taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03ED255D3B3EFFAB99BF681AFE345066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFAB99BF681AFE345066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFAB99BF681AFE345066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFAB99BF681AFE345066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFAB99BF681AFE345066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B3EFFB599BF6DFAFEE75707.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAB99BF6EFBFF1D53A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAB99BF6EFBFF1D53A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAA9A056A9BFF235006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAA9A056A9BFF235006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAA9A056A9BFF235006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAB99BF6CBBFB7D5206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAB99BF6CBBFB7D5206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B20FFAB99BF6CBBFB7D5206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA99BF6E9CFE245526.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA99BF6E9CFE245526.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA99BF6E9CFE245526.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA99BF6E9CFE245526.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA99BF6E9CFE245526.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA99BF6BFCFE595606.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFAA9A056E3BFB1754A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B21FFA89A05693BFF355786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B23FFA89A056D3BFBCC55E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B23FFA89A056D3BFBCC55E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B23FFAF9A056AFBFF145026.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662819/files/figure.png	https://doi.org/10.5281/zenodo.14662819	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662823/files/figure.png	https://doi.org/10.5281/zenodo.14662823	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6BBBFEFB5786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662825/files/figure.png	https://doi.org/10.5281/zenodo.14662825	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF9A056CBBFC605586.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF9A056CBBFC605586.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF9A056CBBFC605586.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF9A056CBBFC605586.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662846/files/figure.png	https://doi.org/10.5281/zenodo.14662846	Fig. 30 Early Pliocene paleogeography of the Mediterranean and adjacent northeastern Atlantic. Paleogeographic configuration based on Popov et al., (2004; middle Pliocene map) and Pérez-Asensio (2021). Zanclean locations with otoliths marked by white numbers with red frame, Piacenzian locations with otoliths marked by black numbers with orange frame. Number of species and authors annotated	Fig. 30 Early Pliocene paleogeography of the Mediterranean and adjacent northeastern Atlantic. Paleogeographic configuration based on Popov et al., (2004; middle Pliocene map) and Pérez-Asensio (2021). Zanclean locations with otoliths marked by white numbers with red frame, Piacenzian locations with otoliths marked by black numbers with orange frame. Number of species and authors annotated	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662848/files/figure.png	https://doi.org/10.5281/zenodo.14662848	Fig. 31 Biogeographic correlation map showing nearest relationships of Early Pliocene otolith-based taxa from Dar bel Hamri. Red arrows and annotations indicate correlations to extant species also known as fossils from the Rharb Basin and relationships of extinct taxa from the Rharb Basin. Map from Blakey, 2020	Fig. 31 Biogeographic correlation map showing nearest relationships of Early Pliocene otolith-based taxa from Dar bel Hamri. Red arrows and annotations indicate correlations to extant species also known as fossils from the Rharb Basin and relationships of extinct taxa from the Rharb Basin. Map from Blakey, 2020	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662823/files/figure.png	https://doi.org/10.5281/zenodo.14662823	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662825/files/figure.png	https://doi.org/10.5281/zenodo.14662825	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662831/files/figure.png	https://doi.org/10.5281/zenodo.14662831	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662833/files/figure.png	https://doi.org/10.5281/zenodo.14662833	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAF99BF6EFBFEA952A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662819/files/figure.png	https://doi.org/10.5281/zenodo.14662819	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B24FFAD9A056BDBFE025406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAD99BF6A5BFAE65286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAD99BF6A5BFAE65286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAD99BF6A5BFAE65286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAD99BF6A5BFAE65286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAD99BF6A5BFAE65286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAD99BF6A5BFAE65286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B26FFAC9A056CDBFD9D50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B27FFAC99BF6EFBFB2E5666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B27FFAC99BF6EFBFB2E5666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B27FFAC99BF6EFBFB2E5666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B27FFAC99BF6EFBFB2E5666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B27FFAC99BF6EFBFB2E5666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B27FFAC99BF6EFBFB2E5666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA29A056C9BFC8E57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA29A056E1BFB5350A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA29A056D7BFC605246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA09A05681BFD9355E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA09A05681BFD9355E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA09A05681BFD9355E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B29FFA09A05681BFD9355E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA099BF6ABBFEFB5606.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA099BF6ABBFEFB5606.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA099BF6ABBFEFB5606.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA099BF6ABBFEFB5606.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA09A056E1BFBD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA09A056E1BFBD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA79A056AA8FE025466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA79A056AA8FE025466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA79A056AA8FE025466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA79A056AA8FE025466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662819/files/figure.png	https://doi.org/10.5281/zenodo.14662819	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2BFFA79A056AA8FE025466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA799BF693BFAD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA799BF693BFAD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA799BF693BFAD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA799BF693BFAD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA799BF693BFAD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA799BF693BFAD554C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2CFFA79A05691BFC6B5606.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2EFFA59A056D3BFB0453A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2EFF9B9A056CFBFF1454A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662819/files/figure.png	https://doi.org/10.5281/zenodo.14662819	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B2EFF9B9A056CFBFF1454A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B10FF9B9A056CDBFC6B5446.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662819/files/figure.png	https://doi.org/10.5281/zenodo.14662819	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B10FF9B9A056E9BFB6D5206.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662819/files/figure.png	https://doi.org/10.5281/zenodo.14662819	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	Fig. 17 Beryciformes and Holocentriformes. a–d Centroberyx vonderhochti n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.219 (reversed), a, c, d paratypes,SMF PO 101.277 (a, c reversed). e–f Centroberyx affinis (GÜnther, 1859), extant, New Zealand (e reversed). g, Sargocentron hastatum (Cuvier, 1829), SMF PO 101.278, Dar bel Hamri. h, Myripristis jacobus Cuvier, 1829, extant, ZMH, Mona 1891. i–l, Myripristis ouarredi n. sp., Dar bel Hamri, Zanclean, i holotype, SMF PO 101.220 (reversed), j–k paratypes,SMF PO 101.221 (j, l reversed). m–n, Liza sp., SMF PO 101.279, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B12FF989A056ADBFD395266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662823/files/figure.png	https://doi.org/10.5281/zenodo.14662823	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF989A056D7BFB995486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF9899BF6CBBFB7F50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF9899BF6CBBFB7F50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF9899BF6CBBFB7F50E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF9E9A05691BFE7F5762.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662823/files/figure.png	https://doi.org/10.5281/zenodo.14662823	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B13FF9E9A05691BFE7F5762.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E9A056CDBFB5B5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662825/files/figure.png	https://doi.org/10.5281/zenodo.14662825	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E9A056CDBFB5B5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E99BF697EFBCC5246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662823/files/figure.png	https://doi.org/10.5281/zenodo.14662823	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	Fig. 19 Gobiiformes. a–c Buenia pulvinus van Hinsbergh & Hoedemakers,2022,SMF PO 101.226 (c reversed),Dar bel Hamri. d Deltentosteus quadrimaculatus (Vlenciennes, 1837), extant, ICM-O 132.8 from the Catalanian sea, Spain,TL =32 mm. e Deltentosteus collonianus (Risso, 1820), extant, ZMUC 14, Mediterranean 8.12.1891. f–j Deltentosteus planus n. sp. Dar bel Hamri,Zanclean, f holotype, SMF PO 101.227, g–j paratypes, SMF PO 101.228 (h, j reversed). k–m, Deltentosteus quadrimaculatus (Valenciennes, 1837),SMF PO 101.229 (reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E99BF697EFBCC5246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E99BF697EFBCC5246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662831/files/figure.png	https://doi.org/10.5281/zenodo.14662831	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E99BF697EFBCC5246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662833/files/figure.png	https://doi.org/10.5281/zenodo.14662833	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E99BF697EFBCC5246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9E99BF697EFBCC5246.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9C9A05695BFD915386.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662825/files/figure.png	https://doi.org/10.5281/zenodo.14662825	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9C9A05695BFD915386.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9C9A05695BFD915386.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B15FF9C9A05695BFD915386.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF9C99BF6DDBFD635406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662825/files/figure.png	https://doi.org/10.5281/zenodo.14662825	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF9C99BF6DDBFD635406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF9C99BF6DDBFD635406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF9C99BF6DDBFD635406.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF9C9A056D7BFC385566.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF9C99BF6A9CFAB250E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662825/files/figure.png	https://doi.org/10.5281/zenodo.14662825	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	Fig. 20 Citharidae and Bothidae. a–c Citharus balearicus Bauza-Rullan,1955, SMF PO 101.230,Dar bel Hamri. d–g Arnoglossus kokeni (Bassoli, 1906), SMF PO 101.231,Dar bel Hamri. h–k Arnoglossus quadratus Schwarzhans, 1999, Dar bel Hamri, Zanclean, j holotype, SMF P 9317, h, i, k paratypes, SMF P 9318. l–o Laeops rharbensis Schwarzhans, 1999, paratypes, SMF P 9320, Dar bel Hamri, Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF939A056BBBFDA65798.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF939A056BBBFDA65798.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662821/files/figure.png	https://doi.org/10.5281/zenodo.14662821	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	Fig. 18 Gobiiformes. a–e Hoeseichthys brioche (Lin et al., 2015), b–e SMF PO 101.222 (b, e reversed), Dar bel Hamri, a SMF PO 101.223,Sidi Mohamed ech Chleuh. f–j Lesueurigobius stazzanensis Schwarzhnans, Agiadi & Carnevale,2020,SMF PO 101.224 (g–i reversed), Dar bel Hamri. k–m Vanneaugobius? sp., k–l SMF PO 101.225 (k reversed),Dar bel Hamri, m, refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°40’N, 08°54’W	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF939A056BBBFDA65798.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF939A056BBBFDA65798.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF939A056BBBFDA65798.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B17FF939A056BBBFDA65798.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B18FF9299BF6874FEA650A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B19FF929A056D61FB95552F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B19FF929A056D61FB95552F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B19FF929A056D61FB95552F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B19FF9299BF6B5BFD7A5766.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B19FF9299BF69BBFA9850E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662827/files/figure.png	https://doi.org/10.5281/zenodo.14662827	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	Fig. 21 Soleidae and Cynoglossidae.a–d Quenselia cornuta Schwarzhans, 1999, Dar bel Hamri, Zanclean, b holotype, SMF P 9325, a, c–d paratypes, SMF P 9326. e–f Dicologlossa hexophthalma (Bennett, 1831), f SMF PO 101.232, Dar bel Hamri, e extant,Lagos fish market, Portugal. g Synaptura sp. SMF PO 101.234,Dar bel Hamri. h Synapturichthys kleini (Risso,1827), SMF PO 101.235, Dar bel Hamri. j–j Microchirus variegatus (Donovan,1808), SMF PO 101.280,Dar bel Hamri. k Vanstraelenia chirophthalma (Regan, 1915), SMF PO 101.236, Dar bel Hamri. l–o Cynoglossus obliqueventralis Schwarzhans, 1999, Dar bel Hamri,21 l holotype, SMF P 9327, m paratype,SMF P 9328, n–o newly collected specimens, SMF PO 101.281	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1BFF9099BF6E9BFBF850E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1BFF979A056D3BFD4E5619.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1CFF969A0569B4FE545086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF695BFB1657A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662831/files/figure.png	https://doi.org/10.5281/zenodo.14662831	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF6D9BFD8D5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF6D9BFD8D5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF6D9BFD8D5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF6D9BFD8D5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF6D9BFD8D5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1DFF9699BF6D9BFD8D5486.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662831/files/figure.png	https://doi.org/10.5281/zenodo.14662831	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6FDBFE185506.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF9599BF6B5BFF0654C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662831/files/figure.png	https://doi.org/10.5281/zenodo.14662831	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	Fig. 23 Haemulidae. a–c Parapristipoma bethensis n. sp., Dar bel Hamri, Zanclean, a holotype, SMF PO 101.285, b–c paratypes,SMF PO 101.286. d– h Pomadasys incisus (Bowdich, 1825), SMF PO 101.241 (d–g reversed),Dar bel Hamri. i, Pomadasys sp. SMF PO 101.244,Dar bel Hamri. j–l, Pomadasys zemmourensis n. sp., Dar bel Hamri, Zanclean, j holotype, SMF PO 101.242 (reversed),k–l paratypes, SMF PO 101.243 (k reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF8B9A0569BBFB805266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662833/files/figure.png	https://doi.org/10.5281/zenodo.14662833	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF8B9A0569BBFB805266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B1EFF8B9A0569BBFB805266.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662829/files/figure.png	https://doi.org/10.5281/zenodo.14662829	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	Fig. 22 Sphyraenidae,Carangidae and Acropomatidae. a Sphyraena sp., SMF PO 101.237,Dar bel Hamri. b–d Trachurus insectus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.283,c–d paratypes,SMF PO 101.284 (reversed). e–g Caranx rharbensis n. sp., Dar bel Hamri, Zanclean, e holotype, SMF PO 101.238,f–g paratypes, SMF PO 101.282 (reversed).h–k Trachurus mediterraneus (Steindachner, 1868), SMF PO 101.239 (h, j reversed), Dar bel Hamri. l–o Verilus mutinensis (Bassoli, 1906), SMF PO 101.240 (m, n reversed), Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662833/files/figure.png	https://doi.org/10.5281/zenodo.14662833	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B00FF8A9A056C7BFEBB5066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B01FF8A99BF6B5BFEFB5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662833/files/figure.png	https://doi.org/10.5281/zenodo.14662833	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	Fig. 24 Cepolidae and Triglidae. a–d Cepola macrophthalma (Linnaeus, 1758),a extant, southwestern France (reversed), b–d Dar bel Hamri, Zanclean, SMF PO 101.247. e–f Cepola pauciradiata Cadenat, 1950, extant, e, CAS 235490, 14°11’N, 17°22’W, f refigured from Schwarzhans (2013a), dredge in the Gulf of Guinea, 04°39’N, 01°08’W. g, Cepola schlegelii Bleeker, 1854, NSMT 110130,Taiwan. h Cepola haastii (Hector, 1881), New Zealand (reversed).i–n Cepola lombartei n. sp., Dar bel Hamri,Zanclean, i holotype, SMF PO 101.245, j–n paratypes, SMF PO 101.246. o Owstonia sp. SMF PO 101.248, Dar bel Hamri.p Peristedion cataphractum (Linnaeus, 1758), SMF PO 101.249, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B01FF8A99BF693BFB725006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B01FF8A99BF693BFB725006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B01FF8A99BF693BFB725006.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B01FF8A9A056E5BFC6B53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B02FF889A0569ADFF065066.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF889A056EDCFBCA5466.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B03FF8E9A056A7CFEE35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B05FF8E99BF69DCFACE53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B05FF8E99BF69DCFACE53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B05FF8D9A056D9CFDB85086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662835/files/figure.png	https://doi.org/10.5281/zenodo.14662835	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	Fig. 25 Trachinidae and Uranoscopidae. a Trachinus armatus Bleeker, 1861, SMF PO 101.250, Dar bel Hamri. b–d Trachinus maroccanus n. sp., Dar bel Hamri, Zanclean, b holotype, SMF PO 101.251, c–d paratypes,SMF PO 101.252 (25c reversed). e–h Trachinus wernlii n. sp., Dar bel Hamri, Zanclean, 25 g holotype, SMF PO 101.253 (reversed), e–f, h paratypes, SMF PO 101.254 (e, h reversed). i Uranoscopus ciabatta Girone et al., 2010, SMF PO 101.287, Dar bel Hamri.j–k Uranoscopus scaber Linnaeus,1758, Dar bel Hamri, SMF PO 101.255 (reversed). l–n Uranoscopus vanhinsberghi n. sp., Dar bel Hamri, Zanclean, l holotype, SMF PO 101.256 (reversed), m–n paratypes, SMF PO 101.257 (n reversed). o Uranoscopus archionema Regan, 1921, Madagascar (reversed). p–q Uranoscopus hoedemakersi n. sp., Dar bel Hamri, Zanclean, p holotype, SMF PO 101.258,q paratype,SMF PO 101.288	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D99BF6A9BFA705086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D99BF6A9BFA705086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D99BF6A9BFA705086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D99BF6A9BFA705086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D99BF6A9BFA705086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D99BF6A9BFA705086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D9A056D1BFC975446.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D9A056D1BFC975446.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D9A056D1BFC975446.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D9A056A5BFB9B57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D9A056A5BFB9B57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8D9A056A5BFB9B57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B06FF8C9A0569DBFD6A53C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C99BF6D9BFE8A5426.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C99BF6D9BFE8A5426.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C99BF6D9BFE8A5426.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C99BF6D9BFE8A5426.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C99BF6ABBFBDD5366.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C99BF6ABBFBDD5366.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF8C9A056D7BFAEB57C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B07FF829A0569DCFEA65326.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662837/files/figure.png	https://doi.org/10.5281/zenodo.14662837	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	Fig. 26 Sparidae.a–b Boops boops (Linnaeus, 1758), a SMF PO 101.259 (reversed), Dar bel Hamri, b SMF PO 101.260 (reversed), Asilah. c Oblada melanura (Linnaeus, 1758), SMF PO 101.262 (reversed), Dar bel Hamri. d–f Diplodus bellottii (Steindachner,1882), SMF PO 101.261 (e–f reversed), Dar bel Hamri. g–i Pagellus bogaraveo (BrÜnnich, 1768), SMF PO 101.264 (reversed),Dar bel Hamri. j Pagellus acarne (Risso, 1827), SMF PO 101.289 (reversed),Dar bel Hamri. k Pagellus bellottii Steindachner, 1882, SMF PO 101.263 (reversed), Dar bel Hamri.l Pagellus erythrinus (Linnaeus,1758), SMF PO 101.265,Dar bel Hamri. m–o Pagrus pagrus (Linnaeus, 1758), SMF PO 101.290, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF6DFBFE1C54A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF6DFBFE1C54A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF6DFBFE1C54A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF693BFC7350E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF693BFC7350E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF693BFC7350E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF693BFC7350E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF8299BF693BFC7350E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B09FF819A056D3BFE655346.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0AFF8199BF691CFB8150A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0AFF819A056D3CFC5452E6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0AFF819A056B3BFA705666.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0AFF879A05687BFF0650A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662839/files/figure.png	https://doi.org/10.5281/zenodo.14662839	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	Fig. 27 Sparidae.a Spicara smaris (Linnaeus,1758), SMF PO 101.292,Dar bel Hamri. b–c Spondyliosoma tingitana n. sp., Dar bel Hamri, Zanclean, c holotype, SMF PO 101.293 (reversed),b paratype,SMF PO 101.294 (reversed). d, Spicara alta (Osório, 1917), SMF PO 101.291, Dar bel Hamri. e–f Dentex canariensis Steindachner,1881, SMF PO 101.266 (f reversed), Dar bel Hamri. g Opsodentex angolensis Poll & Maul, 1953, SMF PO 101.268.h–i Dentex dentex (Linnaeus, 1758), SMF PO 101.267 (h reversed). j–m Opsodentex mordax n. sp., Dar bel Hamri, Zanclean,j holotype, SMF PO 101.269 (reversed),k–m paratypes, SMF PO 101.295 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662842/files/figure.png	https://doi.org/10.5281/zenodo.14662842	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF879A056D1BFBB35786.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0CFF869A05681BFD765286.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662842/files/figure.png	https://doi.org/10.5281/zenodo.14662842	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF8699BF6B1BFEF054A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662842/files/figure.png	https://doi.org/10.5281/zenodo.14662842	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662842/files/figure.png	https://doi.org/10.5281/zenodo.14662842	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662801/files/figure.png	https://doi.org/10.5281/zenodo.14662801	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662805/files/figure.png	https://doi.org/10.5281/zenodo.14662805	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	Fig. 10 Anguilliformes. a–d Panturichthys cf. fowleri (Ben-Tuvia, 1953), SMF PO 101.184 (a reversed), Dar bel Hamri. 10e, Echelus myrus (Linnaeus, 1758), SMF PO 101.216 (reversed), Dar bel Hamri. f–h, Saurenchelys silex van Hinsbergh & Hoedemakers, 2022, Dar bel Hamri, Zanclean, f SMF PO 101.186, g–h tentatively assigned specimens,SMF PO 101.815 (reversed). i–j, Saurenchelys cancrivora Peters, 1864, extant,West Pacific, i BSKU 69087, j BSKU 75296.k, Conger sp. juv., SMF PO 101.187 (reversed), Dar bel Hamri. l, Gnathophis mystax (Delaroche, 1809), SMF PO 101.188 (reversed),Dar bel Hamri. m, Japonoconger africanus (Poll, 1953), SMF PO 101.189 (reversed), Dar bel Hamri. n, Paraxenomystax cf. bidentatus Reid, 1940, SMF PO 101.273, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662807/files/figure.png	https://doi.org/10.5281/zenodo.14662807	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	Fig. 11 Anguilliformes and Siluriformes. a–c Rhynchoconger carnevalei n. sp. 10a paratype, MGPT-PU 130,452 (reversed),Borelli, Piedmont, Italy, late Tortonian, b holotype, SMF PO 101.191, Agrigento,Sicily,Italy, Zanclean, c paratype, SMF PO 101.336 (reversed), Orciano near Pisa,Italy, Zanclean. d, e Rhynchoconger pantanellii (Bassoli & Schubert,1906), SMF PO 101.190 (e reversed),Dar bel Hamri. f Carlarius cf. latiscutatus (GÜnther, 1864), SMF PO 101.192,Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662809/files/figure.png	https://doi.org/10.5281/zenodo.14662809	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	Fig. 12 Myctophidae. a–c Myctophum fitchi (Schwarzhans, 1978), a, c SMF PO 101.194 (a reversed), Dar bel Hamri, b Asilah. d, Hygophum cf. hygomi (LÜtken,1892), SMF PO 101.193,Dar bel Hamri. e Ceratoscopelus maderensis (Lowe, 1839), SMF PO 101.195,Dar bel Hamri. f, Lampanyctus sp. SMF PO 101.196,Dar bel Hamri. g, Diaphus adenomus Gilbert, 1905, SMF PO 101.274 (reversed), Dar bel Hamri. h–l Diaphus cavallonis Brzobohatý & Nolf, 2000, SMF PO 101.197 (j reversed),Dar bel Hamri. m–o Diaphus dirknolfi Schwarzhans,1986, n SMF PO 101.198,Dar bel Hamri, m, o SMF PO 101.199, Sidi Mohammed ech Chleuh	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662811/files/figure.png	https://doi.org/10.5281/zenodo.14662811	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	Fig. 13 Myctophidae. a–c Diaphus draconis Schwarzhans, 2013, SMF PO 101.200 (c reversed), Kef Nsour,Messinian.d–j Diaphus maghrebensis n. sp. f holotype, SMF PO 101.201, Dar bel Hamri,Zanclean, paratypes, h–j SMF PO 101.202 (h, j reversed), Dar bel Hamri, Zanclean, e SMF PO 101.203 (reversed),Kef Nsour, Messinian, d, g SMF PO 101.204, Sidi Mohamed ech Chleuh, Zanclean. k–m, Diaphus taaningi Norman, 1930, extant, k–l MCZ 150978, 36°17’N, 74°49’W, 13 m refigured from Schwarzhans (2013b), dredge in the Gulf of Guinea, 00°20’S, 08°46’E	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662813/files/figure.png	https://doi.org/10.5281/zenodo.14662813	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	Fig. 14 Gadiformes.a–c Physiculus sp. SMF PO 101.205,Dar bel Hamri. d, Bregmaceros sp. SMF PO 101.206, Dar bel Hamri. e, Nezumia ornata (Bassoli, 1906), SMF PO 101.207 (reversed),Dar bel Hamri. f, Trachyrincus scabrus (Rafinesque, 1810), SMF PO 101.275, Dar bel Hamri. g, Merluccius merluccius (Linnaeus, 1758), extant (reversed), Marmara Sea,Turkey. h, Merluccius senegalensis Cadenat, 1950, extant (reversed), CAS 235494, 14°14’N, 17°32’W. i, Merluccius polli Cadenat, 1950, extant, CAS 235466,13°53’N, 17°34’W.j–l, Merluccius polli Cadenat, 1950, SMF PO 101.209 (l reversed), Dar bel Hamri, Zanclean. m–p, Paratrisopterus glaber Schwarzhans,2010, SMF PO 210, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662815/files/figure.png	https://doi.org/10.5281/zenodo.14662815	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	Fig. 15 Ophidiiformes. a Carapus acus (BrÜnnich, 1768), SMF PO 101.211, Dar bel Hamri. b–d, Echiodon dentatus (Cuvier,1829), b, c SMF PO 101.212, Dar bel Hamri, d SMF PO 101.213 (reversed), Sidi Mohamed ech Chleuh. e, Echiodon dentatus, extant, ICM-O 258.6,off Tarragona.f–i, Echiodon praeimberbis (Weiler, 1971), SMF PO 101.214 (g, h reversed), Dar bel Hamri. j–k, Echiodon sp., Holocene dredge samples off Guinea (15j) and Nigeria (k), refigured from Schwarzhans (2013a)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0DFF859A056B1BFE565086.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662817/files/figure.png	https://doi.org/10.5281/zenodo.14662817	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	Fig. 16 Ophidiiformes. a, Brotula aff. multibarbata Temminck & Schlegel, 1846, SMF PO 101.215 (reversed), Dar bel Hamri. b–g, Ophidion tuseti n. sp., Dar bel Hamri, Zanclean,b holotype, SMF PO 101.217 (reversed),c–g paratypes,SMF PO 101.218 (c, e–g reversed). h, Ophidion lozanoi Matallanas, 1990, Holocene dredge samples off Nigeria, refigured from Schwarzhans (2013a). i Ophidion saldanhai Matallanas & Brito, 1999, extant, Gabon, leg. ZMUC (reversed). j, Ogcocephalidae indet., SMF PO 101.276, Dar bel Hamri	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662842/files/figure.png	https://doi.org/10.5281/zenodo.14662842	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662791/files/figure.png	https://doi.org/10.5281/zenodo.14662791	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662793/files/figure.png	https://doi.org/10.5281/zenodo.14662793	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662795/files/figure.png	https://doi.org/10.5281/zenodo.14662795	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662797/files/figure.png	https://doi.org/10.5281/zenodo.14662797	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662799/files/figure.png	https://doi.org/10.5281/zenodo.14662799	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662787/files/figure.png	https://doi.org/10.5281/zenodo.14662787	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662789/files/figure.png	https://doi.org/10.5281/zenodo.14662789	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662844/files/figure.png	https://doi.org/10.5281/zenodo.14662844	Fig. 29 Paleogeographic situation in northern Morocco and southern Spain during Messinian prior to the closure of the Mediterranean (a) and the Zanclean after opening of the Strait of Gibraltar (b), after Martin et al. (2014) and Pérez-Asensio (2021). Otolith-bearing localities are indicated by asterisks (Estepona from van Hinsbergh & Hoedemakers, 2022). Number of species at location in black printing; number of shared species on correlation arrows	Fig. 29 Paleogeographic situation in northern Morocco and southern Spain during Messinian prior to the closure of the Mediterranean (a) and the Zanclean after opening of the Strait of Gibraltar (b), after Martin et al. (2014) and Pérez-Asensio (2021). Otolith-bearing localities are indicated by asterisks (Estepona from van Hinsbergh & Hoedemakers, 2022). Number of species at location in black printing; number of shared species on correlation arrows	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662846/files/figure.png	https://doi.org/10.5281/zenodo.14662846	Fig. 30 Early Pliocene paleogeography of the Mediterranean and adjacent northeastern Atlantic. Paleogeographic configuration based on Popov et al., (2004; middle Pliocene map) and Pérez-Asensio (2021). Zanclean locations with otoliths marked by white numbers with red frame, Piacenzian locations with otoliths marked by black numbers with orange frame. Number of species and authors annotated	Fig. 30 Early Pliocene paleogeography of the Mediterranean and adjacent northeastern Atlantic. Paleogeographic configuration based on Popov et al., (2004; middle Pliocene map) and Pérez-Asensio (2021). Zanclean locations with otoliths marked by white numbers with red frame, Piacenzian locations with otoliths marked by black numbers with orange frame. Number of species and authors annotated	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662803/files/figure.png	https://doi.org/10.5281/zenodo.14662803	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662848/files/figure.png	https://doi.org/10.5281/zenodo.14662848	Fig. 31 Biogeographic correlation map showing nearest relationships of Early Pliocene otolith-based taxa from Dar bel Hamri. Red arrows and annotations indicate correlations to extant species also known as fossils from the Rharb Basin and relationships of extinct taxa from the Rharb Basin. Map from Blakey, 2020	Fig. 31 Biogeographic correlation map showing nearest relationships of Early Pliocene otolith-based taxa from Dar bel Hamri. Red arrows and annotations indicate correlations to extant species also known as fossils from the Rharb Basin and relationships of extinct taxa from the Rharb Basin. Map from Blakey, 2020	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
03ED255D3B0EFFF599BF6D1BFB415405.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14662851/files/figure.png	https://doi.org/10.5281/zenodo.14662851	Fig. 32 Conceptual map of marine fish bioprovinces during the Early Pliocene. From west to east: transitional tropical Maghrebian bioprovince; subtropical Mediterranean bioprovince; temperate northeastern Atlantic bioprovince;Paratethyan bioprovince. Paleogeographic configuration based on Popov et al. (2004); delimitation of bioprovinces altered from Avila et al. (2016)	Fig. 32 Conceptual map of marine fish bioprovinces during the Early Pliocene. From west to east: transitional tropical Maghrebian bioprovince; subtropical Mediterranean bioprovince; temperate northeastern Atlantic bioprovince;Paratethyan bioprovince. Paleogeographic configuration based on Popov et al. (2004); delimitation of bioprovinces altered from Avila et al. (2016)	2023-04-13	Schwarzhans, Werner		Zenodo	biologists	Schwarzhans, Werner			
