taxonID	type	description	language	source
03868B5FFF92FF89FF399EA74E26BAD5.taxon	diagnosis	Amended diagnosis (following Fujimoto 2015 with additions in bold)	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF89FF399EA74E26BAD5.taxon	description	Halechiniscidae with cylindrical body; primary clava and lateral cirrus arise from common cirrophore; primary clava longer than lateral cirrus; secondary clava undeveloped; leg IV sense organ as papilla; legs terminate in digits without peduncles, proximal pads, pretarsi or wrinkles but may have folds; internal digits longer than external digits; each digit terminates in sheathed, small, crescent-shaped claw with minute calcar; pair of ventrally opening seminal receptacles each with slender, sinuous duct terminating in spherical vesicle.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF89FF39980F48F3BC49.taxon	diagnosis	Amended diagnosis (following Fujimoto 2015 with additions in bold)	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF89FF39980F48F3BC49.taxon	description	Quisarctinae with cylindrical body; primary clava and lateral cirrus arise from common cirrophore; primary clava longer than lateral cirrus; secondary clava undeveloped; eyes may be present; stylet supports may be present; leg IV sense organ as elongate papilla with distal spine; legs terminate in digits without peduncles, proximal pads, pretarsi or wrinkles but may have folds; internal digits longer than external digits; each digit terminates in sheathed, small, crescent-shaped claw with minute calcar; pair of ventrally opening seminal receptacles each with slender, sinuous duct terminating in spherical vesicle.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF8DFF3999B34E5EBEAB.taxon	description	(Figs 2 – 8 A, B, Table 2) urn: lsid: zoobank. org: act: 812 F 34 F 1 - 7 EEF- 4 BD 9 - B 9 CB-A 62021 A 42 BB 7	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF8DFF3999B34E5EBEAB.taxon	diagnosis	Diagnosis Quisarctus with sexual dimorphism in the shape of cirri E: female has flagellum-shaped cirri, male has lance-shaped cirri. Body cuticle with punctation and dorsal transverse folds. Buccal apparatus consists of buccal tube, stylets and a pharyngeal bulb with three placoids. Legs I – III sensory organs as tripartite spines. Leg IV sensory organ developed as an elongated papilla with tubular tip. Digits may have proximal folds. Seminal receptacles open ventrally.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF8DFF3999B34E5EBEAB.taxon	materials_examined	Type material The holotype (Figs 2 A, 3) is an adult female (specimen ID M 150 - 123 - 1 - SG _ 2 v 2, SMF 1313) with seminal receptacles, and rosette gonopore. The allotype (Figs 2 B, 4) is an adult male (specimen ID M 150 - 161 - 1 - 2 _ 2 v 48, SMF 1323) with one seminal vesicle filled with spermatozoa and oval gonopore. Sixteen further specimens are paratypes (Figs 5 – 7). Two of them, M 150 - 226 - 1 - 6 _ 5 v 6, SMF 1384 (female, Fig. 6) and M 150 - 226 - 1 - 6 _ 3 v 6, SMF 1382 (adult male exuvium, Fig. 7) were prepared for the SEM investigation. The other 14 specimens are three males, four females, two juveniles and five specimens with undetermined sex. The whole type series is deposited at the Senckenberg Natural History Museum as part of the Tardigrada collection under inventory numbers as specified in Table 2. Type locality Princess Alice Bank, Azores; holotype: 37 ° 59.968 ' N, 029 ° 17.054 ' W; 151 m depth, bioclastic and volcanogenic sediment (Fig. 1 C), collected on 5 th September 2018; allotype: 38 ° 01.061 ' N, 029 ° 19.691 ' W; 306 m depth, bioclastic sediment; collected on 6 th September 2018.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF8DFF3999B34E5EBEAB.taxon	etymology	Etymology The name armatus (from Latin; means armed) refers to lance-shaped cirrus E in males and dorsal transverse folds forming a pattern of dorsal armour plating in females.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF8DFF3999B34E5EBEAB.taxon	description	Description of holotype (SMF 1313) (Figs 2 A, 3, Table 2) Adult female tardigrade with cylindrical body (95 µm long × 30.8 µm wide) and loose, punctated cuticle. A number of transverse folds with less punctated margins on the dorsal side (Fig. 3 A). The punctation of the dorsal side is more prominent; the ventral surface, distal portions of legs, head cuticle and the margins of dorsal transverse folds have less marked or no cuticle punctation. Long primary clavae have a mid-portion of folded cuticle (Fig. 3 A) and an apical pore. Primary clava and the cirrus A arise from a common cirrophore. Cirri A situated antero-dorsally to primary clavae. Cephalic cirri present as paired internal and external cirri plus an unpaired median cirrus; all cirri, including cirri A, consist of three parts: scapus, tubular portion and short flagellum. The internal cirri are dorsolateral; the external cirri are ventrolateral. Secondary clavae are absent. Lipoid eyespots present about at the position of the median cirrus (Fig. 3 C). Mouth cone is retracted (Fig. 3 C). Pharyngeal bulb is macerated. Leg I – III sensory organs present as tripartite spines with scapus, tubular portion and short flagellum (Fig. 3 D, E). Sensory organ of the leg IV as an elongated papilla with a short distal tubular tip (Figs 2 A, 3 E, 8 A). Cirri E are flagellum-shaped with a proximal portion, distal portion and spine, arising from a cirrophore (Figs 2, 3 A). All legs have two internal digits longer than two external digits. Each digit terminates in a simple crescent-shaped claw with calcar. Leg IV digits are larger than legs I – III digits. Proximal part of digits sometimes has folds (Fig. 2 A). Rosette-shaped female gonopore present (Fig. 3 G). A pair of seminal receptacle openings ventrally at the bases of legs IV is connected to slender, slightly S-shaped ducts and terminates as vesicles (Figs 2 A, 3 F). Anus is 4.7 µm posterior to gonopore and appears as a zigzag cuticle fold (Fig. 3 G).	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF92FF8DFF3999B34E5EBEAB.taxon	discussion	Remarks on the allotype (SMF 1323) (Figs 2 B, 4, Table 2) Male with cylindrical body (118.4 µm long × 29.0 µm wide) and loose, thin, punctated cuticle, which forms transversal folds on the dorsal side. Long primary clavae with folded cuticle (Fig. 4 A, D). Secondary clavae absent. Lipoid eyespots present (Fig. 4 D). Mouth cone is rather narrow (diameter: 3.8 µm) and directed anteriorly (Fig. 4 A). About 25 µm long stylets without supports are present in the head section and inside the mouth cone. Three placoids (Fig. 4 E) are visible in the oval pharyngeal bulb (10 × 8.5 µm). The pharyngeal bulb is followed by a globular oesophagus (Fig. 4 A). Leg I – III sensory organs present as tripartite spines with scapus, tubular portion and short flagellum (Fig. 4 F – H). Sensory organ of leg IV is an elongated papilla with a short distal tubular tip (Figs 2 B, 4 I, 8 B). Cirri E arise from a cirrophore and have a lance-shaped distal portion (Fig. 4 J). All legs have two internal digits longer than two external digits (Fig. 4 K). Each digit terminates in a simple crescent-shaped claw with calcar. The claw sheath completely covers the claw. Leg IV digits are larger than legs I – III digits. Proximal part of digits sometimes has folds. One seminal vesicle with spermatozoa present (Fig. 4 L). Oval male gonopore present a short distance anterior to leg IV insertions (Fig. 4 M). Anus slightly posterior to gonopore and appears as a zigzag cuticle fold (Fig. 4 M). Remarks on paratypes (Figs 5 – 7, Table 2) A number of 16 paratype specimens were either investigated with light microscopy, or with SEM in order to study further structural details and character variability. All paratypes exhibit morphology coherent with that of the holotype and the allotype (Fig. 5). Males are 96 – 123 µm in body length (n = 2) and the primary clavae are 27 – 30 µm (n = 2) long. Females are 92 – 118 µm in body length (n = 3) and the primary clavae are 32 – 35 µm (n = 3) long. Juvenile specimens are 75 – 93 µm in body length (n = 2) and the primary clavae are 24 µm (n = 1) long. The primary clavae of the investigated females appeared significantly longer than that of the males (Mann-Whitney test, z = 2.653, p = 0.002). The juveniles have cirri E in the shape of a flagellum, which may indicate either the potential late development of the lance-shaped cirri E specific to males, or that the juvenile specimens are immature females. During SEM investigation, we were able to better examine and confirm the cuticular structures, especially dorsal transverse folds forming a pattern of segmentation or dorsal armour plating (Fig. 6 B) and the epicuticular pillars (Fig. 7). Furthermore, SEM imaging helped to investigate the differences in cirri E between sexes (Figs 6 G, 7 D, E), and the fine structure of the sensory organs and digits folds (Figs 6 F, 7 C). The sexual dimorphism in Quisarctus armatus sp. nov. is observable in the difference between the length of primary clavae; the shape of leg IV sensory organ, and the shape of cirri E: males have spine-like cirri with a lance-shaped distal end, while females have flagellum-shaped tripartite cirri. Males have oval gonopores situated close to the anus, and females have rosette-shaped gonopores opened further from the anus.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF9AFF81FF3999914F73BE1B.taxon	description	Fig. 9; N = 43; 295 – 306 m bsl	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF9AFF81FF3999914F73BE1B.taxon	discussion	Remarks Phenotypically remarkable tardigrades (115 – 188.5 µm in body length, n = 12) with segmented body and three pairs of lateral body processes (Fig. 9 A). All the specimens matched the original description (Kristensen et al. 2015). Mid-dorsal spines sometimes were broken, but we confirmed the presence of ventral body plates, always one pair of ventral cervical spines (Fig. 9 B), the presence of accessory spine on each claw (Fig. 9 C), and the absence of eyes. Measurements of selected specimens presented in Table 3. As we mentioned earlier, this species has been described from the Condor seamount, Azores (Kristensen et al. 2015), and has not been reported yet outside of its type locality until this current record from the PAB.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF9DFF86FF399FF54BCDBC94.taxon	description	Fig. 10; N = 9; 295 – 306 m bsl	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF9DFF86FF399FF54BCDBC94.taxon	discussion	Remarks Robust tardigrades (78 – 120 µm in body length, n = 6) with distinct epicuticular pillars (Fig. 10 A). Primary clavae are 12 – 16 µm long (n = 6). The main trait for this species is the so-called fan-shaped tarsus, with all the digits attached and aligned in one line (Fig. 10 C). Claws with calcar (Fig. 10 C). Following the original description (Renaud-Mornant 1984), we searched and observed the unpunctated zone close to the mouth cone — presumably secondary clavae or their rudiments (Fig. 10 B). This species has a cosmopolitan distribution in the Mediterranean sea, Atlantic Ocean and Indian Ocean (Kaczmarek et al. 2015).	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
03868B5FFF9DFF86FF3999CD48D6BFF5.taxon	discussion	Remarks We investigated five deteriorated tardigrades, probably juveniles (63 – 99 µm in body length, n = 4), which could be assigned to the family Halechiniscidae. However, further determination to genus or species level is not possible because of the lack of morphological resolution.	en	Trokhymchuk, Roman Ruslanovych, Arbizu, Pedro Martínez, Kristensen, Reinhardt Møbjerg, Kieneke, Alexander (2025): Marine tardigrades of Princess Alice Bank (Azores, Northeast Atlantic) with a description of a new species of Quisarctus Fujimoto, 2015. Zootaxa 5711 (1): 125-139, DOI: 10.11646/zootaxa.5711.1.6, URL: https://doi.org/10.11646/zootaxa.5711.1.6
