaemus tetraspis is a widely distributed rainforest species in West Africa,from Senegal to Cabinda Province (Waitkuwait 1986). It is associated with ponds, swampy areas or water pools, frequently far away from main rivers or open waters (Waitkuwait 1986). The photographed specimen was found in a small rain pool under shade in the primary rainforest in Upper Mayombe. Samples from this study were obtained from specimens found in street markets in the Littoral Peneplain, being sold as bushmeat.</p><p>Squamata</p><p>Chamaeleonidae</p></div>	https://treatment.plazi.org/id/03E787B8FF98B738FF71321BFEA19EBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9CB73DFF713700FC3E99D5.text	03E787B8FF9CB73DFF713700FC3E99D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panaspis breviceps (Peters 1873)	<div><p>Panaspis breviceps (Peters, 1873)</p><p>Figure 8L</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3326&amp;materialsCitation.latitude=-5.1076" title="Search Plazi for locations around (long 12.3326/lat -5.1076)">Dinge</a> and Chindende; –5.1076, 12.3326; 54 m a.s.l.; EI 758; GenBank: PQ456012. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1046, P2.205; GenBank: PQ456009–10. • Mayombe NP, Vaku; – 4.6323, 12.8937; 319 m a.s.l.; FKH 0938; GenBank: PQ456011.</p><p>Identification. Panaspis breviceps can be distinguished from other Angolan species based on the presence of movable lower eyelids and number of midbody scales (Ceríaco et al. 2020). The specimens reported here were identified genetically, differing by &lt;3% (16S p -distance) from material collected from between Bamenda and Douala, Cameroon (GenBank: KY683582) and Ivindo, Gabon (GenBank: OL457697) from where the species was described, and by ∼3% from material collected from Tshopo, DRC (GenBank Ascension: OL457698). Specimens from Cabinda Province cluster with the 16S lineage from western, central and eastern populations of the Congo Basin recognised by Lokasola et al. (2021).</p><p>Biology and distribution. The species is widely distributed in West and Central Africa from Cameroon to west DRC (in the west) and Cabinda Province (in the south), while some historical records suggest that the species’ geographic range may extend much further south along the western Angolan escarpment but requiring further examination and molecular validation (Ceríaco et al. 2020). In Cabinda Province, the species was found along fast-flowing forest streams in the rainforest of Upper Mayombe, often seen moving swiftly among pebbles on the sides of streams or getting into the water to escape.</p></div>	https://treatment.plazi.org/id/03E787B8FF9CB73DFF713700FC3E99D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9DB73DFF7133C0FBB49F93.text	03E787B8FF9DB73DFF7133C0FBB49F93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panaspis cabindae (Bocage 1866)	<div><p>Panaspis cabindae (Bocage, 1866)</p><p>Figure 8M</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.328&amp;materialsCitation.latitude=-5.5297" title="Search Plazi for locations around (long 12.328/lat -5.5297)">Nganzi</a>; –5.5297, 12.3280; 96 m a.s.l.; EI-1135; GenBank: PQ456013 .</p><p>Identification. A small-sized Panaspis (max. SVL = 38.4 mm) with pre-ablepharine eyes. The specimen reported here represents the first molecular sample of the species from near the type locality (Chinchoxo, =Landana). It differs by 2.4–3.9% (16S p -distance) from material from other parts of Angola (GenBank: MN846689 −93) and by 4.3% from material from DRC (GenBank: KU236750 −53).</p><p>Biology and distribution. The species is distributed across central Africa southwards to the Angolan plateau and constitutes the most widely distributed Panaspis in Angola (Ceríaco et al. 2020). Usually found moving among leaf litter in savannah and woodland habitats.</p></div>	https://treatment.plazi.org/id/03E787B8FF9DB73DFF7133C0FBB49F93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFAEB730FF713295FBAF9BCD.text	03E787B8FFAEB730FF713295FBAF9BCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phrynobatrachus auritus Boulenger 1906	<div><p>Phrynobatrachus auritus Boulenger, 1906</p><p>Figures 7A–D</p><p>Material. ANGOLA – Cabinda Province • between Dinge and Chindende; –5.1208, 12.3667; 15 m a.s.l.; EI 800, FKH 0158; GenBank: PQ455877–78. • Cacata; –5.3150, 12.4844; 39 m a.s.l.; P1.297; GenBank: PQ455880. • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; FKH 0976; GenBank: PQ455879. • Mayombe NP, Chimbete; –4.6631, 12.5447; 182 m a.s.l.; FKH 1239; GenBank: PQ455881. • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1264, P3.117–18, P3.190–91; GenBank: PQ455882– 86. • Mayombe NP, Mbongo Zimune; –4.7442, 12.6914; 306 m a.s.l.; P3.148; GenBank: PQ455887. • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.155, P3.157; GenBank: PQ455888–89. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; P3.184; GenBank: PQ455890. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0919–20, FKH 0940–43; GenBank: PQ455891–96. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; –4.6403, 12.935; 269 m a.s.l.; P3.167; GenBank: PQ455897 .</p><p>Identification. A medium sized (SVL = 33–40 mm) terrestrial frog with a characteristic facial mask pattern. Specimens reported here were genetically identified and differ between 0.2–0.6% (16S p -distance) from material collected from Nyanga, Gabon (GenBank: KY080313, KY080311), thus we consider our material to be conspecific with P. auritus sensu stricto . The material recovered in this study represents the first confirmed country records of this species (Baptista 2024), although historical material ascribed to the west African species P. plicatus (Günther, 1858), from Chinchoxo, Cabinda Province (Peters 1877) was probably a misidentification of P. auritus due to its similar morphology (Marques et al. 2018).</p><p>Biology and distribution. A widely distributed species in lowland and submontane forests across the Congo Basin and Gulf of Guinea. Phrynobatrachus auritus is widely distributed across the entire study area in forest habitats from the Littoral Peneplain to Upper Mayombe. The species was often found moving among the leaf litter in degraded areas, forest edges or primary moist forest and crossing dirt roads during the day, but at night they can be found perched on leaves up to 1 m above the ground. It was one of the most common frog species in most forest sites.</p></div>	https://treatment.plazi.org/id/03E787B8FFAEB730FF713295FBAF9BCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF90B730FF7131C8FEC49F75.text	03E787B8FF90B730FF7131C8FEC49F75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phrynobatrachus parvulus (Boulenger 1905)	<div><p>Phrynobatrachus aff. parvulus (Boulenger, 1905)</p><p>Figures 7E–H</p><p>Material. ANGOLA – Cabina Province • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; FKH 0979; GenBank: PQ455898. • Tando Zinze; –5.3068, 12.4985; 32 m a.s.l.; FKH 0587–90; GenBank: PQ455899–02. • Cacata; –5.3144, 12.4878; 38 m a.s.l.; FKH 0698–99; GenBank: PQ455903–04. • Mayombe NP, Chimbete; –4.6631, 12.5447; 182 m a.s.l.; P3.135; GenBank: PQ455905. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4407&amp;materialsCitation.latitude=-4.6661" title="Search Plazi for locations around (long 12.4407/lat -4.6661)">Sanga Mongo</a>; –4.6661, 12.4407; 146 m a.s.l.; P2.065; GenBank: PQ455906. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; FKH 1233, P3.219; GenBank: PQ455907–08. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4583&amp;materialsCitation.latitude=-4.6253" title="Search Plazi for locations around (long 12.4583/lat -4.6253)">Sanga Wanda</a>; –4.6253, 12.4583; 221 m a.s.l.; FKH 1257; GenBank: PQ455909.</p><p>Identification. Phrynobatrachus parvulus is a species of small terrestrial frog within the P. mababiensis complex, which is not taxonomically resolved (Zimkus and Schick 2010). The type locality of P. mababiensis is “Tsotsoroga Pan, Mababe Flats”, in Botswana, and the type locality of P. parvulus is “Bange Ngola” = Dande Angola, Malanje Province, Angola. Our material does not seem to be conspecific with anything published to date, being closest to material from Malanje Province, Angola (∼3.5% 16S p -distance; GenBank: MK036476 –78) and Zambia (∼4.2% 16S p- distance; GenBank: MK464392 –93). Thus, we ascribed this material to Phrynobatrachus aff. parvulus until more information becomes available.</p><p>Biology and distribution. Phrynobatrachus aff. parvulus appears to be widely distributed in the Littoral Peneplain and Lower Mayombe, being absent from the Coastal Drier Belt and Upper Mayombe. The species was found among leaf litter at night or moving during the day, often hiding on the forest floor under logs or fallen leaves, sometimes close to streams in mature forest, but also in secondary and degraded forest. Several specimens were found in amplexus during the rainy season at Chimbete, Lower Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF90B730FF7131C8FEC49F75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF81B722FF713472FD8A9BCD.text	03E787B8FF81B722FF713472FD8A9BCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psammophis mossambicus Peters 1882	<div><p>Psammophis mossambicus Peters, 1882</p><p>Figure 8W</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.3677&amp;materialsCitation.latitude=-5.0835" title="Search Plazi for locations around (long 12.3677/lat -5.0835)">Nganzi</a>; –5.5297, 12.3280; 96 m a.s.l.; EI 809; GenBank: PQ456014. • between Dinge and Chindende; –5.0835, 12.3677; 92 m a.s.l.; EI 816; GenBank: PQ456015 .</p><p>Identification. Psammophis mossambicus has remarkable colour polymorphism across its distribution, contrasting with very little genetic variation (Keates 2021). The specimens reported here were genetically identical and differ by &lt;1% (16S p -distance) from other material from Angola (Keates 2021).</p><p>Biology and distribution. A diurnal, widely distributed species in sub-Saharan savannahs but absent from desert, semi-deserts and rainforest (Keates 2021). In Cabinda Province the species is only known to occur in the Littoral Peneplain and the Coastal Drier Belt, from where these new records are reported.</p></div>	https://treatment.plazi.org/id/03E787B8FF81B722FF713472FD8A9BCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF92B733FF713472FDD4996D.text	03E787B8FF92B733FF713472FDD4996D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena anchietae (Bocage 1868)	<div><p>Ptychadena anchietae (Bocage, 1868)</p><p>Figures 5D &amp; 7N</p><p>Material. ANGOLA – Cabinda Province • Fazenda Mandarim; –5.0553, 12.0610; 14 m a.s.l.; P4.070; GenBank: PQ455921. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4407&amp;materialsCitation.latitude=-4.6661" title="Search Plazi for locations around (long 12.4407/lat -4.6661)">Sanga Mongo</a>; –4.6661, 12.4407; 146 m a.s.l.; P2.090; GenBank: PQ455916. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4583&amp;materialsCitation.latitude=-4.6253" title="Search Plazi for locations around (long 12.4583/lat -4.6253)">Sanga Wanda</a>; –4.6253, 12.4583; 221 m a.s.l.; P3.107−08, P3.207−08; GenBank: PQ455917–20. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4714&amp;materialsCitation.latitude=-4.6304" title="Search Plazi for locations around (long 12.4714/lat -4.6304)">Sanga Wanda</a> entrance; –4.6304, 12.4714; 203 m a.s.l.; P3.524 (Tadpoles), P3.536 (Tadpoles); GenBank: PQ455924–25. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4981&amp;materialsCitation.latitude=-4.6875" title="Search Plazi for locations around (long 12.4981/lat -4.6875)">Mbundu</a>; –4.6875, 12.4981; 127 m a.s.l.; P4.102; GenBank: PQ455923. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4597&amp;materialsCitation.latitude=-4.5981" title="Search Plazi for locations around (long 12.4597/lat -4.5981)">Bata Lunhuca</a>; –4.5981, 12.4597; 215 m a.s.l.; P4.092; GenBank: PQ455922. • Mayombe NP, Vaku; –4.6323, 12.8937; 319 m a.s.l.; FKH 0934; GenBank: PQ455926.</p><p>Identification. A species complex of medium-sized grass frogs (SVL = 51−62 mm) with flattened dorsal skin folds that are usually indistinct (Channing and Rödel 2019). Dorsal colouration can vary from very light grey to deep reddish-brown, with vertebral stripe absent. The material from Cabinda Province is almost identical to unpublished topotypic material from Benguela Province, Angola. Therefore, we consider this material to be conspecific with Ptychadena anchietae sensu stricto, despite the high potential for cryptic diversity reported by Hübler (2016).</p><p>Biology and distribution. The Ptychadena anchietae complex is widely distributed in savannahs south of the Congo Basin and in East Africa (Hübler 2016). In mainland Angola, the species is often present in forest habitats, reported along the western escarpment and extending inland in northern moister habitats (Baptista 2024). In Cabinda Province, the species is probably common and present in all ecological regions, from the Coastal Drier Belt to the Upper Mayombe. Our specimens were usually found in clearings, forest edges or disturbed areas.</p></div>	https://treatment.plazi.org/id/03E787B8FF92B733FF713472FDD4996D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF93B733FF7133A8FE2F9E18.text	03E787B8FF93B733FF7133A8FE2F9E18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena mascareniensis OTU	<div><p>Ptychadena mascareniensis OTU 6 (Dumeril and Bibron, 1841)</p><p>Figure 7O</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1031; GenBank: PQ455936. • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.935&amp;materialsCitation.latitude=-4.6403" title="Search Plazi for locations around (long 12.935/lat -4.6403)">Lombe River</a>, Vaku; –4.6403, 12.935; 269 m a.s.l.; FKH 1251; GenBank: PQ455937 .</p><p>Identification. A species complex of medium-sized (SVL = 46–58 mm) grass frogs that includes several new candidate species (Zimkus et al. 2017). Usually with brown to grey dorsal colouration, a green vertebral line and 4–6 continuous rows of dorsolateral and lateral skin folds (Channing and Rödel 2019). In Cabinda Province we recorded two well differentiated (∼3.4% 16S p -distance) mitochondrial lineages within this species complex (see Ptychadena sp. account). Material from Upper Mayombe is conspecific with material from Congo Basin and Gabon ascribed to OTU6 by Zimkus et al. (2017), which is identical to material from Buba, Lake Albert, DRC (GenBank: KX836481) and Kampala, Uganda (GenBank: DQ525931).</p><p>Biology and distribution. The species complex is widely distributed in sub-Saharan savannah and rainforest, including Madagascar. Adapted to a wide panoply of biomes and ecoregions, this species complex has the largest geographic distribution of any amphibian species in Africa. The material mentioned here was collected in clearings on the edge of primary rainforest.</p></div>	https://treatment.plazi.org/id/03E787B8FF93B733FF7133A8FE2F9E18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF93B734FF713475FB51998B.text	03E787B8FF93B734FF713475FB51998B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena perreti Guibe & Lamotte 1958	<div><p>Ptychadena perreti Guibé &amp; Lamotte, 1958</p><p>Figure 7P</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; FKH 1256, P3.206, P3.531; GenBank: PQ455928–30. • Mayombe NP, Bata Lunhuca; –4.5981, 12.4597; 215 m a.s.l.; P4.091; GenBank: PQ455927. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; FKH 1050, P2.215–16, P2.229; GenBank: PQ455931−34.</p><p>Identification. A medium to large grass frog (SVL = 48–64 mm) with compact body and long legs. The specimens reported here present continuous median and post-eyelid folds, and a short but continuous dorsolateral fold, with continuous external fold, and the lack of sacral ridges (Channing and Rödel 2019). Brownish dorsum with black speckles or a green vertebral line from snout to hind limbs (Figure 7P). Material from this work differs by 0.4% (16S p -distance) from material assigned to Ptychadena bibroni collected in Gabon (GenBank: AY517604), from material from Mount Kupe, Cameroon (GenBank: KX671789) ascribed to Ptychadena aff. aequiplicata by Portik et al. (2016), also differs by 0.4–0.6% (16S p -distance) from material from the Republic of the Congo (GenBank: KY080384 –85) ascribed to Ptychadena sp. B by Deichmann et al. (2017). However, Nečas et al. (2024) demonstrated that all those GenBank sequences have been the result of a long-standing confusion within this group, and demonstrate that they all represent indeed material of P. perreti . Thus, we consider Angolan material as conspecific to that from Gabon, Cameroon and Republic of Congo, representing the southwestern most record of the species.</p><p>Biology and distribution. This species is a forest-obligate distributed from Cameroon to Cabinda Province (Channing and Rödel 2019). The specimens collected were found along trails in the forest and near forest streams. All the material from Cabinda Province was collected from Mayombe NP between 200−400 m a.s.l. and represents a new country record.</p></div>	https://treatment.plazi.org/id/03E787B8FF93B734FF713475FB51998B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF94B735FF713567FD1B9A08.text	03E787B8FF94B735FF713567FD1B9A08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptychadena taenioscelis Laurent 1954	<div><p>Ptychadena taenioscelis Laurent, 1954</p><p>Figure 7Q</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.5073&amp;materialsCitation.latitude=-5.3241" title="Search Plazi for locations around (long 12.5073/lat -5.3241)">Tando Zinze</a>; –5.3241, 12.5073; 29 m a.s.l.; P1.141; GenBank: PQ455938 .</p><p>Identification. A small (SVL = 35–40 mm) grass frog with a compact body and long legs. Dorsum brownish with distinctive and continuous skin folds from the posterior part of the eye to lower back. Red to yellow thin vertebral stripe surrounded by numerous black spots (Channing and Rödel 2019). The specimen reported here is identical (16S p -distance) to material from Nyanga, Gabon (GenBank: KY080398) and almost identical (0.2% 16S p -distance) to material from Likouala, Republic of the Congo (GenBank: KY080399). However, the species was originally described from Tanganyika Province, DRC, from where genetic material is not available. Therefore, due to the high diversification of this group we recommend future efforts to obtain topotypic material from Lukuka, Tanganyika Province, DRC, to compare against this available material from west Central Africa. Nevertheless, a preliminary 16S ML tree clusters in the Cabinda Province material with material from Gabon, which has been regarded as nominotypical P. taenioscelis (Conradie 2024) .</p><p>Biology and distribution. Ptychadena taenioscelis is widely distributed in humid savannahs, woodlands and secondary forest in the southern rim of the Congo Basin (Channing and Rödel 2019). In Cabinda Province, the species was only found in the Littoral Peneplain near plantations and degraded secondary forest.</p></div>	https://treatment.plazi.org/id/03E787B8FF94B735FF713567FD1B9A08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF82B722FF713001FE2B9867.text	03E787B8FF82B722FF713001FE2B9867.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Python sebae (Gmelin 1788)	<div><p>Python sebae (Gmelin, 1788)</p><p>Figure 8X</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.498538&amp;materialsCitation.latitude=-5.306833" title="Search Plazi for locations around (long 12.498538/lat -5.306833)">Fazenda Mandarim</a>; –5.0401, 12.0508; 14 m a.s.l.; P4.073 (bushmeat); GenBank: PQ456016. • Tando Zinze; –5.306833, 12.498538; 32 m a.s.l.; photographic record (street market) .</p><p>Identification. A large rock python (max. SVL = 6.5 m) that can be distinguished from its close relative P. natalensis based on the colour pattern on the side of the head and larger scales on the head (see Chippaux and Jackson 2019). The specimens reported here are genetically identical (&lt;0.5% 16S p -distance) to unpublished material (K. Tolley unpublished data) from northern Angola (e.g., Luquembo and Cuanza River).</p><p>Biology and distribution. Widely distributed in West and Central Africa from Senegal eastwards to Tanzania and southwards to Angola (Chippaux and Jackson 2019). The species is frequently found at night and in a wide variety of habitats. The species is frequently used as bushmeat and sold in street markets across Africa. We recorded several pythons in street markets, mostly in the Littoral Peneplain and drier coastal belt, but also in Mayombe NP.</p></div>	https://treatment.plazi.org/id/03E787B8FF82B722FF713001FE2B9867	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA6B706FF7131B0FE449878.text	03E787B8FFA6B706FF7131B0FE449878.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys pusilla (Mertens 1937)	<div><p>Sclerophrys pusilla (Mertens, 1937)</p><p>Figure 4T</p><p>Material. ANGOLA – Cabinda Province • Fazenda Mandarim; –5.0401, 12.0508; 14 m a.s.l.; P4.071; GenBank: PQ455949. • Mayombe NP, Sanga Mongo; –4.6661, 12.4406; 146 m a.s.l.; P2.089, FKH 0966; GenBank: PQ455943−44 • Mayombe NP, Mbundu; – 4.6875, 12.4981; 127 m a.s.l.; P3.129, P3.221, P3.546, P4.101; GenBank: PQ455946−48, PQ455950 • Mayombe NP, Miconge; -4.4860, 12.8780; 377 m a.s.l.; P2.210; GenBank: PQ455945.</p><p>Identification. A medium-sized toad (SVL = 65–80 mm) with flattened parotoid glands and great variation of morphological patterns. Specimens from Cabinda Province are genetically identical to other material from Angola (e.g., Iona NP [GenBank: ON006588] and Caccuchi River [GenBank: KT862862]) and differ by &lt;1% from other material from Republic of the Congo (GenBank: KY079995), DRC (GenBank: KT862859) and Gabon (GenBank: KX260256).</p><p>Biology and distribution. A terrestrial toad with the widest distribution in the genus, present across most sub-Saharan savannahs and woodlands, but also present in forest, from Cameroon to Ethiopia in the east, southwards to Mozambique and eastern South Africa, and westward to Angola and northern Namibia. Although the species is widely distributed in Angola (Marques et al. 2018), this represents the first records from Cabinda Province (Baptista 2024). This species is likely common throughout the province, from the rainforest edge in both Lower and Upper Mayombe to the more arid savannahs of the Coastal Drier Belt.</p></div>	https://treatment.plazi.org/id/03E787B8FFA6B706FF7131B0FE449878	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA6B706FF713295FD769EE8.text	03E787B8FFA6B706FF713295FD769EE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys regularis (Reuss 1833)	<div><p>Sclerophrys regularis (Reuss, 1833)</p><p>Figure 4U</p><p>Material. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4985&amp;materialsCitation.latitude=-5.3068" title="Search Plazi for locations around (long 12.4985/lat -5.3068)">Tando Zinze</a>; –5.3068, 12.4985; 32 m a.s.l.; P1.142; GenBank: PQ455951 .</p><p>Identification. A large toad (SVL = 91–130 mm) with enlarged parotoid glands and large warts scattered across the entire body. The species has low genetic divergence across its entire range. The specimen reported here is genetically identical to material from Pool, Republic of the Congo (GenBank: KY080012), Sambolabo, Cameroon (GenBank: KF665304), Kampala, Uganda ( GenBank: AF220890) and Lake Baringo, Kenya (GenBank: AF220889), and almost identical to material from the type locality in Egypt (GenBank: KF665201).</p><p>Biology and distribution. This toad is known to occur from Senegal eastwards to Ethiopia and northern Tanzania, northwards across the Nile River, and southwards to northern Angola (Channing and Rödel 2019). The species is widely distributed in sub-Saharan savannahs and commonly found in urban and highly impacted areas (e.g. farms and plantations). Only one specimen was found during this study, in degraded forest near farmland in the Littoral Peneplain.</p></div>	https://treatment.plazi.org/id/03E787B8FFA6B706FF713295FD769EE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA6B707FF713725FB579905.text	03E787B8FFA6B707FF713725FB579905.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sclerophrys tuberosa (Gunther 1858)	<div><p>Sclerophrys tuberosa (Günther, 1858)</p><p>Figure 4V</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.4583&amp;materialsCitation.latitude=-4.6253" title="Search Plazi for locations around (long 12.4583/lat -4.6253)">Sanga Wanda</a>; –4.6253, 12.4583; 221 m a.s.l.; FKH 1258; GenBank: PQ455952. • Mayombe NP, Sanga Wanda; –4.6270, 12.4527; 215 m a.s.l.; P3.509; GenBank: PQ455953.</p><p>Identification. A medium-sized toad (SVL = 48–74 mm) with compact body and enlarged globose parotoid glands, and body covered with long spiny warts (Channing and Rödel 2019). The material from the study area differs by &lt;1% (16S p -distance) from material from between Malapa and Mbimb, Cameroon (GenBank: KF665246), and ∼2.5% from material from Lekoumou, Republic of the Congo (GenBank: KY080024).</p><p>Biology and distribution. The species is known to occur in lowland and mid-elevation primary forest to the north of the Congo River, from southeast Nigeria to western DRC (Channing and Rödel 2019). The specimens reported here are the first records for Cabinda Province and Angola (Baptista 2024) and represent the southernmost records for the species. Both specimens collected were found foraging on the ground at the edge of a dirt road surrounded by primary rainforest in Lower Mayombe, at ∼ 200 m a.s.l.</p></div>	https://treatment.plazi.org/id/03E787B8FFA6B707FF713725FB579905	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FFA3B704FF7134AFFEBF9AD8.text	03E787B8FFA3B704FF7134AFFEBF9AD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scotobleps gabonicus Boulenger 1900	<div><p>Scotobleps gabonicus Boulenger, 1900</p><p>Figure 4O</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Chimbete; –4.6631, 12.5447; 182 m a.s.l.; P3.138, FKH 1237–38; GenBank: PQ455954–56 .</p><p>Identification. Scotobleps gabonicus is a relatively large species (SVL = 52–70 mm) and the sole representative of a monotypic genus of Arthroleptidae . It can be distinguished from other Arthroleptidae by having large and small warts on the dorsum, head, and limbs, and webbed feet (Channing and Rödel 2019). Specimens from the study area are almost identical (0.3% 16S p -distance) to material from Ivindo, Gabon (GenBank: MF511944) and identical to material for Lekoumou, Republic of the Congo (GenBank: KY080407). However, they differ by ∼5% (16S p -distance) from material from Cameroon and Nigeria (GenBank: KX671793, MT630558), which may suggest cryptic diversification in this group that warrants further investigation.</p><p>Biology and distribution. The species is widely distributed in the lowland forests of the Gulf of Guinea (Channing and Rödel 2019). This material represents the first records from Cabinda Province, and adds a new amphibian genus to the country list (Baptista 2024). All the specimens were found active at night near water or moving in open areas near a slow-moving stream in Lower Mayombe. Individuals tried to hide under leaf litter when they were approached.</p></div>	https://treatment.plazi.org/id/03E787B8FFA3B704FF7134AFFEBF9AD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9DB73EFF7135FFFE939A08.text	03E787B8FF9DB73EFF7135FFFE939A08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis albilabris (Hallowell 1857)	<div><p>Trachylepis albilabris (Hallowell, 1857)</p><p>Figure 8N–O</p><p>Material. ANGOLA – Cabinda Province • between <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.878&amp;materialsCitation.latitude=-4.486" title="Search Plazi for locations around (long 12.878/lat -4.486)">Dinge</a> and Chindende; –5.0922, 12.3092; 31 m a.s.l.; EI 776; GenBank: PQ456020. • Nganzi; –5.5025, 12.3679; 107 m a.s.l.; Photographic record. • Mayombe NP, Sanga Mongo; –4.6661, 12.4407; 146 m a.s.l.; P2.066; GenBank: PQ456017. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.130; GenBank: PQ456018. • Mayombe NP, Miconge; –4.4860, 12.8780; 377 m a.s.l.; P3.171; GenBank: PQ456019 .</p><p>Identification. A medium-sized skink (max. SVL = 75.8 mm). Dorsum olive-brown with dark flecks, a dark brown to black lateral band running from the snout to the tail, bordered by a white stripe below. Supra- and infralabials characteristically white with a yellowish to orange patch near the forelimb insertion (Figure 8O). In reproductively active males, the white colouration on the labials is replaced by a yellowish colouration (Figure 8N). Specimens from Cabinda Province are genetically identical to material from Lekoumou, Republic of the Congo (GenBank: KY683566).</p><p>Biology and distribution. An arboreal skink widely distributed in West Africa, from Guinea Bissau to Angola (Allen et al. 2019). In Cabinda Province, the species was found in all four ecological zones considered, moving actively during the day on tree trunks and branches.</p></div>	https://treatment.plazi.org/id/03E787B8FF9DB73EFF7135FFFE939A08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9EB73EFF713005FD36985B.text	03E787B8FF9EB73EFF713005FD36985B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis maculilabris (Gray 1845)	<div><p>Trachylepis maculilabris (Gray, 1845)</p><p>Figure 8P</p><p>Material. ANGOLA – Cabinda Province • Caio Cacongo; –5.0618, 12.2588; 43 m a.s.l.; P2.104; GenBank: PQ456021. • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; P3.220; GenBank: PQ456022.</p><p>Identification. A medium-sized skink (max. SVL = 95 mm) with robust limbs and body. Dorsum olive-grey to dark brown, with scattered white to yellowish speckles on the flanks and limbs. Venter, throat and infralabials cream to yellow, uniformly coloured (Ceríaco et al. 2024). Specimens from Cabinda Province differ by ∼1% (16S p -distance) from specimens from Nyanga, Gabon (GenBank: MK792018) and Mambasa, DRC (GenBank: MK496123). However, they differ by ∼3.9% from material south of the Congo River at Cuanza-Sul Province, Angola (GenBank: MK792017).</p><p>Biology and distribution. Trachylepis maculilabris is the arboreal skink with the widest distribution in Africa (Allen et al. 2019). In Cabinda Province, the species was recorded from the coast to Lower Mayombe, being found in clearings, patches of secondary or degraded forest, and in farmland and on buildings.</p></div>	https://treatment.plazi.org/id/03E787B8FF9EB73EFF713005FD36985B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9EB73EFF7132B7FC9E9E63.text	03E787B8FF9EB73EFF7132B7FC9E9E63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trachylepis polytropis (Boulenger 1903)	<div><p>Trachylepis polytropis (Boulenger, 1903)</p><p>Figure 8Q</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Mbundu; –4.6875, 12.4981; 127 m a.s.l.; FKH 1270; GenBank: PQ456023 .</p><p>Identification. A medium-sized skink (max. SVL = 96 mm). Dorsum brown with dark wavy transverse stripes and two black lateral bands running from the snout to the tail, bordered ventro-laterally by a white to yellow stripe. Venter bluish, becoming yellow on the throat. The specimen reported here differs by 1.8% (16S p -distance) from specimens collected near the type locality at Monts Doudou, Gabon (GenBank: KY683558).</p><p>Biology and distribution. The species is typically associated with the great Central African forest block, where the species can be found on the leaf litter, dead trees and roots on soil and dead tree trunks (Chirio et al. 2008). This record represents the first country record for the species. The collected specimen was found active on a rotten trunk of a fallen tree on the forest floor, in Lower Mayombe.</p></div>	https://treatment.plazi.org/id/03E787B8FF9EB73EFF7132B7FC9E9E63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF9CB73CFF713387FBB79F2B.text	03E787B8FF9CB73CFF713387FBB79F2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Varanus niloticus (Linnaeus 1758)	<div><p>Varanus niloticus (Linnaeus, 1758)</p><p>Material examined. ANGOLA – Cabinda Province • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.2371&amp;materialsCitation.latitude=-5.3273" title="Search Plazi for locations around (long 12.2371/lat -5.3273)">Susso</a>; –5.3273, 12.2371; 134 m a.s.l.; roadkill .</p><p>Identification. This is the largest species of monitor lizard in Africa (SVL = 600–800 mm) (Wilms et al., 2021). It has a stout body with powerful limbs, strong claws and an elongated snout. The tongue is dark, almost black. Tail long, laterally compressed, with a low dorsal crest. Limbs spotted, belly and throat paler than dorsum with black bars. Juvenile colouration is black and bright yellow.</p><p>Biology and distribution. This species is well adapted to living in deep river valleys in all sub-Saharan countries from the Nile River in Egypt throughout the network of rivers across most of Africa, reaching southwest to Angola and Namibia (Wilms et al. 2021).</p></div>	https://treatment.plazi.org/id/03E787B8FF9CB73CFF713387FBB79F2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF91B732FF7137BDFE2C99A8.text	03E787B8FF91B732FF7137BDFE2C99A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopus (Silurana) mellotropicalis Evans, Carter, Greenbaum, Gvozdik, Kelley, McLaughlin, Pauwels, Portik, Stanley, Tinsley, Tobias and Blackburn 2015	<div><p>Xenopus (Silurana) mellotropicalis Evans, Carter, Greenbaum, Gvoždík, Kelley, McLaughlin, Pauwels, Portik, Stanley, Tinsley, Tobias and Blackburn, 2015</p><p>Figure 7L</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Quissoki; –4.6028, 12.8736; 336 m a.s.l.; P3.176; GenBank: PQ455960 .</p><p>Identification. A medium-sized (SVL = 51–58 mm) flattened frog with four claws on the feet and smooth skin on the head and dorsum, grey belly with pale yellow markings (Channing and Rödel 2019). The specimens reported here differ by 1.3% (16S p -distance) from the type series from Estuaire, Gabon (GenBank: KT728050) and by 0.8% (16S p -distance) from unpublished sequences from Loufika, Republic of the Congo (Greg Jongsma unpublished data). The specimens from Cabinda Province exhibit an extension of the interocular cream-yellow line reported by Evans et al. (2015), that extends backwards to the insertion of the hind limbs.</p><p>Biology and distribution. Xenopus mellotropicalis is distributed in primary moist forest from Cameroon to southeastern DRC and Cabinda Province (Baptista 2024). The specimen reported here represents the first record of the species for Angola (Baptista 2024). The specimen was found in Upper Mayombe, in a pond next to a large river in syntopy with Hoplobatrachus occipitalis .</p></div>	https://treatment.plazi.org/id/03E787B8FF91B732FF7137BDFE2C99A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF91B731FF71334DFBA29FB0.text	03E787B8FF91B731FF71334DFBA29FB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopus (Xenopus) allofraseri Evans, Carter, Greenbaum, Gvozdik, Kelley, McLaughlin, Pauwels, Portik, Stanley, Tinsley, Tobias and Blackburn 2015	<div><p>Xenopus (Xenopus) allofraseri Evans, Carter, Greenbaum, Gvoždík, Kelley, McLaughlin, Pauwels, Portik, Stanley, Tinsley, Tobias and Blackburn, 2015</p><p>Figure 7J</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda; –4.6253, 12.4583; 221 m a.s.l.; P3.121, FKH 1231; GenBank: PQ455958–59 .</p><p>Identification. A medium-sized (SVL = 38–48 mm) flattened frog with four claws on the feet and smooth skin on the head and dorsum. As with several other members of the group, this species can only be distinguished based on molecular data, calls and subtle morphological features (Evans et al. 2015). Material from Cabinda Province differs by 0.6% (16S p -distance) from the type material from Bioko, Equatorial Guinea (GenBank: KT728120 –28) and is identical to material from Bas-Congo Province, DRC (GenBank: KT728112) ascribed to this species.</p><p>Biology and distribution. An aquatic species widely distributed in forested areas in the Gulf of Guinea from Cameroon to Cabinda Province (Baptista2024). These records represent the first records of the species in Angola (Baptista 2024), and the southernmost for the species. The specimens reported here were collected at night in a small pond on a dirt road after a rainstorm, in primary rainforest in Lower Mayombe. The species was found in sympatry with X. andrei .</p></div>	https://treatment.plazi.org/id/03E787B8FF91B731FF71334DFBA29FB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
03E787B8FF91B731FF7135DDFEB59D50.text	03E787B8FF91B731FF7135DDFEB59D50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xenopus (Xenopus) andrei (Loumont 1983)	<div><p>Xenopus (Xenopus) andrei (Loumont, 1983)</p><p>Figure 7K</p><p>Material. ANGOLA – Cabinda Province • Mayombe NP, Sanga Wanda entrance; –4.6304, 12.4714; 203 m a.s.l.; P3.532–33; GenBank: PQ455960–61 .</p><p>Identification. A medium-sized (SVL = 37–53 mm) flattened frog, with four claws on the feet and smooth skin on the head and dorsum. Specimens are genetically identical (16S p -distance) to material from southern Londgi, Cameroon (GenBank: NC _044878).</p><p>Biology and distribution. Widely distributed from southern Cameroon to northern Angola and western DRC (Evans et al. 2015). The specimens reported here were collected at night in a small pond in a dirt road after a rainstorm. The species was found in sympatry with X. allofraseri .</p></div>	https://treatment.plazi.org/id/03E787B8FF91B731FF7135DDFEB59D50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobón-Rovira, Javier;Lobón-Rovira;Baptista, Ninda L;Clark, Tyron;Verburgt, Luke;Jongsma, Gregory Fm;Conradie, Werner;and, Luis Veríssimo;Vaz, Pedro;Pinto	Lobón-Rovira, Javier, Lobón-Rovira, Baptista, Ninda L, Clark, Tyron, Verburgt, Luke, Jongsma, Gregory Fm, Conradie, Werner, and, Luis Veríssimo, Vaz, Pedro, Pinto (2025): Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology 74 (1): 1-59, DOI: 10.1080/21564574.2024.2421007, URL: https://doi.org/10.1080/21564574.2024.2421007
