taxonID	type	description	language	source
039487E1FF970C19FF4254DDFBD7FDF8.taxon	type_taxon	type species designation: Tepperia sterculiae Lea, 1903); Alonso-Zarazaga & Lyal, 1999: 128 (cataloged; Cryptorhynchini: Cryptorhynchina; type species: “ NYD ”).	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF970C19FF4254DDFBD7FDF8.taxon	diagnosis	Diagnosis. This genus can be distinguished from other genera of Cryptorhynchina by a combination of the following characteristics: rostrum stout; frons wider than half width of an eye; basal margin of each elytron rounded near scutellar shield; legs short; each femur ventrally with a median tooth; meso- and metatibiae angulate basally along outer margins; mesoventral receptacle cavernous, projecting ventrally; and ventrite II nearly as long as III and IV combined (Lea 1903 & 1913; Marshall 1936).	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF970C19FF4254DDFBD7FDF8.taxon	distribution	Distribution. Australia; the Philippines, Taiwan, and Japan — new records.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF970C19FF4254DDFBD7FDF8.taxon	discussion	Remarks. Alonso-Zarazaga and Lyal (1999) noted the type species of Tepperia as “ NYD ” (not yet designated) in their catalog, but Marshall (1936: 195) had previously designated T. sterculiae as the type species, stating that “ T. sterculiae is now fixed as the type of the genus ”. The genus Evaniocis Lea, 1913, which comprises two Australian species and closely resembles Tepperia, was erected based on the “ coarsely faceted eyes and elytra not separately rounded at base ” (Lea 1913); however, the relationship between the two genera warrants attention since states of these characteristics seem to be variable within Tepperia. In the course of his study on New Zealand taxa of Cryptorhynchinae, Lyal (1993) indicated that the presence of a “ spiculum relictum ” can be regarded as a trait of generic importance. On the other hand, Lyal (1993) noted that the shape of the basal sclerite (= copulatory sclerites sensu Lyal 2010) is variable within the subfamily and is sometimes crucial for distinguishing genera. At that time, morphological characteristics of male terminalia had not been studied for Tepperia and its relatives. As described below, we observed structures of the male terminalia of three Tepperia species, T. sterculiae, T. palawanica and T. vagabunda, for the first time, and revealed that their male terminalia are similar in having a spiculum relictum of sternite VIII and a tubular copulatory sclerite in the endophallus. In order to assess the significance of structures of the male terminalia in delimiting the genus Tepperia, it is necessary to examine the character states in other congeners, as well as in members of related genera.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF940C18FF425380FB65FD1C.taxon	description	(Figs. 1 – 11)	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF940C18FF425380FB65FD1C.taxon	diagnosis	Diagnosis. This species is closely similar to T. major Lea, 1903 but can be distinguished from the latter by having the wider frons and pronotum depressed in apical part (Marshall 1936). Supplementary description. Male. Habitus, rostrum, meso- and metaventrites, and venter as in Figs. 1 – 4 & 6. Thorax. Ventral surface of elytra (Fig. 5) with submarginal ridges sublinear in ventral aspect, not arched anteriorly, with two asymmetric files on apico-internal part. Terminalia (Figs. 7 – 11). Tergite VII (Fig. 7) with two distorted rows of setiferous sclerites. Rectum with weakly sclerotized rectal ring. Endophallus (Fig. 8) with tubular copulatory sclerite. Tegminal ring (Fig. 9) with weakly sclerotized parameroid lobes, which are fused basally; diameter of tegminal ring longer than its apodeme. Sternite VIII (Fig. 10) composed of two narrowly separated sublunate sclerites; spiculum relictum present, slightly sclerotized (Fig. 10). Spiculum gastrale (Fig. 11) with apical arms short, widely divergent.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF940C18FF425380FB65FD1C.taxon	materials_examined	Material examined. 1 male, “ Australia ” (handwritten on a white card), “ Samml. K. F. Hartmann / Ankauf 1941.1 ” (typed on a blue card), and “ Staatl. Museum für / Tierkunde Dresden ” (typed on a white card) (SMTD); 1 male, “ Clarence ” (handwritten on a white card), “ N. S. Wales / 15176 ” (typed and partially handwritten on a blue card), and “ Staatl. Museum für / Tierkunde Dresden ” (typed on a white card) (SMTD).	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF940C18FF425380FB65FD1C.taxon	discussion	Remarks. The elytro-tergal stridulatory organ and terminalia of female were not examined.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF950C1DFF4257DCFC31F900.taxon	description	(Figs. 12 – 14, 19 – 21 & 25 – 36)	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF950C1DFF4257DCFC31F900.taxon	diagnosis	Diagnosis. From all known Tepperia and Evaniocis species, this species can be easily distinguished by the pronotum and elytra covered with light-colored scales except several brown scaly markings. In addition, this species is characterized by the rostrum whose dorsal contour is slightly curved and the elytral humeri covered by dark brown scales.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF950C1DFF4257DCFC31F900.taxon	description	Redescription. Male. Measurements (n = 4; in mm): TL: 7.9 – 8.6 (lectotype 7.9, mean 8.1); LP: 2.4 – 2.5 (2.4, 2.4); WP: 2.6 – 2.7 (2.6, 2.7); LE: 5.5 – 6.0 (5.5, 5.6); WE: 3.5 – 3.8 (3.5, 3.6). Integument reddish to dark reddish brown. Body generally covered with dull-white to yellowish brown recumbent scales. Frons and vertex covered with subovate scales; medial part of vertex with ill-defined dark brown patch. Rostrum covered with brownish subovate scales on basal half; apical margin of squamous part shallowly notched; apical half subglabrous, sparsely covered with minute scales. Pronotum covered with subovate scales, lacking suberect scales, with two pairs of dark brown scaly patches on postmedian and median parts along midline; postmedian patches connected with median ones by slightly paler scales, sometimes become obscure; median patches elongate apically with slightly paler scales; apical part covered with darker scales along margins except middle, with imbricate scales medially. Lateral parts of prothorax covered with general scales sparsely mingled with suberect scales, with irregular patch of brownish scales on middle of each side; prosternal canal mostly glabrous, but apical part densely covered with light-colored slender scales. Scutellar shield glabrous. Elytra mostly covered with subovate scales, usually with three pairs of irregular patches of dark brown scales on basal part between 2 nd and 5 th intervals, near humeri, and behind middle between 2 nd and 4 th intervals, sometimes with additional pair of patches on subapical swellings; basal patches indistinctly marginated and connected laterally with humeral patches by paler scales; sutural intervals covered with smaller scales, which become sparser on basal part; strial punctures each bearing whitish subrecumbent spathulate scale. Mesoventral receptacle glabrous on inside of receptacle canal, densely covered with minute scales on apical margin. Mesepimera, mesanepisterna, and lateral parts of mesoventrite mostly covered with subovate scales. Metaventrite and metanepisterna also covered with light-colored subovate scales; metaventrite glabrous on postmedial part; sclerolepidia absent. Femora irregularly mottled with light-colored to brown scales on anterior surfaces; each femur with triangular glabrous portion on apico-ventral part. Abdominal ventrites closely covered with pale brown scales, sparsely mingled with subrecumbent ones. Head (Fig. 19). Apical part of vertex gently flattened medially; frons flattened, 0.8 times as wide as base of rostrum at narrowest point between eyes. Rostrum stout, LR / WRM 2.4 – 2.5, LR / LP 0.7, in profile slightly curved ventrad, very slightly attenuate toward apex, excepting apical part more strongly attenuate; dorsal surface mostly slightly convex but sometimes slightly carinate along midline, rugosely to densely punctate from base to apical part, excepting impunctate portion along midline; punctures become smaller toward apex; medial part of apex depressed, impunctate, flanked by two shallow oblique furrows; sides gradually attenuate from base to before middle, then subparallel to subapical part, and finally weakly bulged near apex; apical margin shallowly arcuate in median part; dorsal contour slightly arcuate in profile. Antennae inserted at middle; scape strongly clavate, slightly shorter than funicle; funicle with 1 st and 2 nd segments subconical, with 3 rd to 7 th barrel-shaped, successively dilated distally; approximate length ratio of 1 st to 7 th as follows: 3: 2: 1: 1: 1: 1: 1.3; club fusiform, 2.0 – 2.5 times as long as wide, 0.6 – 0.7 times as long as funicle, loosely segmented. Mandibles robust, each with rounded outer margin and large inner tooth. Thorax. Prothorax subconical, LP / WP 0.9, weakly constricted subapically; pronotum densely punctate, uniformly convex, indistinctly carinate along midline; sides parallel or slightly attenuate from base to behind middle, then roundly convergent to subapical constriction, and finally slightly attenuate to apical margin, which is strongly arched in dorsal view; lateral parts rugosely punctate, obscurely granulate on middle. Mesoscutellum with scutellar shield round to subovate, approximately as wide as elytral sutural interval, weakly convex. Elytra subovate, LE / WE 1.5 – 1.6, with weakly produced humeri, wider than prothorax, WE / WP 1.3 – 1.4, evenly convex excepting subbasal part weakly prominent between 2 nd to 4 th intervals, with weak subapical swellings; sides subparallel from humeri to apical 2 / 5, then gently arcuately convergent to subapical swellings, and finally rounded apically, slightly emarginate at apices. Intervals weakly convex, wider than striae, partially granulate; sutural ones faintly depressed near base; 8 th and 9 th irregularly granulate on humeri; 9 th granulate behind humeri to just before middle, then granulate-costate to subapical part; 10 th flattened basally, granulate-costate excepting base. Striae shallowly sulcate excepting middle of 9 th more deeply sulcate; 10 th relatively short, terminated before middle; ventral surface with submarginal ridges sublinear in ventral aspect, not arched anteriorly, with two asymmetric files on apico-internal part. Femora robust, gently expanded from base to before middle, not clavate, with ventral tooth near middle, shallowly ventrally sulcate from middle to apex; dorsal contour weakly arcuate; profemora slightly longer than either meso- or metafemora. Tibiae simple along outer margins, not tuberculate; pro- and mesotibiae (Fig. 21) gently attenuate toward apex, obtusely angulate basally on outer margins; mesotibiae slightly shorter than either pro- or metatibiae, hardly to barely curved inward; metatibiae subparallel-sided, more weakly angulate basally on outer margins than on pro- and mesotibiae. Tarsi stout; 1 st tarsomere subconical, 1.6 times as wide as long, approximately twice as long as and as wide as 2 nd, 1.1 – 1.3 times as long as 3 rd; 2 nd subrectangular, 1.3 – 1.5 times as wide as long, half as wide as 3 rd; 3 rd subcordate, 1.5 times as wide as long; 5 th clavate, slightly longer than 3 rd. Rostral canal reaching level between anterior margins of mesocoxae; prosternal canal approximately as wide as procoxa, gradually expanded toward apex; mesoventral receptacle subtrapezoidal, 1.3 – 1.4 times as wide as long, shallowly concave along anterior margin, strongly bulged ventrally, overhanging anteriorly above canal, declined posteriorly, on declivity, indistinctly costate along midline and compressed laterally. Metaventrite flattened or slightly depressed medially, 1.3 – 1.6 times as long as mesoventrite along midline; apical part triangularly depressed; apical depression declined medially, slightly impressed along midline. Abdomen. Ventrites irregularly punctate; ventrite I with abdominal process narrowly acuminate, approximately 1.4 times as long as II along midline, flattened medially, shallowly emarginate along apical margin; II slightly longer than III and IV combined; III and IV subequal in length, each punctate in distorted row; V subtrapezoidal, approximately 3.0 times as wide as long, with apical margin subtruncate. Terminalia. Tergite VII (Fig. 25) transverse, subtrapezoidal, with two longitudinal distorted rows of setiferous tubercles on medial part. Rectum with rectal ring. Penis (Figs. 26 & 27) with body much shorter than its apodemes, approximately 1.5 times as long as wide, weakly sclerotized dorsally, subparallel-sided from base to subapical part, in lateral view flattened dorso-ventrally, moderately curved ventrally; apex (Fig. 28) rounded, flanked by several minute setae; endophallus spinulate basally, with tubular copulatory sclerite which is widened apically, dorsally produced in two laminae apically. Tegminal ring (Fig. 29) with weakly sclerotized parameroid lobes, which are fused basally; diameter of tegminal ring longer than its apodeme. Sternite VIII (Fig. 30) composed of two narrowly separated sublunate sclerites; spiculum relictum present, slightly sclerotized (Fig. 30). Spiculum gastrale (Fig. 31) shorter than penis; apical arms short, widely divergent, with narrow basal sclerotized flange. Female. Measurements (n = 3; in mm): TL 6.7 – 8.7 (mean 7.8), LP 2.1 – 2.6 (2.4); WP 2.3 – 2.9 (2.6); LE 4.6 – 6.1 (5.4); WE 3.0 – 3.9 (3.5). Rostrum (Fig. 20) LR / WRM 2.7, LR / LP 0.6 – 0.7; lateral parts of dorsal surface with sparser punctures on apical half; sides more strongly attenuate from base to middle. Prothorax LP / WP 0.9 – 1.0. Elytra LE / WE 1.5 – 1.6, WE / WP 1.3 – 1.4. Ventrite I slightly inflated medially excepting abdominal process flattened. Terminalia. Tergite VII (Fig. 32) subtrapezoidal, slightly wider than long, with two longitudinal distorted rows of setiferous tubercles on medial part; apical margin gently arcuate; tergite VIII (Fig. 35) slightly longer than wide; sides rapidly tapered apicad; apical margin weakly arcuate, notched at middle. Spiculum ventrale (Fig. 36) with apical plate subovate, gradually tapered apicad, notched at middle of apical margin; apodemes short, approximately as long as apical plate. Ovipositor (Fig. 33) with styli slender, approximately 1 / 3 length of gonocoxite; vagina and bursa copulatrix simple, neither spinulate nor sclerotized. Spermatheca (Fig. 34) c-shaped; cornu short, gradually attenuate and slightly curved apically; collum obliquely convex downward; ramus vestigial; insertions of duct and gland widely separated; gland short, sclerotized basally.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF950C1DFF4257DCFC31F900.taxon	materials_examined	Type material examined. Lectotype (here designated; SMTD), male: “ typus ” (surface, typed) and “ palawani- / cus Heller ” (back side, handwritten) (red card), “ Acc. No. 6108 / Lot / Bu. of Sci., P. I. ” (typed and partly handwritten on a white card), “ Collected by / E. D. Merrill. ” (typed on a pale brown card); “ 1911 / 7 ” (typed and partially handwritten on a yellow card), “ Staatl. Museum für / Tiekunde Dresden ” (typed on a white card), and “ LECTOTYPE / Cyamobolus / palawanicus / Heller, 1913 / Des. N. Tsuji & H. Yoshitake, 2025 ” (typed on a red card; newly attached). Paralectotype, 1 female: “ Acc. No. 6108 / Lot / Bu. of Sci., P. I. ” (typed and partly handwritten on a white card), “ Collected by / E. D. Merrill. ” (typed on a pale brown card); “ 1911 / 7 ” (typed and partially handwritten on a yellow card), “ syntypus ” (typed on a reddish orange card), “ Staatl. Museum für / Tiekunde Dresden ” (typed on a white card), “ bei Metacymia? / Heller det: 1934 ” (partially handwritten and stamped on a white card), and “ PARALECTOTYPE / Cyamobolus / palawanicus / Heller, 1913 / Des. N. Tsuji & H. Yoshitake, 2025 ” (typed on a red card; newly attached). Non-type material examined. The Philippines: [Palawan Is.] 3 males & 2 females, Taytay, W. Schultze leg. (SMTD).	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF950C1DFF4257DCFC31F900.taxon	distribution	Distribution. The Philippines (Cebu Island and Palawan Island). Natural history. Schultze (1916: 184) mentioned as “ Very injurious to seeds of Heritiera littoralis Dryand. ”, suggesting that T. palawanica larvae develop in fruits of H. littoralis.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF900C10FF4257D8FCECFD80.taxon	description	(Figs. 15 – 18, 22 – 24 & 37 – 54) http: // zoobank. org / urn: lsid: zoobank. org: act: B 1 F 814 A 4 - 4 E 1 E- 4 C 16 - 8809 - 763 B 9 BFF 9037	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF900C10FF4257D8FCECFD80.taxon	diagnosis	Diagnosis. This species is similar to T. palawanica in general appearance but can be clearly distinguished from it by having paired brown longitudinal scaly bands on the pronotum and a thicker rostrum whose dorsal contour is more strongly arcuate.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF900C10FF4257D8FCECFD80.taxon	description	Description. Male. Measurements (n = 10; in mm): TL 7.1 – 8.2 (holotype 7.9, mean 7.7), LP 2.1 – 2.6 (2.5, 2.4); WP 2.4 – 2.8 (2.7, 2.7); LE 5.0 – 5.6 (5.4, 5.3); WE 3.3 – 3.9 (3.8, 3.7). Most parts of body covered with dull-white to yellowish brown recumbent scales. Vertex with paired medial patches of dark brown scales behind eyes. Rostrum covered with brownish subovate to oblong scales, which become smaller and slenderer toward apex. Pronotum mainly covered with close light-colored subovate scales, sparsely mingled with whitish and brownish suberect scales, which become slenderer toward apex; medial part bearing paired narrow triangular patches of brownish scales along midline; apical margin with paired tufts of brownish suberect scales on middle. Lateral parts of prothorax densely covered with light-colored scales, sparsely mingled with dark brown suberect scales; prosternal canal mainly glabrous except apical part densely covered with light-colored slender scales. Scutellar shield subglabrous, except apico-lateral part fringed with minute scales. Elytra mostly covered with light-colored to brownish subovate scales, with the following two paired markings of dark suberect scales: obscure stripes on subbasal parts of 4 th intervals and spots on behind middle of 4 th intervals; dark spots sometimes extending to 3 rd intervals; strial punctures each bearing whitish subrecumbent spathulate scale. Mesoventral receptacle densely covered with minute scales on apical margin. Metaventrite and metanepisterna also covered with light-colored subovate scales, except glabrous patch on post-median part of metaventrite. Legs with femora and tibiae irregularly mottled with light-colored to brown scales. Profemur with triangular glabrous patch on apico-ventral part; meso- and metafemur glabrous ventrally along ventral sulci. Abdominal ventrites closely covered with pale brown recumbent scales and sparsely mingled with subrecumbent scales. Head (Fig. 22). Rostrum LR / WRM 2.6 – 2.9, LR / LP 0.7 – 0.8, thicker, in profile more strongly curved ventrad; sides more weakly attenuate from base to before middle; dorsal contour more strongly arcuate. Antennae inserted just behind middle; approximate length ratio of 1 st to 7 th funicular segments as follows: 2.8 – 3.5: 2: 1: 1: 1: 1.2: 1.2 – 1.4; club 2.3 – 2.6 times as long as wide, 0.7 – 0.8 times as long as funicle. Thorax. Prothorax LP / WP 0.8 – 0.9; sides slightly attenuate from base to behind middle. Elytra LE / WE 1.4 – 1.5, WE / WP 1.4 – 1.5, with subbasal part weakly separately prominent on sutural intervals and 3 rd and 4 th intervals. Sutural intervals with minute irregular granules; 9 th weakly granulate-costate from about middle to subapical part; 10 th weakly granulate-costate. 10 th striae more deeply sulcate on entire length. Meso- and metafemora with ventral sulci arising from base. Tibiae weakly tuberculate basally on outer margins, then subparallel-sided apically; mesotibiae (Fig. 24) slightly curved inward. Tarsi with 1 st tarsomere 1.5 – 2.3 times as wide as long, 1.7 – 1.9 times as long as and as wide as 2 nd, 1.5 – 1.8 times as long as 3 rd; 2 nd subquadrate, as long as wide; 3 rd 1.6 – 1.8 times as wide as long; 5 th almost as long as 3 rd. Metaventrite approximately 1.3 times as long along midline as maximum length of mesoventrite. Abdomen. Ventrites irregularly punctate; ventrite I slightly longer, 1.6 – 1.9 times as long as II along midline, hardly to barely emarginate along apical margin. Terminalia (Figs. 37 – 45). Penis with body approximately 1.4 times as long as wide, subparallel-sided or slightly expanded in basal 1 / 3, then gently convergent apically, in profile slightly thinner; apex more weakly arcuate, faintly notched at middle. Female. Measurements (n = 10; in mm): TL 5.4 – 8.3 (mean 7.6), LP 1.6 – 2.5 (2.3); WP 1.8 – 2.8 (2.6); LE 3.8 – 5.8 (5.2); WE 2.6 – 3.9 (3.7). Rostrum (Fig. 23) LR / WRM 2.5 – 3.0, LR / LP 0.7 – 0.8; dorsal surface with punctures sparser on apical half; sides more strongly attenuate from base to middle. Prothorax LP / WP 0.9. Elytra LE / WE 1.4 – 1.5, WE / WP 1.4 – 1.5. Ventrite I slightly inflated medially excepting abdominal process flattened. Terminalia (Figs. 46 – 50). Tergite VIII with apical margin simple, not notched at middle. Spiculum ventrale with apical plate subparallel-sided basally. Spermatheca thicker, with cornu more strongly curved apicad.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF900C10FF4257D8FCECFD80.taxon	etymology	Etymology. The specific name, “ vagabunda ”, is a Latin adjective meaning vagabond, referring to its dispersal habit with fruits of Heritiera littoralis.	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF900C10FF4257D8FCECFD80.taxon	materials_examined	Type material. Holotype, male (KUM): “ (Ryukyus) / Nakaragawa River / Iriomote I. / I. Matoba ”, “ 28. IV. 2005 / サキシマスオウノキ / DZƗよりī 化 ”, “ HOLOTYPE / Tepperia / vagabunda / Tsuji et Yoshitake, 2025 ” (typed on a red card; newly attached). Paratypes. Japan: the Ryukyu Islands: [Ishigakijima Is.] 1 female, Omoto-dake, 16. III. 1964, Y. Miyatake leg. (KUM). 1 ex., Omoto, 8. VI. 1975, T. Takahashi leg. (KUM). [Iriomotejima Is.] 1 ex., Haiminaka, 22. III. 2023, K. Fukui, S. Katoh, H. Hoshito & K. Nishida leg., as a fruit of Heritiera littoralis, emerged 6. V. 2023 (PCKF). 10 exs., Komi, 25. XII. 2004, J. Kantoh leg. (PCHY); 1 female, 20. V. 2005, I. Matoba leg., emerged from a fruit of H. littoralis (KUM); 2 males & 1 female, 21. V. 2005, I. Matoba leg., emerged from fruits of H. littoralis, (KUM); 1 male, 7. VI. 2005, I. Matoba leg., emerged from a fruit of H. littoralis, (KUM); 7 exs., 29. III. 2020, H. Yoshitake leg., as fruits of H. littoralis, emerged 15. IV – 1. V. 2020 (PCHY). 1 male, Tudwumari-hama (Tsukigahama) Beach, 5. III. 2015, M. Wakabayashi leg., as a fruit of H. littoralis, emerged 27. IV. 2015 (KUM); 1 ex., 26. III. 2022, M. Maruyama, T. Hashizume & S. Imada leg., as a fruit of H. littoralis, emerged 5. V. 2022 (KUM). 1 male & 3 females, same data as the holotype (KUM); 2 males & 1 female, same locality as the holotype, 25. IV. 2005, I. Matoba leg. (KUM); 2 males & 2 females, 26. IV. 2005, I. Matoba leg. (KUM & PCIM); 1 male & 7 females, 1. V. 2005, I. Matoba leg. (KUM). 1 male, “ Sonae ” [sic!] (Sonai), 30. V. 1977, J. Okuma leg., at light (KUM). 1 male, Amitori, 29. X. 1988, M. Iwata leg. (KUM). 2 exs., east coast, 16 – 19. X. 2011, S. Tetsu leg., as a fruit of H. littoralis, emerged 10. XII. 2011 (PCHY). Taiwan: 1 male, Kuraru, 31. VIII. 1921, T. Esaki leg. (KUM). Non-type material examined. Japan: the Ryukyu Islands: [Iriomotejima Is.] 1 female, same data as the holotype (KUM; an abnormal specimen with distorted elytral apices).	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
039487E1FF900C10FF4257D8FCECFD80.taxon	distribution	Distribution. Japan: the Ryukyu Islands (Ishigakijima Island and Iriomotejima Island); Taiwan. Natural history. Tepperia vagabunda is associated with Heritiera littoralis. The type series emerged from H. littoralis fruits washed ashore (Fig. 51). Larvae grew in and fed on H. littoralis seeds (Figs. 52 & 53) and pupated in the seeds when they fully matured. One or more individuals were found inside a seed (Fig. 53). New adults emerge through holes made in H. littoralis pericarps (Figs. 53 & 54).	en	Tsuji, Naomichi, Yoshitake, Hiraku (2025): Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae). Zootaxa 5613 (1): 126-138, DOI: 10.11646/zootaxa.5613.1.5, URL: https://doi.org/10.11646/zootaxa.5613.1.5
