taxonID	type	description	language	source
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	description	(Figs. 1, 2)	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	materials_examined	Material examined. Holotype: female (CL 5.0 mm, PCL 3.6 mm), FLMNH UF 68644, Saudi Arabia, Red Sea, about 100 km off Al Haridhah, Red Sea Decade Expedition Leg 2, ROV dive CHR 0233, 17.387135, 40.827209 to 17.387619, 40.828850, depth: 104 – 248.5 m, Chimaera Dive Team, processed by F. Benzoni and F. Barreca, 28.03.2022 (CHR 0233 / OCX- 072) [GenBank Acc. No: PV 627001]. Paratype: male (CL 3.7 mm, PCL 2.5 mm), FLMNH UF 68645, Saudi Arabia, Red Sea, about 5 km east of Shibara Island, Red Sea Decade Expedition Leg 4, ROV dive CHR 0308, 25.407367, 36.775886, depth: 210.1 m, Chimaera Dive Team, processed by S. Bähr and N. Dunn, 16.06.2022 (fcn CHR 0308 / OCX- 147).	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	description	Description. Carapace. Sparsely setose, as long as broad; cervical groove distinct; dorsal surface nearly horizontal from anterior to posterior. Each side of carapace with 3 pairs of epigastric spines and 1 parahepatic spine; 3 uninterrupted ridges present on gastric region posterior to epigastric spines; mid-transverse ridge of carapace uninterrupted, preceded by cervical groove. Posterior branchial region laterally with 4 ridges (exclusive of mid-transverse ridge and posterior-most transverse ridges directly anterior to posterior margin of carapace). Dorsal surface with short uniramous and scattered long iridescent setae arising from transverse ridges. Lateral margins slightly convex, subparallel, with 6 spines: 1 st spine anterolateral, directed straight forward, followed by 1 spine on hepatic margin, 3 spines on anterior branchial margin, and 1 – 2 spines on posterior branchial margin. Rostrum moderately broad, as long as wide, about 0.4 times as long as carapace; dorsal surface nearly horizontal in lateral view, concave from side to side, with dorsal longitudinal carina, without setiferous striae; rostral spine 2.4 times longer than wide (measured at sinus between rostral and anterior lateral spines). Pterygostomian flap rugose, with sparse setae, bluntly produced anteriorly. Thoracic sternum. Sternal plastron as long as broad; lateral limits divergent posteriorly. Sternite 3 about 3.4 times as broad as long, anterior margin moderately convex. Sternite 4 about 3.9 times longer and twice as broad as preceding sternite, 1.9 times broader than long; surface with 1 anterior stria flanking midline. Following sternites smooth. Lateral parts of sternite 7 smooth. Pleon. Smooth. Somite 2 unarmed; somites 2 – 4 without transverse ridges other than anterior one, latter bearing both short and scattered long, uniramous, iridescent setae. Telson with 10 plates. Eye. Peduncle as long as broad. Cornea slightly dilated; maximum corneal diameter about 0.4 times distance between bases of anterolateral spines; eyelashes short, not reaching mid-length of cornea. Antennule. Article 1 with distal margin armed with 3 strong and 1 very small distal spines, distolateral spine longer than distodorsal one, distomesial spine much smaller than distolateral and distodorsal ones, distomesial spine minute or obsolescent; lateral margin unarmed; ventral surface unarmed. Antenna. Article 1 with distomesial small spine not reaching distal margin of article 2. Article 2 with subequal, well-developed, distolateral and distomesial spines, reaching mid-length of article 3. Article 3 with distomesial spine reaching mid-length of article 3, distolateral angle unarmed. Article 4 unarmed. Mxp 3. Ischium with small distal spine on flexor margin; extensor margin unarmed; crista dentata with about 27 small denticles, proximal-most minute. Merus subequal in length to ischium, flexor margin with 2 subequal small spines: distal spine terminal, proximal spine at mid-length; extensor margin unarmed. Carpus unarmed. P 1. Twice as long as carapace, stout, with uniramous iridescent setae along lateral and mesial margins of merus, palm and fingers; dorsal and ventral surfaces of palm and fingers with scattered short setae. Merus about 0.7 times as long as carapace, 2.5 times as long as carpus, with dorsal and lateral spines, and 1 strong distomesial spine. Carpus 0.6 times as long as palm, 0.5 times as long as broad, with 2 rows of small spines on dorsal surface, 1 prominent spine on mesial margin and 3 prominent spines along lateral margin. Palm 0.9 times as long as broad; mesial and lateral margins with strong spines continuing onto fingers, dorsal surface with small spines. Fingers distally spooned; movable finger as long as palm, its mesial margin with 3 spines. P 2 – 4. Somewhat compressed mesiolaterally, sparsely covered with uniramous iridescent setae and some plumose setae. Meri successively shorter posteriorly; P 2 merus 0.8 times as long as carapace, 3.5 times as long as broad, 1.1 times longer than P 2 propodus; P 3 merus 3.8 times as long as broad, 1.2 times length of P 3 propodus; P 4 merus 3.8 times as long as broad, 0.9 times length of P 4 propodus; dorsal margins of meri with row of proximally diminishing spines in P 2 and P 3, unarmed in P 4; dorsolateral surface unarmed in P 2 and P 3, with 1 spine on P 4; ventrolateral margins with prominent terminal spine. Carpi with 3 spines on extensor margin in P 2 and P 3, unarmed in P 4, distal-most spine largest; lateral surface with 1 spine or unarmed; flexor margins with distal spine. Propodi subequal in length on P 2 – 4, each 4.6 – 5.2 times as long as broad; extensor margin with 1 – 2 proximal spines, lateral side unarmed; flexor margin with 6 – 7 slender movable spines in P 2 – 4. Dactyli subequal in length, 0.7 – 0.8 times length of propodi, ending in incurved strong sharp spine; flexor margin with 5 – 6 obsolescent teeth, each with stout spiniform movable spine. Colour in life. Ground colour of carapace and pleon yellow orange. Carapace with several transverse bands of lavender fringed with deep purple on gastric and cardiac regions, some bands complete, others interrupted near midline; epigastric and post-rostral regions with similar, lavender or lilac-purple patches and spots. Rostrum speckled with small red dots; rostral spines with reddish tips. Pleonites 2 – 4 with some symmetrically arranged, whitish and orange spots on anterior ridge; pleonites 2 – 3 whitish with pale orange tinge, pleonites 4 – 6 bright yellow orange. Eyes brownish yellow. P 1 whitish with large pale orange areas and bright red markings on spines; chelae whitish proximally, mostly orange distally, with reddish spots on spines; fingertips whitish. P 2 – 4 with alternating whitish and pale orange transverse bands or rings, orange areas with white spots (Fig. 2).	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	etymology	Etymology. The new species is named after our dear friend and colleague, Francesca Benzoni (King Abdullah University of Science and Technology), for making all the benthic crustacean samples from OceanX expeditions available for study and supporting one of the authors’ (AA) taxonomic work on various groups of Decapoda.	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	distribution	Distribution and bathymetry. Red Sea, at depths of 104 – 248.5 m.	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	biology_ecology	Ecology. Both specimens were found as “ by-catch ” in sample baskets with coral rubble and living octocorals.	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6BFF86FF26FD7A1A64CC86.taxon	discussion	Remarks. Sadayoshia benzoniae sp. nov. belongs to a small species group within Sadayoshia, which is characterised by (1) the pleonites devoid of spines, smooth, with an anterior ridge only; (2) the shape of the sternite 3, which is more than three times as broad as long; and (3) the antennular article 1 with an unarmed lateral margin. This species group includes only two other species, namely S. actaea Macpherson & Baba, 2012 from New Caledonia (Lifou), and S. latisternata Macpherson & Baba, 2010 from French Polynesia (Austral Islands), New Caledonia (Lifou) and Mauritius (Macpherson & Baba 2010, 2012). The new species can be distinguished from both species by (1) the rostrum with a dorsal carina, which is absent in S. actaea and S. latisternata; (2) the proportionally longer rostral spine, about 2.4 times longer than wide vs. 2.1 times longer than wide in S. actaea and S. latisternata; (3) the antennular article 1 with distomesial spine minute or obsolescent vs. well developed in S. actaea and S. latisternata; (4) the gastric region of the carapace with 3 uninterrupted ridges posterior to the epigastric spines vs. 4 ridges in S. actaea and S. latisternata; and (5) the dorsal surface of the carapace with 3 pairs of epigastric spines vs. 4 pairs in S. actaea and S. latisternata, although this last character may be somewhat variable and more specimens need to be examined to confirm its validity (cf. Fig. 1; Macpherson & Baba 2010, 2012). The presence or absence of a distomesial spine on the antennal article 3 (present in the new species, absent in S. actaea and S. latisternata) represent another distinguishing character of the new species (cf. Fig. 1 D; Macpherson & Baba 2010, 2012). In addition, the colour pattern of S. benzoniae sp. nov. differs from that of S. latisternata (cf. Fig. 2; Macpherson & Baba 2010); the life colour of S. actaea remains unknown. Molecular results were fully congruent with morphological evidence. The new species was recovered as sister species of S. actaea, although with low support, in ML analyses (Fig. 3). This clade further clustered with S. latisternata. The COI genetic divergence between the new species and its closest relatives exceeds 8 %.	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6FFF89FF26FC5A1B91C94E.taxon	description	(Figs. 4, 5, 6 B)	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6FFF89FF26FC5A1B91C94E.taxon	materials_examined	Material examined. 1 male (CL 5.7 mm, PCL 4.0 mm), FLMNH UF 68646, Saudi Arabia, Red Sea, about 60 km off Al Birk, Red Sea Decade Expedition Leg 2, Neptune Sub dive NTN 0131, 17.820978, 40.709915, depth: 220 m, Neptune Dive Team, processed by F. Benzoni and F. Barreca, 30.03.2022 (NTN 0131 - Bio 2 B / OCX- 091) [GenBank Acc. No: PV 626998].	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6FFF89FF26FC5A1B91C94E.taxon	description	Description. Red Sea specimen. Carapace. About as long as wide, greatest width measured at posterior third, 1.6 times distance between anterolateral spines. Dorsal surface setose, unarmed, moderately convex. Lateral margins medially convex, with 8 spines; 1 st anterolateral, directed straight forward, situated lateral to small orbital spine, separated from latter by its basal width, overreaching orbital spine; 2 nd and 3 rd spines small, inserted on hepatic margin, 3 rd spine smaller; 4 th – 7 th spines on branchial margin, well developed; 8 th and last spine very small. Rostrum narrowly triangular, trifid, with small subterminal spine on each side, straight horizontal, 1.7 times as long as wide, 0.6 times as long as carapace length; with about half-distance between anterolateral spines, and half carapace width measured at posterior margin. Pterygostomian flap anteriorly produced into sharp spine; surface covered with small spines on anterior half, diminishing in size ventrally. Thoracic sternum. Sternal plastron slightly longer than broad; lateral margins subparallel; sternite 5 slightly narrower than sternite 4; sternite 6 subequal in width to sternite 4; sternite 7 slightly broader than sternite 6. Surface of sternite 3 shallowly depressed in ventral view, anterolaterally rounded, with 2 small processes on lateral margin; anterior margin concavely excavated, with U-shaped median notch, without distinct flanking spines; lateral end with small spine. Sternite 4 with anterolateral margin anteriorly produced into spine, overreaching anterior end of sternite 3, about 1.2 times as long as posterolateral margin. Anterolateral margins of sternite 5 subparallel, anteriorly rounded, as long as posterolateral margin of sternite 4. Pleon. Smooth. Pleonite 2 with tergite 3.3 times as wide as long; pleuron posterolaterally blunt. Pleuron of somite 3 posterolaterally blunt. Telson half as long as broad; posterior plate 1.4 times longer than anterior plate, posterior margin feebly convex or feebly concave. Eye. Peduncle proximally broader, 1.7 times as long as broad, reaching half of rostrum distally. Cornea not dilated, half as long as remaining eyestalk. Antennule and antenna. Ultimate article of antennular peduncle 3.5 times longer than high. Antennal peduncle overreaching eye and tip of rostrum. Article 2 with distinct distolateral spine. Antennal scale not overreaching article 5, 1.4 times as wide as article 5. Articles 4 and 5 each with strong distomesial spine. Article 5 1.3 times as long as article 4, its width 0.8 times that of ultimate article of antennule. Mxp 3. Ischium without distinct distoventral spine lateral to rounded flexor distal margin; crista dentata with about 12 denticles of variable size, some very small. Merus 1.2 times as long as ischium, mesial face flattish, flexor margin sharply ridged along distal third, with small distolateral tooth; extensor margin with subacute spine in distal half, with strong distal spine. Carpus with 3 spines on lateral and extensor surfaces, in addition to prominent distolateral spine. P 1. Moderately elongate, 3.4 times carapace length, densely covered with fine long setae, mostly in tufts. Ischium dorsally with 2 spines, distal spine strong, proximal spine very small; ventromesial margin with strong subterminal spine followed by several proximally diminishing, smaller spines. Merus about 1.3 times as long as carapace, with distinct spines as following: 4 ventral, 1 distomesial, 1 ventro-distomesial, 1 ventro-distolateral, and 3 obliquely arranged proximal ventromesial spines. Carpus subcylindrical, 1.5 times as long as merus, ventrally with 2 distal spines (mesial and lateral). Palm 3.4 times as long as broad, 1.2 times as long as carpus; mesial and lateral margins bluntly ridged. Fingers not gaping, distally slightly incurved; movable finger almost 0.5 (left) or 0.4 (right) times length of palm, opposable margin with subtriangular blunt proximal process, latter fitting into hiatus proximal to low eminence on opposable margin on fixed finger; remaining opposable margin of fixed finger sinuous. P 2 – 4. Meri compressed mesio-laterally, moderately broad; mesial face somewhat flattened, successively shorter posteriorly (P 3 merus 0.9 times length of P 2 merus, P 4 merus 0.8 times length of P 3 merus), subequally broad in P 2 – 4; length-width ratio: 3.9 in P 2, 3.8 in P 3, 3.1 in P 4; dorsal margin with few small proximal spines, latter always distinct on P 2, occasionally obsolescent on P 3, obsolete on P 4; ventrolateral margin with 2 spines in distal half, distally produced into strong spine in P 2, unarmed and distally lobe-like, not sharply produced in P 3 – 4; P 2 merus about 0.8 times length of carapace, slightly longer (about 1.1 times) than P 2 propodus; P 3 merus as long as P 3 propodus; P 4 merus 0.8 times length of P 4 propodus. Carpi relatively short, subequal (slightly longer in P 2 than in P 3 – 4, or successively slighter shorter posteriorly), length less than half that of propodus; distal margin armed with spines in P 2, unarmed in P 3 – 4. Propodi subequal in P 3 – 4, slightly shorter and stouter in P 2; flexor margin slightly concave, with 1 pair of terminal movable spines preceded by 3 – 5 single movable spines on flexor margin. Dactyli subequal in length in P 2 – 4, slightly shorter in P 2 compared to P 3 – 4, slightly longer than carpi (dactylus-carpus length ratio: 1.1 in P 2, 1.2 in P 3, 1.3 in P 4); flexor margin nearly straight, bearing slender terminal spine preceded by 3 subtriangular spines, diminishing in size proximally and almost perpendicular to margin, and 3 more slender, somewhat inclined proximal spines; diminishing in size proximally. Colour in life. Body and appendages ivory white, somewhat translucent (Fig. 6 B).	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6FFF89FF26FC5A1B91C94E.taxon	distribution	Distribution and bathymetry. Western Indian Ocean: from southern Madagascar to the Red Sea, currently known depth range: 220 – 333 m (Baba et al. 2024; present study).	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6FFF89FF26FC5A1B91C94E.taxon	biology_ecology	Ecology. The holotype was found clinging to a gorgonian octocoral, Melithaea sp. (Alcyonacea, Melithaeidae, species under study), strongly suggesting a symbiotic association (Fig. 6 A, C).	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B6FFF89FF26FC5A1B91C94E.taxon	discussion	Remarks. Within the speciose genus Uroptychus, U. deliquus most closely resembles U. galene Baba, Corbari & Macpherson, 2024, which was described from Madagascar (Baba et al. 2024), and also has affinities with U. tridentatus (Henderson, 1885) from the Philippines and Indonesia, U. oxymerus Ahyong & Baba, 2004 from north-western Australia, and U. annae Baba, 2018 from New Caledonia and Vanuatu (Baba 2005, 2018). All these species share (1) the general shape of the carapace; (2) the rostrum with subterminal spines; and (3) the presence of a median notch on the anterior margin of the third sternite in the sternal plastron. The Red Sea specimen herein assigned to U. deliquus (hereafter U. deliquus — RS) is morphologically somewhat different from the type specimens of U. deliquus from Madagascar (hereafter U. deliquus — M). The two most noticeable differences between the Red Sea and Madagascan populations are (1) the third sternite is anterolaterally rounded, with 2 small processes on the lateral margin in U. deliquus — RS vs. anterolaterally sharply produced and bifid distally in U. deliquus — M; and (2) the flexor margin of the P 2 – 4 propodi is armed with 3 – 5 spines in the distal half in U. deliquus — RS vs. armed with 1 – 3 spines in in U. deliquus — M (cf. Figs. 4, 5; Baba et al. 2024). The genetic divergence in COI sequences between U. deliquus — RS and U. deliquus — M ranged between 2.5 % and 3 %. Our phylogenetic analysis recovered all populations of U. deliquus within a single clade (Fig. 3), providing no support for the Red Sea population as a genetically distinct lineage that could be considered as a separate species. Nevertheless, the subtle morphological discrepancies, biogeographical criteria (Red Sea endemism among deep-water taxa, see Türkay 1996; Anker et al. 2023), and a relatively long branch of U. deliquus — RS may reflect an incipient speciation process.	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B64FF8DFF26FE0A1C6DCA12.taxon	description	Galathea aegyptiaca Paulson, 1875 Galathea ahyongi Macpherson & Robainas-Barcia, 2015 Galathea anepipoda Baba, 1990 Galathea australiensis Stimpson, 1858 Galathea boucheti Macpherson & Robainas-Barcia, 2015 Galathea brevimana Paul’son, 1875 Galathea celiae Macpherson & Robainas-Barcia, 2015 Galathea erythrina Macpherson & Robainas-Barcia, 2015 Galathea eulimene Macpherson & Robainas-Barcia, 2015 Galathea genkai Miyake & Baba, 1964 Galathea lemaitrei Macpherson & Robainas-Barcia, 2015 Galathea longimana Paulson, 1875 Galathea parvula Macpherson & Robainas-Barcia, 2015	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B64FF8DFF26FB5E1B7CCB0E.taxon	description	Phylladiorhynchus hylas Rodriguez-Flores, Macpherson & Machordom, 2021	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
03EF09628B64FF8DFF26F91E1D38C9DE.taxon	description	Trapezionida dispar (Macpherson & Baba, 1993) Trapezionida eudora (Macpherson & Baba, 1993) Trapezionida janetae (Tirmizi & Javid, 1992) Trapezionida japonica (Stimpson, 1858) Trapezionida roshanei (Tirmizi, 1966)	en	Anker, Arthur, Rodríguez-Flores, Paula C., Bähr, Susanne, Barreca, Federica, Dunn, Natalie, Rodrigue, Mattie, Qurban, Mohammed, Duarte, Carlos M., Pieribone, Vincent (2025): Previously undetected diversity of mesophotic squat-lobsters in the Red Sea, exemplified by a new species of Sadayoshia Baba, 1969 and a new record of Uroptychus deliquus Baba, Corbari & Macpherson, 2024 (Anomura: Galatheoidea and Chirostyloidea). Zootaxa 5686 (4): 572-586, DOI: 10.11646/zootaxa.5686.4.7, URL: https://doi.org/10.11646/zootaxa.5686.4.7
