taxonID	type	description	language	source
03F387F0FFFFFFC1FF57F87EFD190098.taxon	diagnosis	Diagnosis. Cryptocolea imbricata var. minima is characterized by very small size of plants, numerous very long and dense rhizoids, transversely elliptical leaves with cells of leaves almost not differentiated, female bracts not reflexed, and in differentiation in ITS 1 - 2, trn L-F and rbc L nucleotide sequences.	en	Konstantinova, N. A., Vilnet, A. A., Mamontov, Yu. S. (2023): The phylogenetic affinity, distribution and variability of Cryptocolea imbricata r. m. Schust. (Marchantiophyta). Arctoa 32 (2): 137-150, DOI: 10.15298/arctoa.32.11, URL: https://doi.org/10.15298/arctoa.32.11
03F387F0FFFFFFC1FF57F87EFD190098.taxon	materials_examined	Holotype: RUSSIA: Krasnoyarsk Territory, Taimyr Peninsula, Bataika river valley, spotty Carex arctisibirica - lichen moss tundra, 71 ° 12 ' 48 " N, 92 ° 41 ' 26 " E, 29 July 2021, with Trilophozia scitula, Anthelia juratzkana, E. D. Lapshina 213 E / 4 - 3 - 21 [KPABG 124714]. Other specimens examined: Yamalo-Nenets Autonomous Okrug (Yugra), Nadymskiy District, dwarf shrub-lichen spotty tundra, on clay spot, 66 ° 32 ' 25 " N, 73 ° 52 ' 9 " E, 06. VIII. 2019, E. D. Lapshina 385 E / 3 - 4 - 19 [KPABG 124450].	en	Konstantinova, N. A., Vilnet, A. A., Mamontov, Yu. S. (2023): The phylogenetic affinity, distribution and variability of Cryptocolea imbricata r. m. Schust. (Marchantiophyta). Arctoa 32 (2): 137-150, DOI: 10.15298/arctoa.32.11, URL: https://doi.org/10.15298/arctoa.32.11
03F387F0FFFFFFC1FF57F87EFD190098.taxon	etymology	Etymology. The name reflects the very small size of plants. The variety can be separated by the key below:	en	Konstantinova, N. A., Vilnet, A. A., Mamontov, Yu. S. (2023): The phylogenetic affinity, distribution and variability of Cryptocolea imbricata r. m. Schust. (Marchantiophyta). Arctoa 32 (2): 137-150, DOI: 10.15298/arctoa.32.11, URL: https://doi.org/10.15298/arctoa.32.11
03F387F0FFFFFFC1FF57F87EFD190098.taxon	biology_ecology	Ecology. The species occurs in wet calcareous sites both in lowland and mountain tundra belt. In the alpine belt of northern mountains the species grows on humuscovered or calcareous moist basic rocks, at base or in crevices of rocks where it occurs mostly mixed with oth- er bryophytes, but can sometimes form almost pure mats, as e. g. in the Kamchatka Peninsula and the mountains of South Siberia. Its associates are other not common liverworts like Gymnomitrion revolutum, Radula prolifera, Pseudotritomaria heterophylla, Frullania subarctica, or in moist calcareous sites in northern regions widespread and more or less common Calycularia laxa, Mesoptychia heterocolpos, M. sahlbergii, Scapania crassiretis, Trilophozia quinquedentata, Blepharostoma brevirete, etc. In “ difficult ” habitats such as spots in spotted dwarf shrub-lichen or moss lichen tundra with low moisture the species occurs as the small dwarf variety minima. In such sites the described variety occurs as isolated scattered shoots among other bryophytes, e. g., Tritomaria scitula, Anthelia juratzkana, and Jungermannia sp.	en	Konstantinova, N. A., Vilnet, A. A., Mamontov, Yu. S. (2023): The phylogenetic affinity, distribution and variability of Cryptocolea imbricata r. m. Schust. (Marchantiophyta). Arctoa 32 (2): 137-150, DOI: 10.15298/arctoa.32.11, URL: https://doi.org/10.15298/arctoa.32.11
03F387F0FFFFFFC1FF57F87EFD190098.taxon	distribution	Distribution. Cryptocolea imbricata was described by Schuster (1969) from an isolated location in Minnesota and then recorded by him from one station in Michigan and many localities in Ellesmere Island and Greenland (l. c.). The species was later found in Alaska (Steere and Inoue, 1978), Northwest Territories of Canada (Damsholt, 2007), a single locality in Europe (northern Sweden, Schuster & Mårtensson, 1978), one in Svalbard (Frisvoll & Elvebakk, 1996) and many localities in Asia, including its northern part (Konstantinova et al., 2009, Schuster & Konstantinova, 1996; Fedosov et al., 2015; Fedosov et al., 2020; Konstantinova et al., 2023), mountains of Yakutia (Sofronova, 2018), South Siberia (Mamontov, 2013), and Far East of Russia (Bakalin, 2010, 2015, Bakalin et al., 2021) (Fig. 6). The species occurs from almost sea level in the tundra zone, e. g. in the Yamal Peninsula (Potemkin, 1993) up to 2000 m alt in the Kodar Range and Tardoki-Yani Mountains (South Siberia, Bakalin, 2015) or Kamchatka Peninsula (see specimens examined). Thus, Cryptocolea imbricata can be characterized as an arctomontane, almost circumpolar species.	en	Konstantinova, N. A., Vilnet, A. A., Mamontov, Yu. S. (2023): The phylogenetic affinity, distribution and variability of Cryptocolea imbricata r. m. Schust. (Marchantiophyta). Arctoa 32 (2): 137-150, DOI: 10.15298/arctoa.32.11, URL: https://doi.org/10.15298/arctoa.32.11
