taxonID	type	description	language	source
03E8ED412F5ADC2CFF21F961FE4A5F95.taxon	description	Figs. 1 – 2	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5ADC2CFF21F961FE4A5F95.taxon	materials_examined	Type material. Holotype: ♀ adult: COLOMBIA, Antioquia, San Roque, vereda Guacas Abajo, quebrada El Topacio, 880 – 1100 m, bosque fragmentado, 6 ° 29 ' 27 " N, 74 ° 57 ' 5 " W. Pitfall baited with human feces. 4 – 7. IV, 2013. A. Vélez-Bravo leg. (Colo 19) (CEUA). Paratypes: ♀ adult: as holotype, except: (Colo 17) (USNM); ♀ adult: as holotype, except: (Colo 18) (CEMHS).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5ADC2CFF21F961FE4A5F95.taxon	etymology	Etymology. The specific name is a reference to the very short proximal flagellomeres.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5ADC2CFF21F961FE4A5F95.taxon	diagnosis	Diagnosis. Adult female. Only extant species of Corethrella with flagellomeres I – III globular, IV – XIII very elongated (Fig. 1 A); wings, legs with broad scales (Fig. 1 B, D).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5ADC2CFF21F961FE4A5F95.taxon	description	Description Adult female (n = 3). Head (Fig. 1 A). Dark brown. Outline of head in anterior view laterally elongate; head W / L 1.34 – 1.39. Coronal suture complete, reaching ventral margin of interocular space (Fig. 2 A). Clypeus somewhat elongate, 0.76 – 0.88 times as wide as long, with 14 – 18 dorsal setae. Palpus more lightly pigmented than head; third segment swollen at midlength, about 2 times length of fifth. Mandibles serrate, lacinia sclerotized with smooth margins. Antenna (Fig. 1 A): pedicel dark brown, with at least one thicker, distinctive, elongate dorsal seta; flagellum more lightly pigmented than head; flagellomeres I – III very short; XI – XIII elongate; flagellomere XIII with apical bifurcation. Sensilla coeloconica distribution: 1 (XII – XIII), 2 (II, VIII – XI), 3 ([IX]), 4 (I). Cranial sensilla (Fig. 2 A): Ocular row with 2 thick offset setae at ventral part; 9 – 10 thick setae extending shortly beyond vertex; vertex without setae; few slender setae on frons. Subocular row inconspicuous. Postgenal row with 9 – 10 slender / intermediate setae. With 1 ventromedial thick seta. Thorax (Fig. 1 C). Medium / dark brown. Prescutal suture short, ending halfway towards dorsocentral row of setae (Fig. 2 B). Anterior anepisternum divided longitudinally by sinuous suture. Posterior anepisternum divided by a diagonal suture, inferior portion triangular with anterodorsal margin thick. Sensilla (Fig. 2 B): Antepronotum with about 4 slender setae anteriorly. Postpronotum with 1 thick dorsal seta, 2 slender more posterior. Scutum, prescutal area with 2 thick setae anteroventrally and group of 3 thick setae near prescutal suture. Antealar area with cluster of 11 – 12 thick / intermediate setae centrally located. Supraalar area with 2 – 3 thick setae anteroposteriorly aligned, 4 – 6 intermediate / slender setae surrounding. Dorsocentral row, posterior part with 9 – 12 thick offset, longitudinally aligned setae; about 16 thick setae, 18 intermediates completing the row. Scutellum with 10 thick setae. Posterior anepisternum with 1 – 2 slender setae at dorsal part. Anepimeron without setae. Wings (Fig. 1 D). Light brown, without any pattern of pigmentation. Non-marginal veins with broad scales. Apex of R 2 equal to apex of M 1. Halter as dark as scutellum. Wing length 1.46 – 1.51 mm. R 3 / R 1: 0.66 – 0.68. R 2 + 3 / R 2: 0.31 – 0.36. Legs (Fig. 1 B). Light brown, base of midfemur somewhat darker; hind tibia uniformly pigmented. Femora, tibiae, tarsi with broad scales. Midleg tarsomeres 1 – 3 with thick subapical setae. Tarsal claws equal; apically inserted. Empodium with intermediate thickness; intermediate length; with 6 bifid branches (Fig. 2 C). Ta 1 / Ta 2: 2.36 – 2.48. Ta 3 / Ta 4: 1.29 – 1.4. Abdomen (Fig. 1 E). Medium / dark brown, with cercus more lightly pigmented. Adult male: Unknown Immatures: Unknown	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5ADC2CFF21F961FE4A5F95.taxon	discussion	Remarks. This species keys to the couplet 18 - C. ramentum Borkent, 2008 (peruviana group) in Amaral et al. (2023). Corethrella compacta sp. nov. belongs within the peruviana species group. The synapomorphic states shared with the other species in the group, as recognized by Borkent (2008), are the following: wing without pattern of pigmentation; ventral portion of ocular row with two setae; anepimeron without setae; femora with scales; and base of midfemur more darkly pigmented. Moreover, as discussed by Amaral et al. (2023), the arrangement of the scutellar setae on the antealar area is unique for the species in the peruviana group. Within the peruviana group, only C. truncata Borkent has very short flagellomeres I – III, however unlike C. compacta sp. nov., which has an elongate flagellomere IV, C. truncata also has a very short flagellomere IV. The two species can also be distinguished by the distribution of sensilla coeloconica on the flagellum; the presence of posterior anepisternal setae; the scales on the legs, which are broad in C. compacta sp. nov. and narrow in C. truncata (Borkent 2008: 381, Fig. 74 A); and the pattern of pigmentation of legs.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5FDC22FF21FDCAFE0D5A5C.taxon	description	Figs. 3 – 4	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5FDC22FF21FDCAFE0D5A5C.taxon	materials_examined	Type material. Holotype: ♀ adult: CUBA, Guantanamo province, Baracoa Municipality, Alejandro de Humboldt National Park, Monte Iberia, “ Pez Pega ” stream, site 3, 20 ° 29 ' 21 " N 74 ° 44 ' 35 " W. Malaise trap. 29 – 31. X. 2015, Orestes Bello-González leg. (Cuba 02) (PERC). Paratypes: as holotype, except: 20 ° 27 ' 22 " N, 74 ° 44 ' 17 " W, VI. 2017. (Cuba 24) (CEMHS).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5FDC22FF21FDCAFE0D5A5C.taxon	etymology	Etymology. The specific name is a reference to the presence of the coronal suture, which differentiates it from other similar species.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5FDC22FF21FDCAFE0D5A5C.taxon	diagnosis	Diagnosis. Adult female. Only extant species of Corethrella with flagellomeres I – III moderately short (in comparison to subsequent flagellomeres; Fig. 3 A); third segment of palpus short (about 0.75 length of fifth), cylindrical; posterior anepisternum undivided, dorsoventrally elongate (Fig. 3 C). Description Adult female (n = 2). Head (Fig. 3 A). Medium brown. Outline of head in anterior view somewhat laterally elongate; head W / L 1.40 – 1.42. Coronal suture complete, reaching ventral margin of interocular space. Clypeus wide; 1.49 – 1.52 times wider than long, with 4 dorsal setae (Fig. 4 A). Palpus with same degree of pigmentation as head; third segment roughly cylindrical, about 0.75 length of fifth. Mandibles serrate, lacinia sclerotized. Antenna (Fig. 3 A): pedicel dark brown; with at least one thicker, distinctive elongate dorsal seta; flagellum as dark as head; flagellomeres I – III moderately short, XI – XIII short; flagellomere XIII with apical bifurcation. Sensilla coeloconica distribution: 1 (9 – 13), 4 (1). Cranial sensilla (Fig. 4 A): Ocular row with 1 thick offset seta at ventral part; 14 thick setae extending shortly beyond vertex; vertex with several setae. Subocular row inconspicuous. Postgenal row with 6 slender / intermediate setae. With 2 ventromedial thick setae.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5FDC22FF21FDCAFE0D5A5C.taxon	description	Thorax (Fig. 3 C). Medium brown. Prescutal suture short, ending halfway towards dorsocentral row of setae (Fig. 4 B). Anterior anepisternum divided longitudinally by sinuous suture. Posterior anepisternum undivided, dorsoventrally elongate. Sensilla (Fig. 4 B): antepronotum with 4 – 5 intermediate setae distributed on anterior, ventral portions. Postpronotum with 1 thick dorsal seta, 1 – 3 intermediate more posteriorly. Scutum, prescutal area, 3 thick setae dorsoventrally aligned near prescutal suture, with 4 – 5 intermediate / thick setae more anteriorly. Antealar area anterior portion with 2 thick setae dorsoventrally aligned, with 3 thick, 13 slender / intermediate setae mostly distributed on ventral portion. Supraalar area, 3 – 4 thick setae longitudinally aligned, 4 – 5 intermediate surrounding. Dorsocentral row, posterior part with 2 – 3 thick setae; about 12 thick, 24 slender / intermediate setae completing the row. Scutellum with 9 thick setae. Posterior anepisternum without setae. Anepimeron with 2 slender setae. Wings (Fig. 3 D). Medium brown, without any pattern of pigmentation. Non-marginal veins with very narrow scales, resembling thick setae. Apex of R 2 basal to apex of M 1. Halter as dark as scutellum. Wing length 1.20 mm. R 3 / R 1: 0.74 – 0.75. R 2 + 3 / R 2: 0.24 – 0.28. Legs (Fig. 3 B). Medium brown; hind femur with basal 0.75 pale; hind tibia uniformly pigmented. Legs without scales. Midleg tarsomeres 1 – 3 with thick subapical setae. Tarsal claws equal; apically inserted. Empodium with intermediate thickness; intermediate length; with 5 bifid branches. Ta 1 / Ta 2: 3.12. Ta 3 / Ta 4: 1.11. Abdomen (Fig. 3 E). Uniformly medium brown. Adult male: Unknown Immatures: Unknown	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F5FDC22FF21FDCAFE0D5A5C.taxon	discussion	Remarks. This species keys (loosely) to couplets 36 or 45 in Amaral et al. (2023) but reaches no further. Corethrella coronata Amaral & Pinho sp. nov. shares synapomorphies with species in the rotunda group proposed by Borkent (2008). These are: the posterior anepisternum undivided (unique apomorphy) and wing without pattern of pigmentation (homoplastic across the family). The combination of short flagellomeres I – III (in relation to remaining flagellomeres) with a short and cylindrical third segment of palpus is unique within the species group. Two other species in the rotunda group have short flagellomeres I – III, C. brevivena Borkent and C. globosa Borkent, but in both of these species, flagellomeres I – III are much shorter, the third segment of the palpus is distally enlarged, and they lack a coronal suture.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F51DC21FF21FB83FA2B5CB4.taxon	description	Figs. 5 – 6	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F51DC21FF21FB83FA2B5CB4.taxon	materials_examined	Type material. Holotype: ♂ adult: CUBA, Guantanamo province, Baracoa Municipality, Alejandro de Humboldt National Park, Monte Iberia, “ Pez Pega ” stream, site 2, 20 ° 29 ' 44 " N, 74 ° 43 ' 32 " W. Malaise trap. 29 – 31. X. 2015, Orestes Bello-González leg. (Cuba 05) (PERC). Paratypes: ♂ adult: as holotype, except: Forest, site 4, 20 ° 28 ' 57 " N, 74 ° 43 ' 17 " W. (Cuba 06) (PERC); ♂ adult: as holotype, except: 20 ° 27 ' 22.30 " N, 74 ° 44 ' 16.88 " W, VI. 2017. (Cuba 23) (CEMHS); ♂ adult: as previous, except: (Cuba 27) (CEMHS).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F51DC21FF21FB83FA2B5CB4.taxon	etymology	Etymology. The specific name is a combination of the ancient Greek words for spear, dory, and phallus, referring to the pointed tip of the aedeagus.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F51DC21FF21FB83FA2B5CB4.taxon	diagnosis	Diagnosis. Adult male. Only extant species of Corethrella with posterior anepisternum undivided, dorsoventrally elongate (Fig. 5 C); wing with non-marginal veins with only setae (Fig. 5 D); vein R 2 about same length or longer than R 2 + 3; R 2 reaching C near anterior margin of wing; aedeagus strongly tapered subapically, with pointed apex (Fig. 6 E). Description Adult male (n = 4). Head (Fig. 5 A). Dark brown. Outline of head in anterior view laterally elongate; head W / L 1.34 – 1.59. Coronal suture complete, reaching ventral margin of interocular space (Fig. 6 A). Clypeus nearly square, 0.88 – 1.12 times as wide as long, with 1 – 3 dorsal setae. Palpus more lightly pigmented than head; third segment clavate; about 1.14 length of fifth. Mandible, lacinia weakly sclerotized, with smooth margins. Antenna (Fig. 5 A): pedicel dark brown, with at least one thicker, distinctive elongate dorsal seta; flagellum more lightly pigmented than head; flagellomeres I – III moderately elongate, XI – XIII elongate; flagellomere XIII with apical bifurcation. Sensilla coeloconica distribution: 1 (X – XIII), 2 (I). Cranial sensilla (Fig. 6 A): Ocular row with 1 thick offset seta at ventral part; 12 – 13 thick setae extending shortly beyond vertex; vertex with few setae. Subocular row inconspicuous. Postgenal row with 9 – 13 intermediate setae. With 1 ventromedial thick seta.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F51DC21FF21FB83FA2B5CB4.taxon	description	Thorax (Fig. 5 C). Medium / dark brown. Prescutal suture short, ending halfway towards dorsocentral row of setae (Fig. 6 B). Anterior anepisternum divided longitudinally by sinuous suture. Posterior anepisternum undivided, dorsoventrally elongate. Sensilla (Fig. 6 B): Antepronotum with about 3 intermediate setae anteriorly. Postpronotum with 1 thick dorsal and 2 more posterior intermediate setae. Scutum, prescutal area, with 2 thick setae, about 4 intermediate / slender more anteriorly. Antealar area with 1 thick seta near prescutal suture, with 4 thick setae on posteroventral region and about 20 slender / intermediate setae spread from ventral to dorsal portions. Supraalar area with 2 thick setae longitudinally aligned; 2 – 3 slender setae surrounding. Dorsocentral row, posterior part with 2 thick setae; about 20 thick, 20 slender / intermediate setae completing the row. Scutellum with 6 thick setae. Posterior anepisternum with 2 – 3 intermediate setae dorsally. Anepimeron with 1 slender seta. Wings (Fig. 5 D). Medium brown, without any pattern of pigmentation. Non-marginal veins with setae. Apex of R 2 basal to apex of M 1. Halter as dark as scutellum. Wing length 1.03 – 1.10 mm. R 3 / R 1: 0.45 – 0.53. R 2 + 3 / R 2: 0.78 – 1.04. Legs (Fig. 5 B). Medium brown; hind femur with basal 0.6 pale; hind tibia uniformly pigmented. Legs without scales. Midleg tarsomeres 1 – 3 with thick subapical setae. Tarsal claws on foreleg unequal; mid-, hind leg equal; apically inserted. Empodium thick; intermediate length; with 6 bifid branches (Fig. 6 C). Apices of fore-, midtarsi not bilobed. Ta 1 / Ta 2: 2.40 – 2.90. Ta 3 / Ta 4: 1.15 – 1.48. Abdomen (Fig. 5 E). Medium brown, with posterior portion of segment 8 pale. Genitalia (Fig. 5 E). Gonocoxite (Fig. 6 D) uniformly medium brown, elongate; without distinctively elongate spicules on anteromedial area; dorsomedial row well-defined. With 5 dorsal to dorsomedial setae, somewhat thicker than other setae on gonocoxite, strongly tapering distally. With 1 dorsomedial stout seta, tapering from base, located at 0.46 gonocoxite length from base; without sclerotized plate at base. Gonostylus slender, curved subapically; subbasal seta stout, simple; dorsally located. Apical seta peg-shaped, blunt, squat (with its length only slightly exceeding its diameter). Aedeagus slender, tapering from base, with pointed apex; margins fused apically (Fig. 6 E). Adult female: Unknown Immatures: Unknown	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F51DC21FF21FB83FA2B5CB4.taxon	discussion	Remarks. This species keys to the couplet 30 - C. yucuman Caldart & Pinho in Amaral et al. (2023), but differs in the following traits: vein R 2 longer than, or subequal to R 2 + 3; the apex of CuP discretely distal to the base of M 4; and the presence of a sensillum coeloconicum on the tenth flagellomere. Corethrella doryphallica sp. nov. shares synapomorphies with species in the rotunda group proposed by Borkent (2008). These are: the posterior anepisternum undivided (unique apomorphy), wing without pattern of pigmentation (homoplastic), and non-marginal veins with only setae (without scales). The latter only occurs in the clade composed of C. kerrvillensis (Stone), C. remiantennalis Borkent, C. blandafemur Borkent, C. brevivena Borkent, and C. globosa Borkent, plus the unplaced but probably related species, C. yucuman Caldart & Pinho and C. xokleng Amaral, Pinho & Mariano.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F52DC24FF21FCEAFDB95CED.taxon	description	Figs. 7 – 8	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F52DC24FF21FCEAFDB95CED.taxon	materials_examined	Type material. Holotype: ♂ adult: COLOMBIA: Amazonas, Leticia, 11 km, Tanimboca Forest Reserve, 4 ° 7 ' 10 " S, 69 ° 57 ' 4 " W. 7 – 14. XI. 2017, Malaise Trap, G. Kvifte leg. (Colo 08) (CEUA). Paratypes: ♂ adult: as holotype, except: (Colo 06) (PERC); ♂ adult: as holotype, except Cerca Viva Natural Reserve, 4 ° 7 ' 22 " S, 69 ° 56 ' 53 " W. 3 – 7. XI. 2017 (Colo 09) (CEMHS).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F52DC24FF21FCEAFDB95CED.taxon	etymology	Etymology. The specific epithet is a reference to the blunt (obtuse) dorsomedial seta of the gonocoxite.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F52DC24FF21FCEAFDB95CED.taxon	diagnosis	Diagnosis. Adult male. Only extant species of Corethrella with antenna with one sensillum coeloconicum on flagellomeres II – III, VIII – XIII (Fig. 7 A); complete coronal suture; ocular row with one thick ventral seta (Fig. 8 A); palpus as pigmented as head, with third segment only slightly longer than fourth, fifth segments; wing with distinct midlength band of dark scales, with more basal dark scales on C, Sc, and M (Fig. 7 D); halter pale; gonocoxite with dorsomedial seta stout and apically blunt (Fig. 8 D).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F52DC24FF21FCEAFDB95CED.taxon	description	Description Adult male (n = 3). Head (Fig. 7 A). Light brown. Outline of head in anterior view laterally elongate; head W / L 1.57 – 1.93. Coronal suture complete, reaching ventral margin of interocular space (Fig. 8 A). Clypeus nearly square, 1.12 – 1.13 times wider than long, with 1 – 2 dorsal setae (Fig. 8 A). Palpus with same degree of pigmentation as head; third segment roughly cylindrical, about 1.2 length of fifth. Mandibles not clearly visible, lacinia weakly sclerotized. Antenna (Fig. 7 A): pedicel medium brown, with at least one thicker, distinctive elongate dorsal seta; flagellum as dark as head; flagellomeres I – III moderately elongate, or I elongate, II – III moderately elongate; XI – XIII elongate; flagellomere XIII with apical bifurcation. Sensilla coeloconica distribution: 1 (II – III, [VII], VIII – XIII), 2 (I). Cranial sensilla (Fig. 8 A): Ocular row with 1 thick offset seta at ventral part; 13 – 14 thick setae extending to posterior part of head; vertex with few slender setae. Subocular row inconspicuous. Postgenal row with 6 – 9 intermediate setae. With 2 ventromedial thick setae. Thorax (Fig. 7 B). Light / medium brown, with darker pigmentation on anterior portion of anepimeron and around anterior spiracle. Prescutal suture short, ending halfway towards dorsocentral row of setae (Fig. 8 B). Anterior anepisternum divided longitudinally by sinuous suture. Posterior anepisternum divided by a faint diagonal suture, inferior portion triangular with anterodorsal margin thick, ventral margin poorly defined. Sensilla (Fig. 8 B): Antepronotum with 4 thick, 2 slender ventral setae. Postpronotum with 1 thick dorsal seta, 1 intermediate, 2 slender more posteriorly. Scutum, prescutal area, about 4 slender / intermediate anterior setae, row of 3 thick setae dorsoventrally aligned near prescutal suture, about 4 intermediate surrounding. Antealar area anterior portion with 3 – 4 thick setae dorsoventrally aligned and with 6 intermediate / thick setae forming U-shape on ventral portion, about 10 slender setae scattered from ventral to dorsal portions. Supraalar area with 1 – 2 thick setae with about 16 slender / intermediate surrounding. Dorsocentral row, posterior part with cluster of 4 thick setae; about 15 thick, 14 intermediate setae completing the row. Scutellum with 14 thick setae. Posterior anepisternum without setae. Anepimeron with 4 – 5 slender setae. Wings (Fig. 7 D). Light brown, with darker midlength band on all veins, subbasal spot on M, basal spot on C, Sc. Non-marginal veins with narrow scales. Apex of R 2 equal to apex of M 1. Halter lightly pigmented, distinctly lighter than scutellum. Wing length 0.96 – 1.05 mm. R 3 / R 1: 0.44 – 0.47. R 2 + 3 / R 2: 0.64 – 0.80. Legs (Fig. 7 C). Light / medium brown, with distal 0.4 of hind femur pale; hind tibia with well-defined basal, apical bands of darker pigmentation. Legs without scales. Midleg tarsomeres 1 – 3 with thick subapical setae. Tarsal claws on foreleg unequal; mid-, hind leg equal; apically inserted. Empodium with intermediate thickness; intermediate length; with 5 bifid branches (Fig. 8 C). Apices of fore-, midtarsi not bilobed. Ta 1 / Ta 2: 2.88 – 3.14. Ta 3 / Ta 4: 0.81 – 0.98. Abdomen (Fig. 7 E). Medium brown, with segment 9 somewhat more darkly pigmented. Genitalia (Fig. 7 E). Gonocoxite (Fig. 8 D) brown at base, pale at apex; elongate; without distinctively elongate spicules on anteromedial area; dorsomedial row well-defined. With 5 dorsal to dorsomedial setae, somewhat thicker than other setae on gonocoxite, strongly tapering distally. With 1 dorsomedial stout seta, apically blunt, located at 0.26 gonocoxite length from base; without sclerotized plate at base. Gonostylus slender, curved subapically; subbasal seta simple, elongate; anteriorly located. Apical seta slender, elongate. Aedeagus elongate; fused apically, with pointed apex (Fig. 8 E). Adult female: Unknown Immatures: Unknown	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F52DC24FF21FCEAFDB95CED.taxon	discussion	Remarks. This species somewhat keys to couplets 84 or 96 - C. selvicola Lane in Amaral et al. (2023). It can be distinguished from C. selvicola by the elongate coronal suture; the abdominal segment IX somewhat more darkly pigmented than the VIII; and a blunt dorsomedial seta on the gonocoxite. Corethrella obtusa Amaral & Pinho sp. nov. has a combination of apomorphic character states shared with the clade which comprises the fulva, appendiculata, quadrivittata, brakeleyi, and wirthi species groups, according to the phylogeny of Borkent (2008). More inclusively, the presence of a sensillum coeloconicum on flagellomere III would place it among the wirthi group. This character is quite variable across the family and is considered a weak indication of relationship. Corethrella obtusa sp. nov. is similar to C. selvicola Lane and C. atricornis Borkent, differing from the first by having an elongate coronal suture, a distinct pigmentation pattern on the thorax, four posterior setae on the dorsocentral row, and a blunt dorsomedial seta on the gonocoxite. Corethrella atricornis is only known by female specimens, so the comparison is limited. Nevertheless, considering the sexual dimorphism and intraspecific variations observed in other species, C. obtusa sp. nov. may be distinguished by the shape of the palpus, its darker coloration, and by having 4 – 5 setae on the anepimeron (15 in C. atricornis).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F57DC3BFF21FD27FBC75FCD.taxon	description	Figs. 9 – 10	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F57DC3BFF21FD27FBC75FCD.taxon	materials_examined	Type material. Holotype: ♂ adult: CUBA, Guantanamo province, Baracoa Municipality, Alejandro de Humboldt National Park, Monte Iberia, “ Pez Pega ” stream, site 2, 20 ° 29 ' 44 " N, 74 ° 43 ' 32 " W. Malaise trap. 29 – 31. X. 2015, Orestes Bello-González leg. (Cuba 04) (PERC). Paratype: as holotype, except: site 3, 20 ° 29 ' 21 " N, 74 ° 44 ' 35 " W. (Cuba 03) (CEMHS).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F57DC3BFF21FD27FBC75FCD.taxon	etymology	Etymology. The specific name refers to the parallel margins of the aedeagus in this species.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F57DC3BFF21FD27FBC75FCD.taxon	diagnosis	Diagnosis. Adult male. Only extant species of Corethrella with posterior anepisternum undivided, dorsoventrally elongate (Fig. 9 C); non-marginal wing veins with only setae (Fig. 9 D); vein R 2 about same length or longer than R 2 + 3; R 2 reaching in steep angle; CuP reaching wing margin; aedeagus with parallel margins, abruptly tapering apically (Fig. 9 D).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F57DC3BFF21FD27FBC75FCD.taxon	description	Description Adult male (n = 2). Head (Fig. 9 A). Medium brown. Outline of head in anterior view laterally elongate; head W / L 1.37. Coronal suture complete, reaching ventral margin of interocular space (Fig. 10 A). Clypeus nearly square, 1.15 – 1.35 times wider than long, with 3 dorsal setae (Fig. 10 A). Palpus more lightly pigmented than the head; third segment clavate, about 1.25 the length of fifth. Mandibles not clearly visible, lacinia weakly sclerotized, with smooth margins. Antenna (Fig. 9 A): pedicel medium brown, with at least one thicker, distinctive elongate dorsal seta; flagellum more lightly pigmented than head; flagellomeres I – II short, III moderately elongate, XI – XIII elongate; flagellomere XIII with apical bifurcation. Sensilla coeloconica distribution: 1 (XII – XIII), 2 (I). Cranial sensilla (Fig. 10 A): Ocular row with 1 thick offset seta at ventral part; 10 – 11 thick setae extending shortly beyond vertex; vertex with few slender setae; Subocular row inconspicuous; Postgenal row with 7 slender setae. With 2 ventromedial thick setae. Thorax (Fig. 9 C). Light / medium brown, with a pale lateral band comprising anepisternum, dorsal portion of anepimeron, laterotergite, and metanepisternum. Prescutal suture elongate, almost reaching dorsocentral row of setae (Fig. 10 B). Anterior anepisternum divided longitudinally by a sinuous suture. Posterior anepisternum undivided, dorsoventrally elongate. Sensilla (Fig. 10 B): antepronotum with 4 intermediate setae distributed on anterior and central portions. Postpronotum with 1 thick dorsal seta and 3 slender / intermediate more posteriorly. Scutum, prescutal area, 2 – 3 thick setae on ventral anterior portion, with about 5 intermediate / thick setae surrounding. Antealar area with 1 thick seta near prescutal suture, with 3 thick setae on posteroventral region and about 17 slender / intermediate setae spread from ventral to dorsal portions. Supraalar area with 2 thick setae longitudinally aligned; about 5 slender setae surrounding. Dorsocentral row, posterior part with 2 thick setae; about 17 thick, 10 slender / intermediate setae completing the row. Scutellum with 6 thick setae. Posterior anepisternum with 3 intermediate setae centrally located. Anepimeron with 1 slender seta. Wings (Fig. 9 D). Light brown, without any pattern of pigmentation. Non-marginal veins with setae. Apex of R 2 basal to apex of M 1. Halter as dark as scutellum. Wing length 0.80 – 0.84 mm. R 3 / R 1: 0.44 – 0.50. R 2 + 3 / R 2: 1.07 – 1.20. Legs (Fig. 9 B). Medium brown; hind femur with basal 0.75 pale; hind tibia uniformly pigmented. Legs without scales. Midleg tarsomeres 1 – 3 with thick subapical setae. Tarsal claws on foreleg unequal; mid-, hind leg equal; apically inserted. Empodium thick, intermediate length, with 6 bifid branches (Fig. 10 C). Apices of fore-, midtarsi not bilobed. Ta 1 / Ta 2: 2.49 – 2.60. Ta 3 / Ta 4: 1.17 – 1.24. Abdomen (Fig. 9 E). Uniformly medium brown. Genitalia (Figs. 9 E; 10 D, E). Gonocoxite (Fig. 10 D) uniformly medium brown, elongate; without distinctively elongate spicules on anteromedial area; dorsomedial row well-defined. With 5 dorsal to dorsomedial setae, somewhat thicker than other setae on gonocoxite, strongly tapering distally. With 1 dorsomedial stout seta, strongly tapered at apex, located at 0.39 gonocoxite length from base; without sclerotized plate at base. Gonostylus slender, curved subapically; subbasal seta stout, simple and dorsally located. Apical seta peg-shaped, blunt and squat (with its length only slightly exceeding its diameter). Aedeagus slender, with parallel margins, abruptly converging in pointed apex (Fig. 10 E). Adult female: Unknown Immatures: Unknown	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F57DC3BFF21FD27FBC75FCD.taxon	discussion	Remarks. This species keys to couplet 30 - C. yucuman Caldart & Pinho in Amaral et al. (2023). Corethrella parallela sp. nov. presents synapomorphies of species in the rotunda group proposed by Borkent (2008). These are: the posterior anepisternum undivided (unique apomorphy), wing without pattern of pigmentation (homoplastic across the family), and non-marginal veins with only setae (without scales). The latter only occurs in the clade composed of C. kerrvillensis (Stone), C. remiantennalis Borkent, C. blandafemur Borkent, C. brevivena Borkent, and C. globosa Borkent, plus the unplaced but probably related species, C. yucuman Caldart & Pinho, C. xokleng Amaral, Pinho & Mariano, and C. doryphallica sp. nov. It is possible to distinguish C. parallela sp. nov. from both C. yucuman and C. doryphallica sp. nov. by the pattern of pigmentation of the thorax: light / medium brown, with a pale band in C. parallela sp. nov., and uniformly medium / dark brown in C. doryphallica sp. nov. Moreover, the aedeagus with parallel margins is characteristic of C. parallela Amaral & Pinho sp. nov.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F48DC39FF21FE07FB77593F.taxon	description	Figs. 11 – 12	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F48DC39FF21FE07FB77593F.taxon	materials_examined	Type material. Holotype: ♂ adult: CUBA, Guantanamo province, Baracoa Municipality, Alejandro de Humboldt National Park, Monte Iberia, 20 ° 27 ' 22.30 " N, 74 ° 44 ' 16.88 " W. Malaise trap. VI. 2017, Orestes Bello-González leg. (Cuba 25) (PERC).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F48DC39FF21FE07FB77593F.taxon	etymology	Etymology. The specific epithet refers to the presence of three bands on the wing.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F48DC39FF21FE07FB77593F.taxon	diagnosis	Diagnosis. Adult male. Only extant species of Corethrella with the wing without dark scales just distal to the arculus (Fig. 11 E); with dark scales making a subapical band (with no dark scales on these veins extending to the apex of the wing); hind femur with the base pigmented, apical 0.4 pale (Fig. 11 C); with abdominal segment VIII with same degree of pigmentation as segment IX (Fig. 11 F).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F48DC39FF21FE07FB77593F.taxon	description	Description Adult male (n = 1). Head (Fig. 11 A). Medium brown. Outline of head in anterior view laterally elongate; head W / L 1.65. Coronal suture complete, reaching ventral margin of interocular space (Fig. 12 A). Clypeus elongate, 0.79 times as wide as long, with 1 dorsal seta (Fig. 12 A). Palpus more lightly pigmented than head; third segment roughly cylindrical, about 1.5 length of fifth. Mandibles serrate, lacinia sclerotized with smooth margins. Antenna (Fig. 11 B): pedicel dark brown, with at least one thicker, distinctive elongate dorsal seta; flagellum medium brown, with apex of flagellomeres I – III more lightly pigmented; flagellomeres I – III moderately elongate; XI – XIII elongate; flagellomere XIII with apical bifurcation. Sensilla coeloconica distribution: 1 (II, X – XIII), 2 (I). Cranial sensilla (Fig. 12 A): Ocular row with 1 thick offset seta at ventral part; 1 more dorsal; 13 thick setae extending shortly beyond vertex; vertex with several setae. Subocular row inconspicuous. Postgenal row with 6 intermediate setae. With 2 ventromedial thick setae. Thorax (Fig. 11 D). Medium / dark brown. Prescutal suture not clearly visible. Anterior anepisternum divided longitudinally by sinuous suture. Posterior anepisternum undivided, dorsoventrally elongate. Sensilla (Fig. 12 B): antepronotum with 3 intermediate ventral setae, 1 posterodorsal. Postpronotum with 1 dorsal thick seta, 2 small / intermediate more ventrally. Scutum, prescutal area, with 4 thick, 4 intermediate setae scattered. Antealar area anterior portion with 2 – 3 thick setae dorsoventrally aligned, with 8 thick / intermediate setae more posteriorly, forming U-shape on ventral portion, with 5 intermediate setae surrounding. Supraalar area with 4 intermediate setae aligned longitudinally. Dorsocentral row, posterior part with cluster of 6 thick setae; about 12 thick, 12 intermediate setae completing the row. Scutellum with 14 thick setae. Posterior anepisternum without setae. Anepimeron with 6 slender setae. Wings (Fig. 11 E). Light brown, with three bands of darker scales: subbasal band on C, Sc, R 1, M; midlength band on all veins; subapical band on all veins. Non-marginal veins with broad scales. Apex of R 2 basal to apex of M 1. Halter lightly pigmented, distinctly lighter than scutellum. Wing length 1.24 mm. R 3 / R 1: 0.40. R 2 + 3 / R 2: 1.00. Legs (Fig. 11 C). Light / medium brown with knees pale; hind femur with apical 0.4 pale. Legs without scales. Midleg tarsomeres 1 – 3 with thick subapical setae. Tarsal claws on foreleg unequal; mid-, hind leg equal; apically inserted. Empodium slender, intermediate length, with 3 bifid branches (Fig. 12 C). Apices of fore-, midtarsi not bilobed. Ta 1 / Ta 2: 3.13. Ta 3 / Ta 4: 0.75. Abdomen (Fig. 11 F). Uniformly medium brown. Genitalia (Fig. 11 F). Gonocoxite (Fig. 12 D) brown at base, pale at apex, slender; without distinctively elongate spicules on anteromedial area; dorsomedial row well-defined proximally, with two distal setae similar in size to other surrounding setae. With 5 dorsal to dorsomedial setae, somewhat thicker than other setae on gonocoxite, strongly tapering distally. With 1 dorsomedial stout seta, tapering from base, located at 0.27 gonocoxite length from base; without sclerotized plate at base. Gonostylus slender, curved subapically; subbasal seta simple, elongate; anteriorly located. Apical seta or peg absent. Aedeagus elongate; fused subapically; with pointed apex (Fig. 12 E). Adult female: Unknown Immatures: Unknown	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F48DC39FF21FE07FB77593F.taxon	discussion	Remarks. This species keys to couplet 5 - C. whartoni Vargas / C. belkini Borkent in Amaral et al. (2023). It can be distinguished from C. whartoni by having one ventral seta on the ocular row (two in C. whartoni); presence of a sensillum coeloconicum on flagellomere II, lacking on IX; posterior portion of dorsocentral row with six thick setae (four in C. whartoni, but this is a highly variable character in other species); subbasal band on wing not extending to posterior part; apical 0.4 of hind femur pale (hind femur completely pigmented in C. whartoni). Corethrella trivittata sp. nov. can be distinguished from C. belkini by lacking uniformly pigmented tarsi (in C. belkini, at least mid-, hind leg tarsomeres 2 – 4 with banding); abdominal segments VIII and IX equally pigmented (segment IX darker in C. belkini); gonostylus evenly tapering from base (with abrupt apical tapering in C. belkini); apical seta of gonostylus absent or very reduced (slender and elongate in C. belkini). The new species shares the distinctive wing pigmentation pattern of the quadrivittata species group, including one synapomorphy recognized by Borkent (2008), which is the presence of a subapical band of dark scales. Similarly to C. belkini, though, it does not have synapomorphies of the quadrivittata group, so its position in the phylogeny of the family remains somewhat uncertain.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FC9EFBA15DA9.taxon	materials_examined	Material examined. 2 ♀♀ adult: COLOMBIA, Amazonas, Leticia, Cerca Viva Natural Reserve, 4 ° 7 ' 22 " S, 69 ° 56 ' 53 " W, Malaise Trap. 3 – 7. XI. 2017, G. Kvifte leg. (PERC).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FC9EFBA15DA9.taxon	distribution	Distribution. Colombia * (Cerca Viva Natural Reserve, Amazonas), Brazil.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FE6AFB115CED.taxon	materials_examined	Material examined. 6 ♂♂, 4 ♀♀ adult: CUBA, Guantanamo province, Baracoa Municipality, Alejandro de Humboldt National Park, confluence btw. Nibujon and Jaragua rivers, 20 ° 28 ' 30 " N, 74 ° 39 ' 58 " W. Light trap. VI. 2017, Orestes Bello-González leg. (PERC). ♀ adult: as previous, except: Monte Iberia, 20 ° 27 ' 22.30 " N, 74 ° 44 ' 16.88 " W. Malaise trap. (PERC).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FE6AFB115CED.taxon	distribution	Distribution. Cuba * (Alejandro de Humboldt National Park, Guantanamo), Costa Rica.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FBE2FC8E5AAD.taxon	materials_examined	Material examined. 2 ♂♂ adult: COLOMBIA, Amazonas, Leticia, 11 km, Tanimboca Forest Reserve, 4 ° 7 ' 10 " S, 69 ° 57 ' 4 " W. 7 – 14. XI. 2017, Malaise Trap, G. Kvifte leg. (PERC)	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FBE2FC8E5AAD.taxon	distribution	Distribution. Panama, Colombia * (Leticia, Amazonas).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FAE6FEED584D.taxon	materials_examined	Material examined. 7 ♀♀ adult: COLOMBIA, Los Patios, Norte de Santander, 7 ° 50 ' 11 " N, 72 ° 30 ' 27 " W. 24. X. 2017, light trap + frog-call trap, G. Kvifte leg. (PERC).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21FAE6FEED584D.taxon	distribution	Distribution. Mexico, Costa Rica, Panama, Trinidad and Tobago, Colombia * (Los Patios, Santander), Guyana, Brazil, Argentina.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21F98DFBE6595B.taxon	materials_examined	Material examined. ♂ adult: COLOMBIA, Antioquia, San Roque, vereda Guacas Abajo, quebrada El Topacio, 880 – 1100 m, bosque fragmentado, 6 ° 29 ' 27 " N, 74 ° 57 ' 5 " W. Pitfall baited with human feces. 4 – 7. IV, 2013. A. Vélez-Bravo leg. (CEUA).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3EFF21F98DFBE6595B.taxon	distribution	Distribution. Costa Rica, Trinidad and Tobago, Colombia * (Leticia, Amazonas).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3FFF21F8ADFD3F5E91.taxon	materials_examined	Material examined. 3 ♀♀ adult: COLOMBIA, Amazonas, Leticia, 11 km, Tanimboca Forest Reserve, 4 ° 7 ' 10 " S, 69 ° 57 ' 4 " W. 7 – 14. XI. 2017, Malaise Trap, G. Kvifte leg. (PERC).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4DDC3FFF21F8ADFD3F5E91.taxon	distribution	Distribution. Mexico, Costa Rica, Panama, Colombia * (San Roque, Antioquia; Leticia, Amazonas), Guyana, French Guiana, Trinidad and Tobago, Brazil.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4CDC3FFF21FEC1FB5E5F85.taxon	materials_examined	Material examined. ♂ adult: COLOMBIA, Amazonas, Parque Nacional Natural Amacayacu, 150 m, 11 - 24. IV. 2000, Malaise trap, A. Felix leg. (PERC).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4CDC3FFF21FEC1FB5E5F85.taxon	distribution	Distribution. Mexico, Costa Rica, Colombia * (Amacayacu National Park, Amazonas), Brazil.	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4CDC3FFF21FDD5FBE85CBE.taxon	materials_examined	Material examined. 2 ♂♂, 5 ♀♀ adult: COLOMBIA, Chocó province, Nuqui, Jubirá, 5 ° 50 ' 38 " N, 77 ° 16 ' 37 " W. 29. IX. 1999, reared from Werauhia rigens (Griseb.) J. R. Grant, Pérez & Wolff leg. (CEUA). ♂, 2 ♀♀ adult: as previous except: Bahía Solano, Corregimiento El Valle, 6 ° 6 ' 11 " N, 77 ° 25 ' 37 " W. 22. II. 1999, reared from Guzmania musaica (Linden & André) Mez, (CEUA).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
03E8ED412F4CDC3FFF21FDD5FBE85CBE.taxon	distribution	Distribution. Mexico, Costa Rica, Colombia * (Bahía Solano and Nuqui, Chocó).	en	Amaral, André P., Campos, Leonardo Leite Ferraz De, González, Orestes Carlos Bello, Wolff, Marta, Bernal, Ximena E., Pinho, Luiz Carlos (2025): Six new species and records of frog-biting midges (Diptera: Corethrellidae) from Cuba and Colombia. Zootaxa 5689 (1): 58-80, DOI: 10.11646/zootaxa.5689.1.2, URL: https://doi.org/10.11646/zootaxa.5689.1.2
