taxonID	type	description	language	source
03F187F8FFC6FFB0FC23F9368463FE7F.taxon	discussion	Remarks. Both Andrewesia apicalis (Chaudoir, 1872) and Andrewesia obesa (Andrewes, 1923) were described from Singapore. Andrewes (1923) was seemingly not aware of A. apicalis when he described A. obesa from Singapore. Liebke (1938) felt that only one species might be involved: “ I have not seen the type, but from Chaudoir’s very detailed description it appears that it is at least closely related to A. obesa, if not the same species ”. I have checked the type of A. apicalis in MNHN, and can confirm that it is the same as the type of A. obesa in NHML (an image of the cotype of the latter is available from the Biodiversity of Singapore archive: https: // singapore. biodiversity. online /).	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC5FFB0FF09FE76879EFBF8.taxon	discussion	Remarks. Actenoncus can be distinguished from other Orthogoniine genera by these characters: the eyes are very large and very prominent, the sutural angle on the elytral apex bears a small but evident denticle, and the elytral intervals are without setiferous pore (in other genera, at least the 3 rd interval has setiferous pores). The absence of a median tooth on the right mandible is no longer considered a diagnostic character of this genus (Tian & Deuve, 2009). Orthogonius solidicornis should be transferred to Actenoncus according to the revised definition of the generic characters of Actenoncus, but this species was somehow overlooked in both the 2009 revision of Actenoncus (Tian & Deuve, 2009), as well as the catalogue of the Oriental Orthogonini (Tian & Deuve, 2006).	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC5FFB0FE87FBB184F8FA06.taxon	discussion	Remarks. Uloma gebieni Schawaller, 2000 is preoccupied by Uloma gebieni Kaszab, 1939. The proposed replacement name for Uloma gebieni Schawaller, 2000 is Uloma grimmi, new name.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC5FFB0FE87FBB184F8FA06.taxon	etymology	Etymology. Dedicated to the late Dr. Roland Grimm, specialist of Palaearctic, Oriental and Afrotropical Tenebrionidae.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC5FFB6FF77F9D8849FF92C.taxon	description	(Figs. 1 A – B, 2 A – C)	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC5FFB6FF77F9D8849FF92C.taxon	materials_examined	Material examined. Derosphaerus metallescens: 1 ♀ (HT): Singapore / TYPE [red label printed] / Encyalesthus metallescens Fairm. 1893. Singapore / MUSEUM PARIS, Collection Léon Fairmaire) [MNHN]; 1 ♀ (PT): Encyalesthus metallescens n. sp., Sumatra / [NHMB, collection Frey]; 1 ♂: Sumatra, Medan / Encyalesthus metallescens F. / [NHMB, collection Frey]. Derosphaerus viriditinctus: 2 ♀♀: Borneo / cum typo comparatum / Encyalesthus viriditinctus Fairmaire / P. ARDOIN DET. 1973. / Museum Paris Coll. P. Ardoin 1978 / [MNHN]; 1 ♀: Borneo / Plesiotype! No. 291 [red label printed] / Encyalesthus viriditinctus Frm. / [NHMB, collection Frey].	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC5FFB6FF77F9D8849FF92C.taxon	discussion	Discussion. Fairmaire (1882) described Encyalesthus (= Derosphaerus) viriditinctus, based on a single specimen from Palembang, Sumatra. Sex was not mentioned but it is female. Gebien (1914) provided additional descriptive notes on E. viriditinctus, based on one female specimen from Borneo in his possession (labelled as “ plesiotype ”). Specifically, he quantified the width of the frons (not reaching the transverse diameter of an eye), noted as important diagnostic features the long legs (elongate meso- and metatarsi) and the presence of a distinct anal impression which finely inflects the apical edge of the last ventrite. Unfortunately, Fairmaire’s holotype in MNHN could not be located. Instead, I have examined Gebien’s specimen in the Frey’s collection, labelled as “ plesiotype ” (Fig. 1 A), and two specimens of D. viriditinctus from Ardoin’s collection in MNHN, labelled as “ cum typo comparatum ” and identified as females (Fig. 1 B). Fairmaire (1893 a) described Encyalesthus metallescens based on two specimens from Singapore (HT, MNHN) and Sumatra (PT, NHMB), without mentioning the sex. However, it seems clear that the specimens are both female as Fairmaire did not note the distinctive dilated metatibia of the male E. metallescens in his description. Gebien (1914) did not describe E. metallescens directly but remarked that E. hauschildi can be distinguished from E. metallescens by “ … ungefurchtes Prosternum, einfache Hintertibien ... ”; which means that E. metallescens has prosternum furrowed and metatibiae dilated. While Gebien did not specifically say that he had a male specimen of E. metallescens, the tibial dilation is clearly a male feature. I follow Gebien’s concept of the male as it matches very well with the female, other than sexual differences that are typical for this genus. Gebien did not give any details of the male specimen on which he based his description. I found in Frey’s collection only one male specimen from Medan, Sumatra, and assume that that was the specimen from Gebien. Figs. 2 A and 2 B are respectively images of the holotype and paratype (both female) of Derosphaerus metallescens, whereas Fig. 2 C is an image of a male D. metallescens featured in Schawaller (2011) (in which it has been synonymised to D. viriditinctus and named as such). Kaszab (1987) provided a checklist of the species in Derosphaerus but D. metallescens is missing in the species list. Schawaller (2011) regarded Derosphaerus metallescens as a junior synonym of Derosphaerus viriditinctus. I note that Schawaller (2011) could not reexamine the types of both taxa, so this synonymy would be open to doubt with some justification. I propose to resurrect the former as a good species based on examination of the type materials and by returning to the descriptions of Fairmaire and Gebien. The view that D. metallescens and D. viriditinctus are unlikely to be synonymous is supported by the following reasons: For D. metallescens and D. viriditinctus to be synonymous, one has to disregard quite significant differences in the female holotypes of these two species. These differences include the more obvious ones like the frons width (difference in IE / TD between the two species more than two times), leg length (specifically meso- and metatarsi), and presence / absence of anal impression in the last ventrite. Subtler differences include longitudinal impression on pronotum, shape of pronotum, and convexity of elytral interval. For instance, compared to D. viriditinctus, D. metallescens typically has a bulkier pronotum relative to elytra (e. g., see PB / EW in Table 1). Admittedly, there is quite a bit of pronotal shape variation across the wide distribution range of D. metallescens. As a result of examining the types mentioned above, it is believed that the derived characters possessed by these two species are not common and that they are not homologous. One also has to consider what was regarded by Gebien as the male D. metallescens; this seems to be a very reasonable male choice for the female holotype, based chiefly on the pronotum shape, its longitudinal impression, IE / TD, etc. If, however, D. metallescens and D. viriditinctus were to be considered synonymous, this would result in the following rather atypical sexual dimorphism for a coleopteran: the frons of the male (IE / TD: 1.5 in the male D. metallescens) is much broader than that in the female (IE / TD: 0.6 – 0.7 in the female D. viriditinctus). Table 1 summarises some of the salient measurements that reflect the difference between the two species D. viriditinctus and D. metallescens. Remarks. Additional materials of Derosphaerus metallescens were also studied in MNHN and NHMB. There is considerable variation in the body size, pronotum shape, degree of shine on dorsum. While the body length shows a considerable variation of almost up to 30 %, most of the pronotum and elytra measurements conform to those in Table 1. However, there are a small number of specimens in which the pronotum is less bulky (PB / EW as low as 0.60) and also more transverse (PW / PL as high as 1.22). I am not sure if the whole series is conspecific or not, although it is difficult to separate them into different morphospecies.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	description	(Figs. 3 A- E)	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	materials_examined	Type specimen. Holotype: Male, Singapore: MacRitchie, Jelutong tower, coll. L. F. Cheong, 27 November 2019.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	description	Measurements. Male (n = 1): BL: 9.6 mm; BW: 3.7 mm; IE / TD: 2.23; PW / PL: 1.39; PW / FW 1.84; EL / EW: 1.52. Description. Colour dark reddish brown; for body shape see Fig. 3 A. Head: Clypeus without an elevated transverse ridge and without horns, truncate at apex, minutely punctate; frontoclypeal suture finely incised and distinct laterally, obscure in middle; frons with a broad Y-shaped depression in middle, surface with large and irregular punctures; genae produced laterad. Antennae (Fig. 3 B) reaching before middle of pronotum; middle antennomeres not produced medially nor pointed medially on distal margin, the latter sharp; antennomeres V to X gradually widening; VII to X extremely transverse; XI semicircular. Mentum (Fig. 3 C) rounded at apex, with basolateral excavations 0.5 times the length of mentum and separated at base, bearing a weak medial impression, and with long arcuate stretch of pubescence on both sides of the median impression. Ligula with a few sparse setae. Pronotum roundly quadrate, convex, widest at middle, as wide as elytra at base; anterior impression broad and deep, with a pair of low protuberances on posterior margin of impression; surface with small and dense punctures, punctation slightly coarser and denser in the anterior impression, surface between punctures with microscopic punctures; anterior margin emarginate, narrowly beaded; lateral margin arcuate, narrowly beaded, with a distinct sulcus along the margin; basal margin unbeaded. Prosternum longitudinally and strongly elevated in middle, coarsely punctate at both sides of the elevation which is Cheong: Nomenclature and taxonomy of Carabidae and Tenebrionidae finely punctate; prosternal process gradually dilated posteriad, adunc in apical half, distinctly sulcate along lateral margins, moderately punctate. Scutellum subtriangular, with minute punctures in basal half. Elytra subparallel-sided, widest at apical third; striate-punctate, punctures small but somewhat broader than the corresponding striae; elytral intervals convex, with microscopic punctures. Mesoventrite coarsely and densely punctate, with V-shaped ridge nearly vertical. Metaventrite convex, flattened and microscopically punctate in middle, coarsely punctate laterally. Last abdominal ventrite unbeaded. Protibia with a distinct longitudinal row of granules on dorsal side (Fig. 3 D). Aedeagus (Fig. 3 E) with parameres strongly convergent in middle, narrow and beak-shaped at apices. Female unknown.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	distribution	Distribution. Singapore.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	diagnosis	Diagnosis. Among the group of species with a distinct longitudinal row of granules on the dorsal side of protibia in males, Uloma schawalleri, new species, is most similar to U. nyctelia Ando in Ando et al., 2017 from Borneo and U. spinipes Kaszab, 1975 from Bhutan and Nepal, with their mentums all having distinct basolateral impressions and arcuate fields of pubescence on the surface. U. nyctelia differs from U. schawalleri, new species, as follows: (1) male mentum sharply truncate at apex, with basolateral excavations connected at base; (2) aedeagus with parameres less strongly convergent in middle, apices less slender; (3) distal five protibial teeth equally large. U. spinipes differs from U. schawalleri, new species, as follows: (1) male mentum more rounded in shape, with a very long and narrow field of pubescence placed just within the lateral margin; (2) parameres with a long narrow tip.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	discussion	Remarks. Other Uloma species in this group can be readily separated by the shape and structure of the mentum in males. Specifically, with respect to the surface of mentum in males, U. masumotoi Schawaller, 2000 (Sumatra, Borneo), U. ichoi Nakane, 1956 (Japan), and U. sauteri Kaszab, 1941 (Taiwan) are without distinct basolateral excavations; U. palopoensis Merkl & Ando, 2018 (Sulawesi), U. pelengensis Merkl & Ando, 2018 (Sulawesi), and U. visca Schawaller, 1994 (Philippines) are without arcuate fields of pubescence. U. sarawakensis Ando in Ando et al., 2017 (Borneo) has the male mentum distinctly raised in T-shape and sinuate at apex. Differences in the shapes of the middle antennomeres and aedeagus can be used to further aid separating U. schawalleri, new species, from these species. For instance, U. masumotoi and U. palopoensis have some of the middle antennomeres pointed medially on the distal margin. U. ichoi, U. sauteri and U. visca have the tip of parameres truncate. Like various other species in this group (e. g. U. nyctelia, U. palopoensis, and U. sarawakensis), the female of U. schawalleri, new species, is unknown. Based on other Uloma species in which the females are known, the anterior impression on pronotum and the row of granules on protibia should be absent in the females of U. schawalleri, new species. The presence of dense pubescence on mentum might also be a male-specific character. The diagnosis above is partly based upon my examination of the rich type materials of Sumatran and Bornean Uloma species in SMNS, upon which the revision of Schawaller (2000) is based. Collecting circumstances. Recorded at light traps.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC3FFB4FEA1F8468361FA62.taxon	etymology	Etymology. The specific name is dedicated to Dr. Wolfgang Schawaller, well-known specialist in Tenebrionidae.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	description	(Figs. 4 A- E)	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	materials_examined	Type specimens. Holotype: Male, Singapore: Chestnut, coll. L. F. Cheong & Y. W. Cheong, 13 May 2014. Paratypes: 1 male, Singapore: Bt. Timah Main Road, coll. L. F. Cheong, 21 September 2021. 1 male, Singapore: Chestnut, coll. L. F. Cheong, 30 November 2020; 1 female, ditto. 1 female, ditto, 11 November 2019.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	description	Measurements. Male (n = 3): BL: 19.5 - 22.6 mm; BW: 7.4 - 9.1 mm; PW / PL: 1.22 - 1.29; EL / EW: 1.56 - 1.68. Female (n = 2): BL: 19.2 - 19.5 mm; BW: 7.2 - 7.4 mm; PW / PL: 1.25 - 1.29; EL / EW: 1.72 - 1.76. Description. Male. Elongate, black, matt. Head with punctures fine, coarser near frontoclypeal suture; clypeus shallowly emarginate at anterior margin; frontoclypeal suture finely incised but not forming deep groove; ocular ridges distinctly elevated, reaching more or less the rear edges of eyes. Antennae short, reaching middle of pronotum. Mentum obtrapezoidal, with pronounced, dense, erect, yellow moustache. Pronotum widest at middle; lateral margins reflexed, slightly sinuate before base, with beads gradually becoming narrow to very narrow forward until just before anterior corners, thence the margins weakly emarginate (Fig. 4 B); all margins distinctly beaded; apical margin interrupted in middle; disk densely and finely punctate, with basally keeled median line, interspaces between punctures weakly microreticulate. Propleura nearly smooth, shallowly rugulose; prosternal process broad, flat, bisulcate, basal portion with moderately dense golden hairs, elsewhere with sparsely scattered short golden hairs. Elytra striate-punctate; intervals convex, slightly shagreened, with microscopic punctures. Metaventrite nearly smooth, with fine punctures or microgranules each of which bears very short hair; disc covered by dense golden pubescence in the middle, with three clusters of elongate granules on each side of midline in posterior half (Fig. 4 C). Abdominal ventrites with fine minute punctures or microgranules each of which bears very short hair; 5 th ventrite furrowed apically. Protibia strongly bent at apical fourth, densely pubescent in apical sixth (Fig. 4 D), with a blunt broadening in the basal region. Aedeagus (Fig. 4 E) slightly curved; parameres gradually narrowed apicad from base, 0.75 times as long as basal piece. Female. Pubescence on prosternal process and metaventrite shorter than male; metaventrite without clusters of elongate granules; protibia not strongly bent at apex, with pubescence there shorter and less dense than male.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	distribution	Distribution. Singapore.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	diagnosis	Diagnosis. The widely distributed Promethis excisa (Gebien, 1914) (recorded from Malaysia, Singapore, Sumatra, Borneo, Sulawesi, Philippines) shares with P. tuberculata, new species, the trait of having their broadly beaded lateral margins of pronotum becoming very finely beaded before the anterior corners so that a shallow emargination is observed on the lateral margins. P. excisa differs from P. tuberculata, new species, as follows: (1) anterior corners of pronotum more distinctly emarginate; (2) frontoclypeal suture with a deep groove; (3) male protibia sharply edged on outer margin near apex, with two blunt broadenings basally; (4) male mesotibia suddenly widened just before apex; (5) metaventrite glabrous in females (not described for males); (6) aedeagus more strongly curved; parameres almost parallel-sided at base, only narrowing in apical third.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	discussion	Remarks. P. tuberculata, new species, belongs to the P. excisa group (sensu Kaszab, 1988 a) on account of its mentum with pronounced, dense, yellow moustache, and 5 th abdominal ventrite furrowed apically, among other characters. Among the extant four species of this group, P. plicifrons (Gebien, 1918) from Malaysia and Singapore is most readily identified by its strikingly short antennae, the presence of a granule on the metaventrite between the mesocoxae in males, and the strikingly flattened mesotibiae in males. The remaining species in this group, together with the new species P. tuberculata, new species, all reach couplet 254 in the key of Kaszab (1988 a) due to the absence of the aforementioned characters of P. plicifrons. P. tuberculata, new species, and P. excisa are then separated from the rest by the distinctive emargination on the lateral margins of pronotum just before the anterior corners. In other words, P. tuberculata, new species, would be inserted into the key at couplet 254, which has to be altered as follows: 254 (255) Lateral margins of pronotum with bead narrowed to very narrowed forward and interrupted far before the anterior corners, before which the margins are more or less distinctly emarginate. 254 a (254 b) Frontoclypeal suture bearing a deep groove; male protibia strongly curved at about apical third, with two blunt broadenings basally and a sharp edge at the outer margin near apex; male mesotibia suddenly widened just before apex; metaventrite glabrous in females ..... P. excisa (Gebien, 1914) 254 b (254 a) Frontoclypeal suture finely incised but without deep groove; male protibia strongly bent at apical fourth, with a blunt broadening basally; male mesotibiae not suddenly widened near apical portion; metaventrite with golden pubescence, males with three clusters of elongate granules on each side of midline in posterior half .............................. P. tuberculata, new species 255 (254) Lateral margin of pronotum with bead not interrupted at the anterior corner, beading continuously into that of anterior margin. The comparative diagnosis above is partly based upon my examination of the type material of P. excisa in NHMB. The holotype of P. plicifrons cannot be located; I have instead studied the photographs of a HNHM specimen identified by Kaszab as P. plicifrons (photographs kindly provided to me by György Makranczy of HNHM). Collecting circumstances. Encountered in rotting wood.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
03F187F8FFC1FFB5FC09FA648314FAF6.taxon	etymology	Etymology. The specific name refers to the coarse granules that occur on the metaventrite of the males.	en	Cheong, Loong Fah (2025): On nomenclature and taxonomy of some Carabidae and Tenebrionidae from Singapore, with descriptions of new species of the genera Uloma Dejean, 1821 and Promethis Pascoe, 1869. Raffles Bulletin of Zoology 73: 233-242, DOI: 10.26107/RBZ-2025-0017
