identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EE8A15D53AFFCFFCC874C9F76562A0.text	03EE8A15D53AFFCFFCC874C9F76562A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudoleskeella rupestris var. tenuis Ignatov & Ignatova 2022	<div><p>Pseudoleskeella rupestris var. tenuis Ignatov &amp; Ignatova,  var. nov.</p><p>Type: Russia, Republic of Sakha /  Yakutia,  EvenoBytantaisky District,  Orulgan Range, upper course of  Aenigan-Toolono Creek, 68°16’, 128°25’E, 900 m alt., S-faced cliffs in narrow gorge, 6 Aug 2011 Ignatov 11- 4533 (MHA 9060832). Holotype MHA, isotype MW. Figs. 2 ‘3120’, 3.</p><p>Diagnosis:  Pseudoleskeella rupestris var. tenuis differs from  var. rupestris in having smaller plants, usually shorter (to 0.5–0.6 vs. 0.6–0.9 the leaf length) and thinner costae in leaves from erect shoots, and short, double vs. single costae in many leaves from creeping shoots.</p><p>Description: Plants small to medium sized, pale green or yellowish-green, in loose mats. Stems creeping to ascending and erect, 1.0– 1.5 cm long, irregularly branched, terete-foliate; hyalodermis absent, central strand present. Leaves from erect shoots straight, 0.5–0.7× 0.2–0.4 mm, from ovate bases ± abruptly tapered into lanceolate, narrow, straight or slightly curved acumina; margins plane, subentire or finely serrulate above; costae single, thin, to 0.5(0.6) the leaf length or, in poorly developed plants, short and double; cells in mid-leaf rounded-polygonal or elongate-rhomboidal, 10–15×5–7 µm, with moderately thickened walls, smooth. Leaves from creeping shoots with wider bases and usually with short and double costae or with short single costae. Sporophytes unknown.</p><p>Other specimens examined:   Perm Province,  Bezgodov 16 Jul 2017 #194 (MHA9049390) ;  28 Jul 2017 #325, #329 (MHA9018309, MHA9018303); 15 Jul 2017 #143 (MHA 9049420);  9.VIII.2005 #106, #53 (MHA9046519, (MHA9049520) .   Altai Republic,  Ignatov 32/29, 32/30, 32/31 (MHA9046619, MHA9046337, MHA9046620). Republic of   Sakha / Yakutia, Orulgan Range,  Ignatov 11-4363, 11-4071 (MHA9046637, MHA9046634)  .</p><p>This  variety grows on rocks (limestones, aleurolites, shists), in the Urals at 180–760 m elev., in Orulgan Range, Yakutia at 670 m, in Altai at 2150–2200 m.</p><p>Differentiation:  Pseudoleskeella rupestris var. tenuis</p><p>tenuis are usually narrower ovate and the acumina are longer (however, these characters occasionally overlap). In optimally developed plants of  P. tectorum costae in the majority of leaves are forked or double, while in  P. rupestris var. tenuis costae are single in leaves from upright shoots and forked or double in leaves from creeping shoots. Similarly, this character separates  P. rupestris var. tenuis from  var. rupestris, the latter  variety always having leaves from all shoots with single, stronger and longer costae (cf. Fig. 2B vs. Fig. 4). The isotype of  P. sibirica (Fig. 4A, C–D) has all leaves with long, single costae, thus it belongs to  P. rupestris var. rupestris .</p><p>3140 Murmansk Prov. E 3152 Khabarovsk Terr.</p><p>Forked or double costae also occur  P. papillosa, but this species is very different in small size of plants and prominent, massive papillae on the dorsal side of the leaves (Fig. 5).</p><p>2) Papillose laminal cells</p><p>This is another character used in keys of  Pseudoleskeella, specifically for the identification of  P. papillosa . This species has conspicuous papillae over most cells (Fig. 5D). Papillae are occasionally observed on leaves of  P. tectorum as well, especially if the stem with undetached leaves is put in the microscope slide (Fig. 5A–C). These</p><p>50 µm</p><p>A</p><p>papillae are less numerous, and not so sharply delimited, as their ‘slopes’ are more gentle.</p><p>3) Brood branches</p><p>Most moss floras key out  P. nervosa as the only species characterized by the presence of fragile branches with minute leaves clustered in leaf axils near shoot tips. However, we found similar brood branches in many collections of  P. papillosa from Russia (Fig. 6). Unlike  P. nervosa, they are less tightly crowded, thus looking less ‘stellate’. Anyway, in some cases strict following of a key may result in erroneous identifications.</p><p>4) Bistratose leaf margins</p><p>Leaf margins are commonly unistratose in most species of  Pseudoleskeella . However, in the Caucasus and Crimea there are plants with partly bistratose leaf margins which were described as  Leskeella incrassata (Lindb. ex Broth.) Broth. This species is indistinguishable from  P. nervosa in all other characters. Similar plants also occur sporadically in southern Europe. Molecular-phylogenetic data do not support the taxonomic recognition of  L. incrassata (specimens from the Caucasus, OK3143– 3146 have bistratose margins, while specimens from Sakhalin and from Nizhhy Novgorod are unistratose).</p><p>5) Thick-walled cells</p><p>This character is somewhat difficult to demonstrate and apply.  Pseudoleskeella catenulata is usually keyed out as a species with thickest cell walls within the genus; it also has short leaf acumina, often with subobtuse apices, but morphotypes with slightly longer acumina and acute apices are not rare. These latter plants can be confused with short-leaved plants of extremely variable  P. rupestris (compare Fig. 7S and Fig. 4C, F). However,  P. catenulata always has leaves with costae to 0.5–0.6 the leaf length, while in  P. rupestris costae extend far above mid-leaf, becoming weakly delimited from adjacent cells in the leaf acumina. Cell wall thickness is also helpful, as leaf cells of  P. rupestris are usually much thinner-walled (cf. Fig. 4D vs. Fig. 9 A’). If it is difficult to estimate if cells are thick- or thin-walled, another distinguishing character can be used: the leaf cell areolation of  P. catenulata is not contrasting between juxtacostal and marginal areas (Fig. 8A), whereas in  P. rupestris rhomboidal and elliptical, thinner-walled cells in mid-leaf differ contrastingly from several rows of quadrate to transversely rectangular, thicker-walled marginal cells (Fig. 8B–C).</p><p>The clade of  P. catenulata in the molecular phylogenetic tree is well supported (Fig. 1), but it includes one specimen from Sakhalin Island differing from other specimens of this clade in morphology. It has very long acuminate leaves and only moderately thickened cell walls, in contrast to leaves with short acumina and thick-walled cells of  P. catenulata . DNA was extracted twice from this specimen in order to exclude possible errors; two identical sequences were obtained. This plant is described below as a new species.</p></div>	https://treatment.plazi.org/id/03EE8A15D53AFFCFFCC874C9F76562A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ignatova, E. A.;Kuznetsova, O. I.;Fedorova, A. V.;Ignatov, M. S.	Ignatova, E. A., Kuznetsova, O. I., Fedorova, A. V., Ignatov, M. S. (2022): The Genus Pseudoleskeella (Bryophyta) In Russia. Arctoa 31 (1): 7-16, DOI: 10.15298/arctoa.31.02, URL: https://doi.org/10.15298/arctoa.31.02
03EE8A15D53EFFC0FCC8736DF1BC6510.text	03EE8A15D53EFFC0FCC8736DF1BC6510.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudoleskeella sachalinensis Ignatov & Ignatova 2022	<div><p>Pseudoleskeella sachalinensis Ignatov &amp; Ignatova,  spec. nov.</p><p>Type: Russia, Sakhalinskaya Province, Sakhalin <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.46666&amp;materialsCitation.latitude=49.866665" title="Search Plazi for locations around (long 143.46666/lat 49.866665)">Island</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.46666&amp;materialsCitation.latitude=49.866665" title="Search Plazi for locations around (long 143.46666/lat 49.866665)">Smirnykh District</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=143.46666&amp;materialsCitation.latitude=49.866665" title="Search Plazi for locations around (long 143.46666/lat 49.866665)">Nature Reserve</a> “Vaida Mountain”, 49°52'N, 143°28'E, 450 m alt., on rocks in fir forest, 21 Aug 2006 Ignatov &amp; Teleganova 06-323 (MHA9046548). Holotype MHA, isotype MW.</p><p>Diagnosis:  Pseudoleskeella sachalinensis is similar to  P. rupestris var. rupestris in having leaves with long acumina, long single costae, and elongate-rhomboidal leaf cells, but differs in having leaves from erect stems with even longer acumina, constituting ca. 0.5–0.7 the leaf length (0.3–0.5 the leaf length in  P. rupestris var. rupestris) and longer cells in distalmost leaf portion: 4–6:1 vs. 2–4:1.</p><p>Description: Plants medium-sized, green or yellowish-green, in loose mats. Stems creeping and ascending to erect, 1.0– 1.5 cm long, hyalodermis absent, central strand present. Leaves from erect shoots straight or slightly falcate-secund, 0.7–1.0× 0.2–0.4 mm, from an ovate base ± abruptly tapered into long, narrow lanceolate, straight or slightly curved acumina constituting ca. 0.5– 0.7 the leaf length; margins plane, entire or serrulate in places; costae single, thin, to 0.6–0.8 the leaf length; cells in mid-leaf elongate-rhomboidal, 19–35×6–8 µm, towards margins shorter, rhomboidal, quadrate and transversely rectangular, firm-walled, smooth; upper cells of leaf acumina oblong, 16–35 µm long, 4–6:1. Leaves from creeping shoots slightly smaller, with wider bases, costa single. Sporophytes unknown.</p><p>Pseudoleskeella sachalinensis is currently known only from the type locality.</p><p>Differentiation:  Pseudoleskeella sachalinensis differs from all other species of the genus by the longest – ca. 0.5–0.7 the leaf length – leaf acumina, and especially long – 4–6:1 – cells in the uppermost leaf portion. Leaf acumina of  P. rupestris are usually shorter, composed of shorter cells. However, in some morphotypes of  P. rupestris leaf acumina in proportion to leaf length approach those of  P. sachalinense and are similarly narrow acuminate (Fig. 7G). In this case, the difference in cell length and width of acumina are important distinguishing characters: a number of long cells in leaf acumina is larger in  P. sachalinensis and the acumina are 2–4 cells wide for a longer distance than in  P. rupestris (cf. Fig. 7C–E vs. F).</p></div>	https://treatment.plazi.org/id/03EE8A15D53EFFC0FCC8736DF1BC6510	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ignatova, E. A.;Kuznetsova, O. I.;Fedorova, A. V.;Ignatov, M. S.	Ignatova, E. A., Kuznetsova, O. I., Fedorova, A. V., Ignatov, M. S. (2022): The Genus Pseudoleskeella (Bryophyta) In Russia. Arctoa 31 (1): 7-16, DOI: 10.15298/arctoa.31.02, URL: https://doi.org/10.15298/arctoa.31.02
03EE8A15D531FFC1FF3E7522F1326336.text	03EE8A15D531FFC1FF3E7522F1326336.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudoleskeella IN	<div><p>KEY TO IDENTIFICATION OF  PSEUDOLESKEELLA IN RUSSIA</p><p>1. All median leaf cells with prominent single papillae at the upper ends .............................. 3.  P. papillosa In European Russia  P. papillosa is known only from the Kola Peninsula and the Urals. In Asiatic Russia the species is known from scattered localities in many relatively well explored mountain areas, except Arctic.</p><p>— Leaf cells smooth or weakly prorate or only a few cells with single, small or large papillae at the upper cell ends ................................................................ 2</p><p>2. Costae single, percurrent; axillary brood branches usually present; corticolous, rarely saxicolous ........ ........................................................... 6.  P. nervosa Pseudoleskeella nervosa is a common epiphyte in the forest zone and in the forest-steppe zone of European Russia and in the Caucasus. In Asiatic Russia it occurs in southern West Siberia and in a few localities in the Russian Far East (Kamchatka; Primorsky Territory; Sakhalin Island and Kunashir Islands).</p><p>— Costae single, forked, or double, extending 0.2–0.9</p><p>the leaf length; axillary brood branches absent; saxicolous, rarely corticolous ...................................... 3</p><p>3. Costae forked or double in most leaves from both upright and creeping stems, occasionally single in some leaves ........................................................... 4</p><p>— Costae single in all leaves or single in leaves from upright shoots and forked or double in leaves from creeping shoots ..................................................... 5</p><p>4. Leaves broadly ovate or broadly ovate-triangular at the base, abruptly tapered to acumina that are 35– 50% the leaf length .......................... 1.  P. tectorum In Russia  P. tectorum is known from most well explored mountain areas and some lowlands where calcareous outcrops are more or less numerous.</p><p>— Leaves ovate or ovate-oblong at base, somewhat abruptly tapered to acumina that are 45–70% the leaf length .............................. 4.  P. rupestris var. tenuis This variety is known from scattered localities in the Urals, Altai Mountains, Yakutia, and Sakhalin Island.</p><p>5. Leaf apices acute, often blunt; acumina 0.2–0.4(– 0.5) the leaf length; leaf cells 1–1.5:1, thick-walled ....................................................... 2.  P. catenulata Pseudoleskeella catenulata occurs mainly in European Russia and the Caucasus with scattered localities in Asiatic Russia.</p><p>— Leaf apices acuminate; acumina 0.3–0.6(–0.7) the leaf length; leaf cells 1.5–3:1, moderately thick-walled .................................................................... 6</p><p>6. Leaves from creeping shoots often with double or forked costae; leaves from upright shoots with costae to 0.3–0.5(–0.7) the leaf length ......................... ....................................... 4.  P. rupestris var. tenuis</p><p>— Leaves from creeping shoots with single costae; leaves from upright shoots with costae to (0.5–)0.7– 0.9 the leaf length ................................................. 7</p><p>7. Leaf acumina 0.5–0.7 the leaf length; upper leaf cells elongate-rectangular, 4–6:1 ...... 5.  P. sachalinensis Russian Far East (Sakhalin Island).</p><p>— Leaf acumina 0.3–0.5(–0.7) the leaf length; upper leaf cells rhomboidal, irregularly polygonal, and rectangular, 2–4:1 ............ 4.  P. rupestris var. rupestris In Russia  P. rupestris var. rupestris is known from the Kola Peninsula, the Urals, the Caucasus, mountains of southern Siberia, and the southern Russian Far East. It occasionally occurs in northern Siberia in areas with calcareous bedrocks.</p></div>	https://treatment.plazi.org/id/03EE8A15D531FFC1FF3E7522F1326336	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ignatova, E. A.;Kuznetsova, O. I.;Fedorova, A. V.;Ignatov, M. S.	Ignatova, E. A., Kuznetsova, O. I., Fedorova, A. V., Ignatov, M. S. (2022): The Genus Pseudoleskeella (Bryophyta) In Russia. Arctoa 31 (1): 7-16, DOI: 10.15298/arctoa.31.02, URL: https://doi.org/10.15298/arctoa.31.02
