taxonID	type	description	language	source
03946B0DFFCF472F247CFF32742AFE11.taxon	materials_examined	Type species: Shinkailepas kaikatensis Okutani, Saito & Hashimoto, 1989, by original designation. Diagnosis: [Modified from Fukumori et al. (2019).] Phenacolepadids with limpet-formed shell as adults. Protoconch purple, larval operculum smooth. Cephalic lappets present. Epipodial fold serrated. Mantle caeca present. Eye lobe separate from neck lobe, or partly to completely merged into it.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	description	(Figs 2 – 4)	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	diagnosis	Diagnosis: Medium-sized Shinkailepas (SL ≤ 8.5 mm) with apex slightly bent to left, positioned near posterior shell margin at about 90 % from anterior edge. Shell sculpture cancellate with concentric sculpture equal in strength or stronger than radial ones, drawn out to form weak nodes where they cross. Protoconch bulbous, smooth except for micro-pustules, about 660 μm in size when measured from adult shell. Diameter of opercular nucleus (see Fukumori and Kano 2014) 475 μm. Ostracum translucent, with silvery sheen. Operculum entirely corneous, not calcified. Epipodial fold with 12 – 15 short, triangular, tentacular projections.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	materials_examined	Type locality: Wocan hydrothermal vent field, Carlsberg Ridge. Type material: Holotype (RSIO 38219; Fig. 2 A), female, 95 % ethanol; SL 5.3 mm, SW 4.2 mm; Wocan vent field (60 ° 31.8 ʹ E, 6 ° 21.6 ʹ N), 2920 m deep, collected by a seven-function manipulator of HOV Jiaolong on dive # 129, R / V Xiangyanghong 9 cruise DY 38, March 2017. Paratype 1 (RSIO 38220; Figs 2 B, 3 D), female, 95 % ethanol; SL 6.5 mm, SW 5.4 mm. Paratype 2 (RSIO 38221; Figs 2 C, 3 C), female, 95 % ethanol; SL 4.5 mm, SW 3.6 mm. Paratype 3 (RSIO 38222; Fig. 3 B), female, 95 % ethanol, radula dissected for SEM (Fig. 4 E – F); SL 4.1 mm, SW 3.1 mm. Paratype 4 (RSIO 38223; Fig. 2 D), male, 95 % ethanol; SL 3.7 mm, SW 2.9 mm. Paratype 5 (RSIO 38224; Fig. 4 C – D), male, 95 % ethanol, periostracum removed for SEM; SL 5.9 mm, SW 3.9 mm. Paratype 6 (RSIO 38225; Fig. 4 C – D), female, 95 % ethanol (Fig. 3 A); SL 4.8 mm, SW 3.7 mm. All paratypes with same collecting data as holotype. Material examined: One shell (NSMT-Mo 79568; Fig. 4 A – B), periostracum of the protoconch removed for SEM, 99 % ethanol; SL 8.4 mm, SW 6.1 mm; Kairei vent field (70 ° 2.4187 ʹ E, 25 ° 19.2315 ʹ S), 2424 m deep, collected by suction sampler mounted on HOV Shinkai 6500 on dive # 1449, R / V Yokosuka cruise YK 16 - E 02, Feb 13 2016. Lot of five specimens (RSIO 38226), 95 % ethanol, up to SL 5.1 mm, SW 3.9 mm; Wocan vent field (60 ° 31.8 ʹ E, 6 ° 21.6 ʹ N), 2970 m deep, collected by a seven-function manipulator of HOV Jiaolong on dive # 125, R / V Xiangyanghong 9 cruise DY 38, March 2017.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	description	Description: Shell (Fig. 2) medium-sized for genus, nearly limpet-form as adults but conspicuously coiled at juvenile stage. Aperture oval, 1.1 – 1.3 times as long as wide. Height about one-third of length. Apex overhanging, close to shell edge at approximately 10 % of length from posterior margin, recurved slightly to left side. Shell sculpture cancellate from crossing of concentric sculpture with radials, forming low protrusions where two directions meet. Concentric sculptures stronger than or equal in strength to radials. Shell margin nearly flat to slightly raised (Fig. 2 A – B). Septum (‘ deck’) large, conspicuous, occupying anterior one-fourth of aperture at posterior (Fig. 2 B, E). Inner shell surface carry numerous shell pores, absent on septum. Ostracum thin, translucent-white with silvery sheen. Shell microstructure (Fig. 4 C) of three layers dorsal to myostracum, outermost homogeneous layer thin, followed by another thin granular layer, innermost crossed-lamellar layer thick. Periostracum thin, translucent, light to bright green (Fig. 2 A-D), typically with some sulphide mineral deposits on top (Fig. 3 D), in extreme cases shell may appear to become reddish in appearance due to rusty deposits, as known for some other Shinkailepas species (Fukumori et al. 2019). Protoconch (Figs 2 F, 4 A) white, covered by dark purple-red periostracum, bulbous, approximately 660 μm in length when measuring exposed part from adult shells, completely smooth except for minute dimples of 1 – 2 μm in size (Fig. 4 B). External anatomy (Figs 2 C – D, 3 A – C) typical for genus (Sasaki et al. 2006). Cephalic tentacles simple conical, tapered, elongate. Cephalic lappets small, positioned immediately anterior of cephalic tentacle. Left cephalic lappet develops into conical penis (Fig. 2 D) in males. Triangular neck lobe present posterior of cephalic tentacles. Eyes present, with dark pigments. Oral disc (Fig. 2 C – D) circular with strongly muscular outer lip, surrounded by well-developed oral lappet as wide as oral disc to either side. Dorsal surface (Fig. 3 A) covered by thin mantle with thickened edge lacking major tentacles. Hypobranchial gland lacking. Shell muscle divided into two, left shell muscle reaching slightly anterior of right shell muscle. Pallial edge slightly anterior of left shell muscle edge. Single ctenidium bipectinate, oblique in position with base at posterior left extending to anterior right. Intestine visible on dorsal mantle wall immediately posterior of ctenidium. Pallial gonoduct attached on dorsal mantle ceiling, surrounded anteriorly by capsule gland and posteriorly by albumen gland in females; in males replaced by prostate anteriorly and annex gland posteriorly. Pericardium triangular, on posterior left just posterior or ctenidium. Part of stomach visible on most posterior part of dorsal visceral mass, surrounded by extensive digestive gland dorsal to extensively developed gonad. Foot (Fig. 2 C) with circular, flattened sole, anterior pedal gland present. Epipodial fold carries 12 – 15 short, triangular tentacles. Operculum (Fig. 3 C) present, attached to dorsal surface of foot below visceral mass, thin, translucent, rather symmetrical in outline but with thickened nucleus area at left side, entirely corneous with no evidence of calcification. Diameter of opercular nucleus (see Fukumori and Kano 2014) 475 μm. Radula (Fig. 4 E – F) rhipidoglossate, formula c. 80 – 4 – 1 – 4 – c. 80. Central tooth oblong-quadrate, twice as long as wide, dorsal surface smooth, distally with very weak anterior ridge, overhanging cusp lacking. Innermost first lateral large, film-like, obliquely positioned with outer edge much higher than central tooth. Anterior edge of first lateral rolled to form weakly overhanging, smooth cusp; outermost edge raised to form vertical ridge. Second lateral small with inwardly recurved shaft, positionally occupying anterior half of first lateral, distally equipped with double overhanging cusps. Anterior cusp of second lateral smooth, posterior cusp with five to six denticles. Third lateral with strongly rolled shaft enveloping second lateral, cusp oblique with single major denticle whose inner side carries irregular serration. Outermost fourth lateral large, 1.5 times as long as third lateral, with slightly inwardly recurved shaft that gradually expands distally to form large, overhanging cusp with one inner, major denticle plus three outer, minor denticles. Inner marginals (Fig. 4 F) as long as fourth lateral, distally serrated into rake-like cusp with five to six denticles. Outer marginals much longer than inner marginals with elongate shafts ending in very finely serrated cusps, increasingly so outwards. Egg capsule (Fig. 3 D) about 0.6 mm in size, oblong oval, semi-transparent, similar in shape to other Shinkailepas egg capsules previously illustrated in literature (Metaxas 2011). Only one found on shell surface (of paratype 1; Fig. 2 B), at least seven embryos can be seen inside the capsule.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	etymology	Etymology: Greek, tiára (‘ headdress’) + asimía (‘ silvercoloured’), feminine adjective. Refers to the silver sheen of the ostracum and its cap-like shape, reminiscent of oriental silver headdresses.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	distribution	Distribution: Known with certainty from hydrothermal vents on both the CR (Wocan field) and the CIR (Kairei field).	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCF472B24B3FE6075ADFCF6.taxon	discussion	Remarks: Morphologically S. tiarasimia most closely resembles S. conspira from the Lau Basin in the southwestern Pacific (Poitrimol et al. 2022, Chen and Sigwart 2023), which exhibits similar shell sculpture with concentric ribs stronger than radial ones equal in strength, a similar apex position near the shell posterior, as well as a similar radula morphology. These two species, however, can be clearly differentiated in that the epipodial lobe of S. conspira is well developed and bears up to 60 paddle-like flaps (Chen and Sigwart 2023) compared to S. tiarasimia where only 12 – 15 small triangular tentacles are present. Shinkailepas conspira is also notably a much larger species reaching up to 21.3 mm in SL (Chen and Sigwart 2023). Another species sharing a similar adult shell sculpture is S. tufari from the Manus Basin vents also in the southwestern Pacific (Poitrimol et al. 2022); however, S. tufari has a much larger septum positioned at approximately the midpoint of the aperture (Beck 1992). While S. myojinensis typically exhibits stronger shell sculpture than S. tiarasimia (Sasaki et al. 2003) this species can be morphologically very variable and some specimens may resemble S. tiarasimia (Yahagi et al. 2017), but the operculum of S. myojinensis is partly calcified (likewise in S. conspira and S. tufari), whereas in S. tiarasimia it is entirely corneous. The protoconch of both S. conspira and S. tufari are both about 800 μm in size (Beck 1992, Chen and Sigwart 2023), much larger than the 660 μm in S. tiarasimia; while that of S. myojinensis is similar and also 660 μm (Sasaki et al. 2003). However, we note that our protoconch size is measured from the exposed part in adult shells unlike those of S. myojinensis and S. tufari which are measured using juveniles (Beck 1992, Sasaki et al. 2003). As the teleoconch of Shinkailepas often envelops the protoconch during growth, our measurement of the protoconch size is likely less accurate than those measured from juveniles. The central teeth of S. tiarasimia also appear to lack a horizontal ridge that is prominent in S. myojinensis (Sasaki et al., 2003).	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	description	(Figs 5 – 7)	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	description	ZooBank registration: urn: lsid: zoobank. org: act: BD 73673 D- FE 2 F- 449 C- 82 CE-DA 2 F 6 A 4 B 989 D	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	diagnosis	Diagnosis: Medium-sized Shinkailepas (SL ≤ 12 mm) with the apex weakly inclined to the right and positioned at approximately 70 % from the anterior shell edge. Shell sculpture weakly cancellate with radial ribs much stronger than concentric ribs, raised nodes are formed where the two directions meet. Septum (‘ deck’) reduced and minute. Epipodial fold carries 15 – 18 small triangular tentacles. Operculum entirely corneous. Diameter of opercular nucleus approximately 500 μm.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	materials_examined	Type locality: Daxi hydrothermal vent field, Carlsberg Ridge. Type material: Holotype (RSIO 38227; Figs 5 B, 6 – 7), male, 95 % ethanol, SL 11.7 mm, SW 8.4 mm, male; Daxi vent field, Carlsberg Ridge, 60 ° 10.8 ʹ E, 6 ° 48.0 ʹ N, 3480 m deep, R / V Xiangyanghong 9 cruise DY 38, collected by a seven-function manipulator of HOV Jiaolong on dive 126, March 2017. Paratype 1 (RSIO 38228; Fig. 5 C), female, 95 % ethanol, shell broken during collection, animal approximately 8.5 mm in length and 5.5 mm in width, same collecting data as holotype. Material examined: One specimen (RSIO 38229; Fig. 5 A), female, 95 % ethanol, SL 6.7 mm, SW 5.0 mm, Wocan vent field (60 ° 31.8 ʹ E, 6 ° 21.6 ʹ N), 2920 m deep, collected by a seven-function manipulator of HOV Jiaolong on dive # 125, R / V Xiangyanghong 9 cruise DY 38, March 2017.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	description	Description: Shell (Fig. 5) medium-sized for genus, adult limpet-formed, with evidence of coiling when juvenile. Shell length 1.2 – 1.4 times of width; shell height approximately onequarter of length. Aperture oval, reduced and minute septum (‘ deck’) present posteriorly. Apex positioned at approximately 30 % from posterior shell edge, weakly recurved to right side. Shell sculpture dominated by very strong radial ribs crossing with concentric growth lines to form weakly cancellate sculpture; raised protuberances form where two directions converge. Radial sculpture with three to five minor ribs between each pair of major ribs. Shell margin rather flat. Shell interior translucent-white, with numerous shell pores. Ostracum moderately thin, translucent. Numerous shell pores present on inner ostracum surface but absent on septum. Shell microstructure (Fig. 7 A) with three layers dorsal to the myostracum, a thin outermost granular layer, followed by a thick simple prismatic layer, then by an equally thick crossed lamellar layer. Protoconch unknown as corroded in all specimens available for study. Periostracum moderately thick, greenish, often covered in rusty mineral deposits. External anatomy (Fig. 6 A – C) typical for genus, most characters agree well with those described under S. tiarasimia above. Epipodial fold (Fig. 5) carries 15 – 18 small triangular tentacles posteriorly. Operculum (Fig. 6 D) large, entirely corneous, occupying over half of dorsal surface of foot, exact shape could not be determined due to its folded condition in examined specimens from preservation issues. Diameter of opercular nucleus approximately 500 μm, but due to its folded state, exact measurement was not possible. Radula (Fig. 7 B – D) rhipidoglossate, overall dentition pattern typical for genus (Fukumori et al. 2019), formula c. 100 – 4 – 1 – 4 – c. 100. Central tooth subquadrate, about 1.5 times as long as wide, with thickened central ridge running horizontally. Innermost first lateral film-like, strongly oblique, twice as wide as central tooth, cusp indistinct, lacking serration, curled over distally to form overhanging flap, outermost side with raised fold. Second lateral narrower than central tooth, hook-shaped with shaft recurved inwards, inner side of shaft with one raised hump; double-cusped with upper one smooth, lower one carrying four sharp minor cusps. Third lateral (Fig. 7 D) narrow with inward-rolled shaft, with one minute denticle, otherwise smooth. Outermost fourth lateral large, with solid, smooth, very slightly inwardly recurved shaft, distally enlarging to form wide cusp divided into five denticles, of which second from inside strongest by far. Inner marginals with shafts similar in length to fourth lateral, serrated into 4 – 5 denticles distally. Outer marginals much smaller, cusp rake-like, serrated into many fine denticles whose numbers increase outwards.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	etymology	Etymology: The specific epithet is a combination of Latin cornū (‘ horn’) and Ancient Greek thaûma (‘ wonder, marvel, astonishment’), referring to its horn-like, curved apex and the wonderful happening of collecting the few specimens known, allowing its formal description. This name was also partly inspired by the magical helm ‘ cornuthaum ’ that appears in the roguelike video game NetHack. Used as noun in apposition.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	distribution	Distribution: Only known from the Daxi and Wocan vent fields on the CR.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
03946B0DFFCB472724A1FCC57182FC5A.taxon	discussion	Remarks: Shinkailepas cornuthauma is clearly distinct from other described Shinkailepas species by its relatively anterior apex position at approximately 30 % of SL from the posterior shell edge, a very reduced septum (‘ deck’), combined with its strong radial shell sculpture with three to five minor ribs between two major ribs. Shinkailepas tollmanni is the only similar species with a similarly reduced condition of the septum, but its sculpture consists solely of fine radial ribs which are smooth instead of strongly granulated as in S. cornuthauma (Beck 1992, Fukumori et al. 2019). Furthermore, the epipodial tentacles of cornuthauma are small, triangular, and number between 15 – 18, which aids to distinguish it from species like S. conspira and S. gigas whose epipodial folds are divided into over 50 large, paddle-like projections (Chen and Sigwart 2023, Senckenberg Ocean Species Alliance et al. 2024). Molecular analyses Our phylogenetic tree reconstructed using the COI gene (Fig. 8) recovered a strongly supported monophyletic genus Shinkailepas (Bayesian Posterior Probability, BPP = 0.9) including S. tiarasimia and S. cornuthauma, supporting their inclusion in this genus. Two major clades were recovered within Shinkailepas, one containing S. kaikatensis, S. gigas, and S. cornuthauma (‘ Clade A’ hereafter) and the other with all remaining species (‘ Clade B’ hereafter); both were strongly supported (BPP = 0.94 and 0.96, respectively). Within Clade A, S. cornuthauma was the earliest diverging species sister to the S. kaikatensis — S. gigas pair, but only with low support (BPP <70). Within Clade B, there was a basal split between the fully supported (BPP = 1) Shinkailepas aff. tufari Woodlark Basin and Shinkailepas sp. Mariana Trough (sensu Poitrimol et al. 2022) pair and all other taxa. Shinkailepas tiarasimia was recovered as sister to S. conspira with strong support (BPP = 0.98). This pair was in turn clustered with S. aff. tufari Lau Basin & Futuna Arc (Poitrimol et al. 2022) with moderate support (BPP = 0.84). These three taxa were weakly supported (BPP <0.70) as sister to the S. myojinensis and S. tollmanni pair. Shinkailepas tufari was found to be the closest relative of these five species, with moderate support (BPP = 0.83). From estimates of genetic divergence between Shinkailepas species using the COI gene (Table 1), the K 2 P distance between S. tiarasimia and other known congeners ranges from 6.65 to 8.86 % and the same for S. cornuthauma ranges from 7.05 to 11.75 %. This is roughly equivalent to the range observed among other Shinkailepas species (5.50 – 11.42 %), and above the lowest value of 5.50 % between two described species (S. tollmanni — S. tufari). These support the recognition of S. tiarasimia and S. cornuthauma as new species of Shinkailepas.	en	Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong, Sun, Jin (2025): Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1): 1-15, DOI: 10.1093/zoolinnean/zlae167, URL: https://doi.org/10.1093/zoolinnean/zlae167
