taxonID	type	description	language	source
03EC543E4C0C9666FF0BFD2A6F6FD8CC.taxon	description	(Fig. 1, 2, 6)	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0C9666FF0BFD2A6F6FD8CC.taxon	materials_examined	Type material examined Holotype, ♀, NHMW, glued to a mounting card, labels: (a) white rectangular cardboard, printed: “ 10 - 15. V. 1963 / Dünen S v. / Beirut. Libanon / Kasy & Vartian ”; (b) red rectangular cardboard, printed: “ TYPUS ”; (c) red rectangular cardboard, printed: “ Ochodaeus / berytensis m. / Petrovitz ”; d) white rectangular cardboard, printed: “ NHMW-ZOO- / COL- 0002640 ”. Other specimens examined (290 ex.) ISRAEL Upper Galilee - Tel Dan, 15. VI. 2002, light trap, V. Kravchenko (1 ex. / SMNH); - Elon, 9. III. 1970, leg. Schweiger / Ochodaeus berytensis, det. Petrovitz (1 ex. / SMNH); - Nahal Keziv, 12. IV. 2006, light trap, V. Kravchenko, V. Chikatunov (1 ex. / SMNH); - Nahal Kziv, 20. II. 1999, M. Finkel (1 ex. / SMNH); ditto, 24. IV. 1999 (1 ex. / SMNH); ditto, 9. IV. 1999 (1 ex. / SMNH); ditto, 8. V. 1999 (1 ex. / SMNH); ditto, 4. VI. 1999 (2 ex. / SMNH); ditto, 17. VI. 2000 (1 ex. / SMNH); - Nahal Keziv, Upper Galilee, close Montfort, 11. V. 2010, trap 4, 33 ° 02 ’ 43 ’’ / 35 ° 13 ’ 24 ’’, semi-dry, leg. C. Drees (1 ex. / CJBH; 1 ex. / CSZF); - Nahal Keziv, 10. V. 2020, M. Finkel & H. Inbar (3 ex. / SMNH); ditto, 28. V. 2020 (2 ex. / SMNH); ditto, 10. VII. 2020 (1 ex. / SMNH); - IL, Nahal Kziv NR, cca 253 m, Montfort castle env., 33 ° 2 ’ 40.48 ’’ N, 35 ° 13 ’ 33.72 ’’ E, 17. - 23. V. 2017, Martin Just leg., / Ochodaeusberytensis [???] [sic!] Petrovitz, 1965, det. A. Bezdĕk, 2019 (1 unsexed / IECA); - Bar’am, 15. XI. 2016, 262154, ex. truffle, O. Danai, N. Ezov (1 ♂ / SMNH; 1 ex. / CJBH); ditto, n ° 262155 (1 ex. / SMNH); - Matityahu Reserve station, on Tuber aestivum, IV- 2019, L. Azoulay leg. (10 ex. / CJBH); - Bar'am forest (Oak), truffle plantation (by trap), IV- 2019, L. Azoulay leg. (82 ex. / MNHN & CJBH); - Bar'am (natural forest), 7 - V- 2019 (by trap), L. Azoulay leg. (50 ex. / MNHN & CJBH); ditto, 27 - VII- 2019 (45 ex. / MNHN & CJBH); - Bar’am forest, old-growth woodland, 262746, 33 ° 02 ’ N 35 ° 25 ’ E, 674 m, 16. VI. 2005, A. Timm, T. Assman (1 ex. / SMNH); ditto, n ° 262745 (1 ex. / SMNH); ditto, 26. IV. 2005, n ° 262739 (1 ex. / SMNH); ditto, 25. VI. 2005, n ° 262720 (1 ex. / SMNH); ditto, 26. XII. 2005, n ° 262774 (1 ♂ / SMNH); ditto, 6. II. 2006, n ° 262790 (1 ♂ / SMNH); ditto, 6. II. 2006, n ° 262791 (1 ex. / SMNH); ditto, 6. II. 2006, n ° 262792 (1 ex. / SMNH); - Ziv’on, batha, 33 ° 01 ’ N / 35 ° 21 ’ E, 85193, 16. IV. 2005, pitfall, A. Timm, T. Assmann (1 ex. / SMNH); - Hurfeish, batha, 33 ° 01 ’ N 35 ° 21 ’ E, 16. IV. 2005, 85221, Pitfall, A. Timm, T. Assmann (1 ex. / SMNH); ditto, 6. VI. 2005 (1 ex. / SMNH); - ' En Ya’aqov, 19. III. 2007, I. Shtirberg (1 ex. / SMNH); - Har Meron, above Field School, 4 - 11. IV. 2007, T. Levanony (2 ex. / SMNH); - Har Meron, 2. VII. 2002, light trap, V. Kravchenko (1 ex. / SMNH): ditto, 12. VII. 2002 (3 ex. / SMNH); - Har Meron, 20. IV. 2002, light trap, V. Kravchenko (2 ex. / SMNH): - Har Meron, 5. X. 2002, light trap, V. Kravchenko, V. Chikatunov (1 ex. / SMNH); Golan Heights - Mas'ade, 25. IV. 2002, V. Kravchenko (16 ex. / SMNH & CJBH); - Odem forest, 33 ° 12 ’ N 35 ° 44 ’ E, 5. VI. 2009, leg. G. Sabatinelli (1 ex. / CSZF); - Buq'ata, 1. VIII. 2002, 4663, V. Kravchenko, V. Chikatunov (7 ex. / SMNH & CJBH); Carmel Ridge - Nahal Oren, 24. II. 1998, light, V. Chikatunov, T. Pavliček (2 ex. / SMNH); - Nahal Oren, 4. VI. 1999, NFS 6, light trap, V. Chikatunov, T. Pavliček (1 ex. / SMNH); - Nahal Oren, 09. V. 2010, trap 6, northfacing slope, 32 ° 42 ’ 52 ’’ / 34 ° 59 ’ 33 ’’, semi-dry, leg. C. Drees (1 ex. / CJBH; 1 ex. / CSZF); ditto, trap 4 (2 ex. / CSZF); Judean Hills - Zur Hadassa, 31. III. 2001, Y. Mandelik (18 ex. / SMNH); - Matta’, 19. IV. 2007, I. Shtirberg (1 ex. / SMNH); Foothills of Judea - ' Adullam, 15. V. 2008, O. Skutelsky / Ochodaeus berytensis, det. V. Chikatunov (1 ♂, 1 ♀ / SMNH); - ' Adullam, 16. V. 2008, O. Skutelsky (2 ex. / SMNH).	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0C9666FF0BFD2A6F6FD8CC.taxon	distribution	Distribution. – Israel, Lebanon, Turkey, Jordan	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0C9666FF0BFD2A6F6FD8CC.taxon	description	Brief redescription Relatively variable in size. Length 4.6 – 7.6 mm. Body unicolor, entirely pubescent, dorsally and ventrally shiny orange-brown to chestnut brown. Head. – Transverse. Clypeus transverse, broadly rounded anteriorly, finely bordered and darkened along margin; clypeofrontal suture straight, deeply impressed, forming a pronounced transverse groove reaching the upper eye margin laterally. Labrum thick, broadly emarginate anteriorly. Eyes convex, subspherical, relatively small, diameter about equal to length of genae. Cephalic punctation coarse, squamose. Antennae: scape orange-yellow; club yellowish except darkened inner segment basally. Pronotum. – Convex, fully bordered, punctation similar to that of the head. Elytra. – With well-marked striae of medium-sized, closely spaced punctures; interstriae flat with 4 – 5 rows of setigerous granules. Humeral callus well developed. Scutellum triangular, tongue-shaped, apex rounded. Legs. – Protibiae tridentate. Profemora with sharp flat tooth near femorotibial joint (inward-directed) and medial, obtusely rounded tooth forming a flattened triangular bulge in male. Meso- and metafemora each with a sharp flat tooth near femorotibial joint; metafemoral tooth larger with denticulate upper edge. Genitalia. – Aedeagus with elongate, dorsoventrally curved phallobase, acuminate at apex; parameres short, symmetrical. Endophallus with three endophallites: two broad serrated plates and one smaller, arch-shaped apical endophallite with denticulate apex. Sexual dimorphism (Fig. 2). – Male: profemur with a medial, obtusely rounded tooth forming a flattened triangular bulge. Pollex [distal tooth located at the junction of the internal apical spur of the protibia] flat in cross-section, slightly concave dorsally in males; round in section, regularly pointed apically in females.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0C9666FF0BFD2A6F6FD8CC.taxon	diagnosis	Comparative diagnosis. – Ochodaeus berytensis is easily distinguished from the two other species by its robust habitus, very transverse clypeus, and the absence of a clypeal horn in both sexes.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0C9666FF0BFD2A6F6FD8CC.taxon	biology_ecology	Ecology and trophic association. – Ochodaeus berytensis has recently been identified as a mycetophagous pest infesting Tuber aestivum truffle orchards in Upper Galilee (Israel) (Huchet et al. 2022). Adult beetles actively feed on mature truffle fruit bodies, boring tunnels and cavities that significantly compromise their commercial value. While the beetles were consistently found within the sporophores, no immature stages were detected, suggesting that the species does not complete its life cycle in the truffle itself, but rather uses it as a feeding resource. This represents the first confirmed association between an Ochodaeidae species and a hypogeous fungus, providing direct evidence of mycophagy within this family. The seasonal infestation peaks between April and July and affects both cultivated and nearby natural habitats, underscoring the potential threat this species poses to truffle farming in the Mediterranean region.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0D9665FC06FD2269F3DE52.taxon	description	(Fig. 3, 4, 6)	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0D9665FC06FD2269F3DE52.taxon	materials_examined	Type material examined Holotype, ♂, (MNHN), pinned specimen, labels: (a) Circular, blue, handwritten (in Marseul’s hand): “ Ochodaeus / n. gigas m. / Ouargla / 1 [8?] 54 ”; (b) Rectangular, pale blue, printedlabel: “ MUSEUM PARIS / coll. / DE MARSEUL ”; (c) Rectangular, red, printed: “ TYPE ”; (d) Rectangular, white, handwritten and printed: “ Holotype / Ochodaeus gigas / Mars. / J. B. Huchet det. 2025 ”. Other specimens examined (28 ex.) ISRAEL Upper Galilee - Har Meron, 15. III. 2002, V. Kravchenko (2 ex. / SMNH); idem but 20. IV. 2002 (1 ♀ / SMNH); Central Coastal Plain - Tel Aviv, 12. XI. 2002, V. Kravchenko, V. Chikatunov (2 ♂ / SMNH); idem but 12. IV. 2003 (2 ♂, 1 ♀ / SMNH); - TelAviv, 3. II. 1978, A. Freidberg (1 ex. / SMNH); Southern Coastal Plain - Nizzanim, 2. IV. 2001, V. Chikatunov (1 ♂, 1 ♀ / CJBH; 1 ♂, 1 ♀, 1 ex. / SMNH); idem but 15. I. 2002 (1 ex. / SMNH); - Nizzanim, 23. I. 2003, light trap, V. Kravchenko, V. Chikatunov (1 ex. / SMNH); North Negev - Retamim, 28. XI. 2002, light, V. Kravchenko, V. Chikatunov (1 ♂ / SMNH); idem but 5. VI. 2003, light trap (1 ex. / SMNH); - Israel, Retamim, 12. VI. 2002, light trap, V. Kravchenko (1 ex. / SMNH); - N. Negev, Retamim, Fenix, 5. V. 2002, V. Kravchenko / Ochodaeus cornifrons det. V. Chikatunov (1 ♀ CJBH; 3 ex. / SMNH); Central Negev - ' Ezuz, 15. III. 2003, lighttrap, V. Kravchenko, V. Chikatunov (1 ♀ / CJBH; 1 ex. SMNH); - ' En Zin, 18. III. 2001, light trap, V. Kravchenko, I. Yarom (1 ♂ / SMNH); - Israel, ' En Zin, 27. II. 2001, V. Chikatunov (1 ex. / SMNH); Arava Valley - Israel, Shezaf Nat. Res., 17. III. 2001, light trap, V. Kravchenko, I. Yarom (1 ♀ / SMNH).	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0D9665FC06FD2269F3DE52.taxon	distribution	Distribution. – Ochodaeus gigas is mainly found in desert regions of North Africa and the Middle East, including Morocco, Algeria, Tunisia, Libya, Egypt, Israel, and Saudi Arabia (new country record). Given its current distribution, the species is likely to be present in adjacent countries, including Jordan, Lebanon, and Syria. Brief redescription Relatively variable in size. Length 4.5 – 9.0 mm. Body light reddish-brown to yellowish-brown; anterior clypeal margin, pronotal margins, elytral suture, and outer margins narrowly darkened. Head. – Distinctly transverse, width across eyes at least three times greater than median length. Punctation dense, coarse, granulose; surface setose. Clypeus very short, anterior margin flattened, not emarginate, with small central horn laterally compressed and connected to margin by a ridge. Clypeofrontal suture obscured by base of horn. Labrum deeply emarginate, setose. Eyes large, globose, laterally produced.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0D9665FC06FD2269F3DE52.taxon	description	Pronotum. – Coarsely, irregularly punctate; punctation dense and rugose on disc, granulose antero-laterally, surface with short, erect setae. Elytra. – With large humeral calli; striae fine, distinct; interstriae densely and deeply punctate, each puncture anteriorly bordered by a strong granule; surface with short, erect setae. Scutellum with parallel sides at base. Legs. – Protibiae tridentate, basal tooth smaller. Profemora with small curved tooth at apex of infero-anterior margin; males with additional blunt tooth at mid-infero-anterior edge. Meso- and metafemora with similar tooth at apex of infero-posterior margin. Metatibia slender with median row of spines. Abdomen. – Sternites sclerotized with distinct dorsolateral margin; stridulatory peg present. Genitalia. – Aedeagus with elongate, dorsoventrally curved phallobase, acuminate at apex; parameres short, symmetrical, slightly diverging apically. Endophallus with four serrated endophallites, unequal in size, grouped near base. Sexual dimorphism. – Weakly developed. Pollex usuallyshapedas a small, round-sectioned spur, apically pointed and curved outward in its apical third in males; in females, it takes the form of a small, flat, triangular tooth. Major males typically exhibit a profemur bearing a broad, blunt, yet discreet tooth at mid-length; this tooth may be barely visible or even absent in minor males.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0D9665FC06FD2269F3DE52.taxon	diagnosis	Comparative diagnosis. – In this species, the most distinctive character is the large clypeal horn, from the base of which arises a longitudinal carina extending to the anterior margin of the clypeus, giving a triangular profile in lateral view (Fig. 3, 4). The eyes, large, subspherical and globose, occupy nearly the entire width of the head. In contrast, the eyes are distinctly smaller in O. berytensis, with their diameter equal to or less than the length of the genae, and reniform in lateral view in O. (Diplochodaeus) alleonis).	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0E9664FCF0FB4D6B45DC9A.taxon	description	(Fig. 5, 6)	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0E9664FCF0FB4D6B45DC9A.taxon	materials_examined	Type material examined Lectotype, ♀, NMNHS, specimen glued on paper card, dissected, the genitalia stored in a microvial containing glycerin pinned under the specimen. Labels: (a), rectangular, framed, handwritten: “ Dobrodja ”; (b) rectangular, yellowed paper, handwritten: “ Ochodaeus / Alleonis Fairm. / N. S. ”; (c) white, rectangular, printed: “ ♀ ”; (d) red, rectangular, printed: “ LECTOTYPE ”; (e) red, rectangular, printed: “ Ochodaeus alleonis Fairmaire, 1884, LECTOTYPE, J. - B. Huchet des. 2024 ”. Other specimens examined (5 ex.) ISRAEL Central Coastal Plain - Herzliyya, 13. II., leg. Bytinski. Solz., coll. Petrovitz, inscutellaris (1 ♀ / MHNG); - Herzliyya, hill, 32 ° 11 ʹN 34 ° 49 ʹE, 7. III. 2008, A. Freidberg (1 ♂ / SMNH> CJBH); - Herzliyya, hill, 32 ° 11 ʹN 34 ° 49 ʹE, 17. III. 2012, A. Freidberg, 120418 (1 ♀ / SMNH); - TelAviv, AbuKabbir, 25. III. 2025, G. Pisanty (1 ♂ / SMNH); Jordan Valley - 50 m, Umm Zuqa Nature Reserve, Rt. 5788, 18. III. 2008, V. Chikatunov (1 ♀ / SMNH).	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0E9664FCF0FB4D6B45DC9A.taxon	discussion	Remark about Ochodaeus inscutellaris Pic, 1900 Described from the Jordan Valley, near Jericho, on the bank of the Jordan River (approx. 31 ° 52 ′ 0 ″ N 35 ° 27 ′ 0 ″ E), Ochodaeus inscutellaris was based on a single specimen, the holotype of which is no longer present in Pic’s collection housed at the Muséum national d’Histoire naturelle (MNHN) and is presumed lost. The study of four specimens from Israel that match the original description has enabled clarification of the taxonomic status of this species. As concluded by Huchet & Hillert (2025), O. inscutellaris should be regarded as a junior synonym of O. alleonis Fairmaire, 1884. Brief redescription Relatively variable in size. Length 4.5 – 7.3 mm. Habitus consistently bicolored: pronotum black, elytra reddish brown. Head. – Transverse. Clypeus subrectilinear to slightly concave medially, then strongly sinuate toward the antero-internal eye margins; small horn present in both sexes, contiguous with anterior margin. Labrum deeply emarginate. Eyes reduced, reniform. Mentum with narrow anteromedian depression. Gula convex. Cephalic punctation scale-like with contiguous setiferous granules.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0E9664FCF0FB4D6B45DC9A.taxon	description	Pronotum. – Transverse, fully bordered, black, widest at base; basal margin with dense yellowish setae; disc punctation dense and granulate. Elytra. – Short, punctate-striate; interstriae with rugose background and 3 – 4 more or less aligned rows of punctures. Juxta-sutural interval depressed behind scutellum, then raised into a carina extending to the apex; often bearing a dark triangular patch nearly reaching apex. Apex subrectilinear in posterior view. Humeral callus distinct. Scutellum black, narrow, elongate. Genitalia. – Aedeagus with elongate, dorsoventrally curved phallobase, acuminate at apex; parameres short, symmetrical. Endophallus with three serrated endophallites in transverse plates, serrated along convex margin. Sexual dimorphism. – Males with prominent medial tooth on profemur and an inner notch on protibiae just behind the pollex.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
03EC543E4C0E9664FCF0FB4D6B45DC9A.taxon	diagnosis	Comparative diagnosis. – Ochodaeus alleonis is easily distinguished from the two other species by its bicolored habitus, reniform eyes (regularly convex and globular in Ochodaeus berytensis and O. gigas), elongate scutellum, and the presence of a small clypeal horn in both sexes. O. gigas also bears a clypeal horn, but it is distinctly more developed and not contiguous with anterior margin. In O. berytensis, the clypeus is unarmed.	en	Huchet, Jean-Bernard (2025): The Ochodaeidae of Israel: A taxonomic review with key to species and updated distribution (Coleoptera: Scarabaeoidea). Faunitaxys (London, England) 13 (29): 1-8, DOI: 10.57800/faunitaxys-13(29), URL: http://dx.doi.org/10.5281/zenodo.15933344
