identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039787890F6E6123FD0BA5F2FEC0629C.text	039787890F6E6123FD0BA5F2FEC0629C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anticyphon argentinensis Ruta 2018	<div><p>Anticyphon argentinensis Ruta, new species</p><p>(Figs. 1–2, 7–12)</p><p>Zoobank.org/ urn:lsid:zoobank.org:act: 0FDB4206-157C-46DD-8B0E-B66DC1E6B509</p><p>Type Material. Holotype, male (ANIC):“ Tucumán, R. A. \ 9 – 1949 \ Wygodzinsky col. \ Tafi ad Valle \ 2000m ” [printed &amp; handwritten].</p><p>Etymology. The species is named for terra typica, Argentina.</p><p>Diagnosis. Anticyphon argentinensis resembles Anticyphon ecuadorensis Ruta, A. oyonensis, and Anticyphon peruvianus Ruta in size, coloration, and granulate punctation on the pronotal disc. Anticyphon argentinensis has a more transversely rectangular pronotum with less rounded anterolateral pronotal angles. Definitive identification is possible only on the basis of the male genitalia: trigonium without lateral lobes, as long as parameroids, parameres wide, each with small, triangular denticle in basal portion.</p><p>Description. Male. Body oblong oval, moderately large, slightly flattened, clothed with yellowish procumbent setae. Coloration of dorsum yellowish brown, antennae and legs yellowish. Head small, 1.6X wider than interocular space, covered with minute granulate punctures; eyes relatively big, protuberant; frons with 2 pairs of small tubercles. Antennae filiform, antennomere 2 shortest [antennomeres 9–11 missing in the holotype]. Pronotum small, transversely rectangular, sides straight, widest at posterior angles, anterolateral angles square, subtly produced; disc moderately convex. Pronotal punctation granulate, stronger on sides than on central portion. Pronotum with subtly marked pits along basal margin. Each elytron with 3 subtly marked longitudinal carinae. Elytral punctation irregular, relatively sparse; punctures shallow, separated by ca. 1.0 diameter. Penis (Fig. 7) moderately large (L = 1.30 mm, W = 0.34 mm), parameroids long, straight, wide, with subparallel sides, pointed at apices; trigonium subtriangular, as long as parameroids, without lateral lobes, pala longer than parameroids, moderately wide; tegmen (Fig. 8) moderately large (L = 0.98 mm, W = 0.51 mm), parameres wide in lateral view, covered with microsculpture in apical portion, basal portion of each paramere with small subtriangular denticle; sternite VIII (Fig. 9) small (L = 0.35 mm, W = 0.47 mm), V-shaped, with sparse setae in apical portions; sternite IX (Fig. 10) relatively small (L = 0.48 mm, W = 0.43 mm), suboval, lightly sclerotized in basal portion, apical portion setose; tergite VIII (Fig. 11) (L = 0.77 mm, W = 0.65 mm) with subrectangular apical plate, apical half covered with microsetae, apical margin arcuate, with row of dense, short setae intermixed with sparse, longer ones, basal portion of apical plate darker and more strongly sclerotized than remaining portion of plate, apodemes much shorter than apical plate; tergite IX (Fig. 12) (L = 0.70 mm, W = 0.52 mm) distinctly narrower than tergite VIII, apical plate membranous, margin densely covered with setae.</p><p>Measurements and Ratios. Male (n = 1): TL = 5.30 mm; PL = 0.65 mm; PW = 1.80 mm; EL = 4.70 mm; EW = 3.00 mm; TL/EW = 1.80; PW/PL = 2.80; EL/EW = 1.60; EL/PL = 7.20.</p><p>Female. Unknown.</p><p>Biology. Unknown.</p><p>Distribution. Anticyphon argentinensis is known only from the type locality in northern Argentina (Fig. 23). This species extends the known geographical range of the genus about 2,000 km to the southeast.</p></div>	https://treatment.plazi.org/id/039787890F6E6123FD0BA5F2FEC0629C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruta, Rafał	Ruta, Rafał (2018): Two New Species of Anticyphon Ruta (Coleoptera: Scirtidae) from Argentina and Ecuador. The Coleopterists Bulletin 72 (4): 707-712, DOI: 10.1649/0010-065X-72.4.707, URL: http://dx.doi.org/10.1649/0010-065x-72.4.707
039787890F6C6125FD22A3C9FEFF610E.text	039787890F6C6125FD22A3C9FEFF610E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anticyphon lescheni Ruta 2018	<div><p>Anticyphon lescheni Ruta, new species</p><p>(Figs. 3–6, 13–22)</p><p>Zoobank.org/ urn:lsid:zoobank.org:act: 0ABF94AB-1A94-4DDB-B912-74F7E309DA92</p><p>Type Material. Holotype, male (SEMC): “ ECUADOR: Azuay \ Cajas 25 km NW Cuenca \ 3700 m, 7 Jan. 1992 \ C. Carlton, R. Leschen \ #127 ex: beating” [printed] . Paratypes, 8 females (SEMC), same data as holotype .</p><p>Etymology. The species is dedicated to one of the collectors, Richard Leschen, a leading expert in Coleoptera systematics.</p><p>Diagnosis. Anticyphon lescheni resembles A. ecuadorensis and Anticyphon paramoensis Ruta in size, shape, and coloration. The granulate punctation of the pronotal disc is subtler than in A. ecuadorensis and A. paramoensis . Accurate identification is possible only on the basis of the male genitalia: trigonium wide, with wide lateral lobes, distinctly shorter than parameroids, parameres bifid, with microsculptured membranous apical portion.</p><p>Description. Male. Body oblong oval, small, slightly flattened, clothed with brownish, suberect setae. Elytra brown, scutellary shield lighter, light brown, pronotum light brown with darker spots, antennae dark brown, legs yellowish. Head small, 1.6X wider than interocular space, covered with minute, granulate punctures; eyes relatively big, protuberant; frons with 2 depressions. Antennae filiform, antennomere 2 shortest, antennomere 3 only slightly longer than 2, narrowest. Pronotum small, subrectangular, sides subtly curved, widest at posterior angles, anterolateral angles rounded; disc convex. Pronotal punctation granulate, stronger on sides than on central portion. Pronotum without pits along basal margin. Elytral punctation irregular, relatively sparse; punctures shallow, separated by ca. 1.0–1.5 diameter. Penis (Figs. 13–14) moderately large (L = 1.00 mm, W = 0.26 mm), parameroids long, straight, slightly widened in apical portion, rounded at apices, small lobes armed with small denticles (visible in lateral view) present in basal portion; trigonium widely subtriangular, ventral margin covered with denticles, lateral lobes wide, curved ventrally, of same length as median portion of trigonium, distinctly shorter than parameroids, pala longer than parameroids, moderately wide; tegmen (Fig. 15) moderately large (L = 0.87 mm, W = 0.43 mm), parameres bifid, outer processes extended by membranous structure covered with microsculpture in apical portion; sternite VIII (Fig. 16) small (L = 0.28 mm, W = 0.42 mm), V-shaped, with sparse setae in apical portions; sternite IX (Fig. 17) relatively small (L = 0.34 mm, W = 0.40 mm), suboval, transverse, apical portion setose; tergite VIII (Fig. 18) (L = 0.55 mm, W = 0.59 mm) with transversely subrectangular apical plate, apical half covered with microsetae, apical margin arcuate, with row of dense, short setae basal portion of apical plate darkened, apodemes much shorter than apical plate; tergite IX (Fig. 19) (L = 0.52 mm, W = 0.41 mm) distinctly narrower than tergite VIII, apical plate membranous, margin densely covered with setae.</p><p>Female. Distinctly larger than male. Antennomere 3 distinctly longer than 2. Pronotum often brown with darker spots. Each elytron with 3 subtly marked longitudinal carinae. Sternite VIII elongate (L = 1.32 mm, W = 0.31 mm), with sinuate apodemes not fused basally, apex with sparse setae. Tergite VIII long (L = 1.94 mm, W = 0.40 mm); apical portion subtriangular, with sparse setation on apical margin; apodemes long, 3X longer than apical portion. Ovipositor (Fig. 20) long (L = ca. 3.0 mm); branchlets small; coxites (L = 0.53 mm) membranous, triangularly elongate, narrow, with parallel sides in apical half, covered with sparse, minute setae; styli with 2 tufts of setae at apex. Vulva (Fig. 21) membranous, with 5 sclerites forming large sclerotized structure: 2 small subrectangular sclerites (L = 0.14 mm, W = 0.08 mm); 2 large, elongated sclerites connected with membrane (single sclerite: L = 0.60 mm, W = 0.18 mm); and a subtriangular ventral sclerite (L = 0.20 mm, W = 0.12 mm); bursal sclerites (Fig. 22) consisting of large, lanceolate sclerite (L = 0.40 mm, W = 0.11 mm), smaller sclerite consisting of paired, rodlike sclerites covered with minute denticles (L = 0.14 mm, W = 0.07 mm) and a small, oval sclerite (L = 0.07 mm, W = 0.03 mm); posterior portion of bursella covered with oval, pentagonal and hexagonal cuticular structures; anterior portion of bursella covered with microtrichia.</p><p>Measurements and Ratios. Male (n = 1): TL = 3.75 mm; PL = 0.63 mm; PW = 1.4 mm; EL = 3.25 mm; EW = 2.25 mm; TL/EW = 1.70; PW/PL = 2.20; EL/EW = 1.40; EL/PL = 5.20. Females (n = 8): TL = 4.80–5.50 mm (mean = 5.1 mm); PL = 0.80–0.95 mm (mean = 0.86 mm); PW = 1.70–1.95 mm (mean = 1.8 mm); EL 4.00– 4.70 mm (mean = 4.4 mm); EW = 2.60–3.00 mm (mean = 2.8 mm); TL/EW = 1.74–1.96 (mean = 1.84); PW/PL = 2.00–2.25 (mean = 2.14); EL/EW = 1.48–1.67 (mean = 1.56); EL/PL 4.72–5.63 (mean = 5.10).</p><p>Biology. Unknown.</p><p>Distribution. Anticyphon lescheni is known only from the type locality in southern Ecuador (Fig. 23), not far from the type locality of A. ecuadorensis .</p></div>	https://treatment.plazi.org/id/039787890F6C6125FD22A3C9FEFF610E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruta, Rafał	Ruta, Rafał (2018): Two New Species of Anticyphon Ruta (Coleoptera: Scirtidae) from Argentina and Ecuador. The Coleopterists Bulletin 72 (4): 707-712, DOI: 10.1649/0010-065X-72.4.707, URL: http://dx.doi.org/10.1649/0010-065x-72.4.707
039787890F6A6126FD0AA587FB5865CE.text	039787890F6A6126FD0AA587FB5865CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anticyphon RUTA	<div><p>AN UPDATED IDENTIFICATION KEY TO THE SPECIES OF ANTICYPHON (modified from Ruta 2016)</p><p>Note. Females are known for two species only. Therefore, only males can be identified with the use of the following key.</p><p>1. Body larger (TL = 6.50–6.80 mm); pronotal disc with normal, not granulate punctation; lateral processes of trigonium absent; pala of penis with membranous dorsal process ....................... 2</p><p>REFERENCES CITED</p><p>Chaboo, C. S., and W. D. Shepard. 2015. Beetles (Coleoptera) of Peru: A survey of the families. Clambidae, Eucinetidae, Scirtidae (Scirtoidea). Journal of the Kansas Entomological Society 88 (2): 208–210.</p><p>Klausnitzer, B. 2012. Zur Kenntnis der Gattung Prionocyphon L. Redtenbacher 1858 sensu lato (Coleoptera, Scirtidae) aus Südamerika. Linzer biologische Beiträge 44 (2): 1177–1194.</p><p>Klausnitzer, B. 2013. Neue Arten der Gattung Prionocyphon L. Redtenbacher 1858 aus Südamerika (Coleoptera, Scirtidae). Linzer biologische Beiträge 45 (1): 775–787.</p><p>1 ʹ. Body smaller (TL = 3.80–5.80 mm); pronotal disc with granulate punctation; lateral processes of trigonium absent; pala of penis without dorsal process ........................... 3</p><p>Lawrence, J. F. 2001. A new genus of Valdivian Scirtidae (Coleoptera) with comments on Scirtoidea and the beetle suborders. Special Publication of the Japan Coleopterological Society 1: 351–361.</p><p>2. Trigonium distinctly shorter than parameroids ............... A. santanderensis Ruta</p><p>Libonatti, M. L. 2014. A revision of the genus Ora Clark, 1865 (Coleoptera: Scirtidae) in Argentina (part I) – descriptions of new species. Zootaxa 3884 (1): 27–44.</p><p>2 ʹ. Trigonium almost as long as parameroids .... .................................. A. davidsoni Ruta</p><p>3. Median process of trigonium distinctly shorter than parameroids; parameres bifid at base, with mesal processes ............................. 4</p><p>Libonatti, M. L. 2015. A revision of the genus Ora Clark, 1865 (Coleoptera: Scirtidae) in Argentina (part II) – redescriptions, updated distributions and a key to species. Zootaxa 3985 (1): 69–97.</p><p>3 ʹ. Median process of trigonium as long as parameroids; parameres simple .................... 5</p><p>Libonatti, M. L., and R. Ruta. 2013. Review of the Argentinean species of Pseudomicrocara Armstrong (Coleoptera: Scirtidae). Zootaxa 3718 (2): 137–157.</p><p>4. Parameres more than 3X longer than trigonium ........................ A. peruvianus Ruta</p><p>4 ʹ. Parameres less than 2X longer than trigonium .......... A. lescheni Ruta, new species</p><p>Ruta, R. 2011. Chilarboreus gen. nov., a new genus of Chilean Scirtidae (Coleoptera: Scirtoidea), with descriptions of three new species. Journal of Natural History 45: 1689–1713.</p><p>5. Parameroids subparallel in dorsal view .......... 6</p><p>5 ʹ. Parameroids curved in dorsal view .......... 7</p><p>6. Trigonium with lateral lobes, parameres strongly sclerotized, without denticle in basal portion ................... A. ecuadorensis Ruta</p><p>Ruta, R. 2016. Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation. Zootaxa 4175: 301–318.</p><p>6 ʹ. Trigonium without lateral lobes, parameres membranous, with small, subtriangular denticle in basal portion ............................... ........... A. argentinensis Ruta, new species</p><p>7. Body broadly oval, uniformly brownish; median process of trigonium straight; parameroids widely separated .............. A. oyonensis Ruta</p><p>7 ʹ. Body oblong, contrasting brown and black to totally black; median process of trigonium curved in lateral view; parameroids narrowly separated in apical portion ........................ .............................. A. paramoensis Ruta</p><p>ACKNOWLEDGMENTS</p><p>Adam Ślipiński and Cate Lemann are thanked for their hospitality during the author’ s stay in Canberra and for allowing study in the ANIC collection of Scirtidae . Zack Falin (University of Kansas) is thanked for sending SEMC specimens. Anonymous reviewers are thanked for their helpful suggestions.</p><p>(Received 27 March 2017; accepted 10 October 2018. Publication date 28 December 2018.)</p></div>	https://treatment.plazi.org/id/039787890F6A6126FD0AA587FB5865CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruta, Rafał	Ruta, Rafał (2018): Two New Species of Anticyphon Ruta (Coleoptera: Scirtidae) from Argentina and Ecuador. The Coleopterists Bulletin 72 (4): 707-712, DOI: 10.1649/0010-065X-72.4.707, URL: http://dx.doi.org/10.1649/0010-065x-72.4.707
