identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039E87CD862BD06495D16641FD65F89F.text	039E87CD862BD06495D16641FD65F89F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aechmea cunhambebensis Leme	<div><p>Aechmea cunhambebensis Leme, s p. nov. (Fig. 1 A–J)</p><p>Diagnosis:––This new species is morphologically similar to Aechmea sumidourensis Leme (2015: 87), differing from it mainly by its inflorescence narrower (ca. 5 cm vs. 6–7 cm in diameter), with rachis dark winish-purple (vs. greenish), primary branches with fewer flowers (3–5 vs. 4–10 in number per branch), rachis visible (vs. completely hidden by the floral bracts), and smaller flowers (15–16 mm vs. 19–21 mm long).</p><p>Type: –– BRAZIL. Rio de Janeiro: Parque Estadual de Cunhambebe, Rio Claro, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.16428&amp;materialsCitation.latitude=-22.902056" title="Search Plazi for locations around (long -44.16428/lat -22.902056)">Lídice</a>, trilha para a <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.16428&amp;materialsCitation.latitude=-22.902056" title="Search Plazi for locations around (long -44.16428/lat -22.902056)">Pedra Chata</a>, “Vale”, 1,420 m elev., 22º 54’ 07.4” S, 44º 09’ 51.4” W, 30 July 2015, E . Leme 9034 &amp; C . Mendes (holotype RB!) .</p><p>Description:–– Plant terrestrial or epiphytic, flowering 60–70 cm tall, propagating by stout, basal shoots. Leaves ca. 20 in number, rosulate, suberect, coriaceous, forming a funnelform rosette; sheath elliptic, 20–23 × 10–11 cm, densely and minutely white lepidote on both sides, abaxially greenish, adaxially purplish-wine colored toward the apex; blade narrowly lingulate, inconspicuously if at all narrowed toward the base, 45–62 × 5–5.5 cm, green, densely white lepidote abaxially with trichomes almost completely obscuring the leaf color, adaxialy densely and inconspicuously white lepidote, apex acute to acuminate, ending in spine ca. 8 mm long, margins densely spinose; spines dark brown, narrowly triangular, flat, 1–3 mm apart, subspreading, straight to antrorse, the basal ones 2–3 mm long, 0.8–1.2 mm wide at the base, the upper ones 0.5–1.5 mm long, ca. 0.5 mm wide at the base. Peduncle erect, ca. 40 cm long, ca. 0.7 cm in diameter, dark winish-purple, densely white lepidote, completely covered by the peduncle bracts and not visible; peduncle bracts narrowly lingulate, acute and distinctly apiculate, 15–21 × 3–4 cm, erect, densely and inconspicuously white lepidote, densely imbricate, distinctly exceeding the internodes and completely enfolding the peduncle, distinctly veined, papyraceous, stramineous, remotely spinulose to entire. Inflorescence (fertile part) subcylindric, once-branched, slightly shorter than to slightly exceeding the leaves, erect, ca. 15 cm long, ca. 5 cm in diameter, rachis 0.5–0.6 cm in diameter, straight, densely white lepidote, dark winish-purple; primary bracts resembling the floral bracts, slightly exceeding the stipes; primary branches ca. 45 in number, polystichously and densely arranged, slightly complanate, subspreading, 2–3.8 × 1.2–2 cm (excluding the petals), bearing 3–5 flowers densely and subdistichously arranged, shortly stipitate; stipes 3–5 × 4–5 mm, stout, densely white lepidote, dark winish-purple; rachis slightly flexuous, dark winish-purple, densely white lepidote; floral bracts partially enfolding the basal portion of the flowers, neither adnate to the rachis above its base nor pouch-shaped, equaling to slightly exceeding the ovary, ecarinate but bearing protruding midnerve toward the apex, broadly subtriangular, dark red, 6–9 × 5–8 mm (including the apical spine), thin in texture, distinctly nerved, densely white lepidote mainly abaxially, bearing fimbriate trichomes at the apical margins, acute, distinctly mucronate, ending in a brown mucro 2–2.5 mm long, entire. Flowers 15–16 mm long (including the petals), odorless, sessile; sepals subquadrate, obtuse and distinctly mucronulate, asymmetrical with the membranaceous lateral wing not exceeding the midnerve, erect, dark red, densely white lepidote, ecarinate, connate at the base for ca. 1 mm, ca. 7 × 4 mm, including the 1.5–1.7 mm long, brown, subspreading apical spine; petals narrowly spathulate, apex rounded and emarginate, 10–11 × 2.5–3 mm, free, whitish near the base and lilac toward the apex, suberect forming a subtubular corolla, without noticeable callosities, bearing at the base 2 sublinear, bifid to dentate appendages, ca. 2.5 × 0.3 mm; filaments ca. 6.5 mm long, slightly complanate, not dilated toward the apex, white, the antesepalous ones free, the antepetalous ones basally adnate to the petals for ca. 4 mm; anthers ca. 3 mm long, dorsifixed slightly below the middle, white, base inconspicuously bilobed, apex apiculate; stigma conduplicate-spiral, subcapitate, white, ca. 1.5 mm long, margins shortly lacerate; ovary suboblong, terete, ca. 5.5 × 3.5 mm, red, densely white lepidote; epigynous tube infundibuliform, ca. 1.5 mm long; placentation central to apical; ovules obtuse. Fruits unknown.</p><p>Distribution and habitat:––This new species grows as a terrestrial or epiphytic species in a high elevation Atlantic Forest area (fig. 1 A), at an elevation of 1,420 m, in the State Park of Cunhambebe, in the locality of Lídice, county of Rio Claro, Rio de Janeiro state, southeastern Brazil. This conservation unit shelter a great extension of low to high elevated areas covered by Atlantic Forest in the southern portion of Serra do Mar, comprising the counties of Angra dos Reis, Mangaratiba, Rio Claro, and Itaguaí.</p><p>Aechmea cunhambebensis was found forming dense groups of plants scattered along the final portion of the trail to Pedra Chata, sharing its habitat with Nidularium antoineanum Wawra (1880a: 113), N. purpureum Beer (1857: 75), and Wittrockia superba Lindman (1891: 20), among other bromeliad species.</p><p>Etymology:––The name of Aechmea cunhambebensis is a reference to the State Park of Cunhambebe, where it was found.</p><p>Distinctive characters:— Aechmea cunhambebensis is morphologically related to A. sumidourensis, its likely closest relative, due to its habit, rosette and leaf conformation, inflorescence structure, and color of the flowers. However, the new species differs from it by its broader leaf blades (5–5.5 cm vs. 3.3–4 cm wide), which have spines more densely arranged (1–3 mm apart vs. 3–6 mm apart), inflorescence narrower (ca. 5 cm vs. 6–7 cm in diameter), with the rachis dark winish-purple (vs. greenish) and visible (vs. completely hidden by the floral bracts), primary branches more densely arranged throughout the rachis (vs. the basal ones more sparsely arranged in comparison with the upper ones), with fewer flowers (3–5 vs. 4–10 per branch), floral bracts smaller (6–9 × 5–8 mm vs. 9–11 × 13–15 mm), broadly triangular (vs. suborbicular), flowers smaller (15–16 mm vs. 19–21 mm long), and petals with sublinear, bifid to dentate basal appendages (vs. narrowly spathulate, rounded and crenulate appendages).</p><p>On the other hand, this new species is also related to Aechmea phanerophlebia Baker (1889: 47) . The important morphological differences of A. cunhambebensis are the inflorescence and primary branches with dark winish-purple rachis (vs. rose to reddish), the distinctly smaller flowers (15–16 mm vs. 24–25 mm long), the smaller floral bracts (6–9 × 5–8 mm vs. ca. 10 × 11 mm), sepals shorter (ca. 7 mm long, including the 1.5–1.7 mm long, brown, subspreading apical spine vs. ca. 10 mm long, including the ca. 4 mm long, yellowish-brown, erect or suberect apical spine), asymmetric, with a membranaceous lateral wing not exceeding the midnerve (vs. strongly asymmetric, with the membranaceous lateral wing distinctly exceeding the midnerve to slightly shorter than the apical spine), petals smaller (10–11 × 2.5–3 mm vs. 15–16 × 4.5 mm), anthers shorter (ca. 3 mm vs. ca. 5 mm long), and stigma subcapitate (vs. cylindric), ca. 1.5 mm long (vs. 2.5 mm long).</p></div>	https://treatment.plazi.org/id/039E87CD862BD06495D16641FD65F89F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8629D06B95D1642BFEF7FDC6.text	039E87CD8629D06B95D1642BFEF7FDC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aechmea marcpignalii Leme 2025	<div><p>Aechmea marcpignalii Leme, sp. nov. (Fig. 2 A–J)</p><p>Diagnosis:––This new species differs from Aechmea ramusculosa Leme (1995: 111) by the basal primary branches longer (15–26 cm vs. 10–12 cm long), with 10–15 secondary branches (vs. 2–5), flowers smaller (ca. 20 mm vs. ca. 33 mm long), and petals shorter (15–16 mm vs. ca. 27 mm long).</p><p>Type: –– BRAZIL. Bahia: Itambé, Catolezinho, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.24278&amp;materialsCitation.latitude=-15.13675" title="Search Plazi for locations around (long -40.24278/lat -15.13675)">Duas Barras</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.24278&amp;materialsCitation.latitude=-15.13675" title="Search Plazi for locations around (long -40.24278/lat -15.13675)">Fazenda Santa Teresinha</a>, 299 m elev., 15° 08’ 12.3” S, 40° 14’ 34” W, November 2010, R . Reis Jr. &amp; M . Pignal s.n., cult. E . Leme 8460 (holotype RB!) .</p><p>Description:–– Plant terrestrial or epiphyte, flowering 120–150 cm high. Leaves 10–15 in number, rosulate, suberect-arcuate, coriaceous, forming a broadly funnelform rosette; sheath broadly ovate, ca. 15 × 11 cm, castaneous toward the base, densely pale castaneous lepidote on both sides; blade sublinear, not narrowed toward the base, 80–100 × 5.5–7 cm, green to yellowish-green, densely white lepidote on both sides, trichomes forming a membrane adaxially, apex cuspidate, soon drying, becoming nigrescent, margins densely (toward the base) to sparsely (toward the apex) spinose; spines dark brown, triangular-uncinate, flat, prevailingly retrorse to sometimes nearly straight, the basal ones 4–6 mm long, 3–5 mm wide at the base, 4–10 mm apart, the upper ones 2–3 mm long, ca. 2 mm wide at the base, 15–25 mm apart. Peduncle suberect, ca. 50 cm long, 1–1.5 cm in diameter, red, white lanate; peduncle bracts narrowly lanceolate, the basal ones cuspidate, the distal ones acuminate, 9–12 × 2–2.5 cm, erect, nerved, the basal ones spinulose toward the apex, the distal ones entire, white lepidote toward the base, exceeding the internodes and enfolding the peduncle, pale rose to stramineous, the basal ones thinly coriaceous, the upper ones papyraceous. Inflorescence 4-times branched, paniculate, equaling the leaves, suberect, ca. 60 cm long (fertile part), ca. 35 cm in diameter at the base, 7–15 cm in diameter near the apex; rachis 5–10 mm in diameter, slightly flexuous to nearly straight, white lanate, red; primary bracts narrowly lanceolate, acuminate, entire, nerved, subdensely and inconspicuously white lepidote toward the base, stramineous, thin in texture, suberect to spreading with the branches, decreasing in size toward the inflorescence apex, 4–12 × 0.6–2 cm, distinctly exceeding the stipes but shorter than the branches; primary branches ca. 30 in number, polystichous, the basal to median ones laxly arranged, the upper ones subdensely to densely arranged, suberect to spreading, distinctly decreasing in length toward the inflorescence apex, the basal ones 15–26 cm long, bearing 10–15 secondary branches densely arranged toward the apex, the median to apical ones 4–10 cm long, bearing 3–6 secondary branches, the branches near the inflorescence apex resembling the secondary branches, sessile; stipes of the primary branches 3–6 × 0.4–0.7 cm, ebracteate, complanate, red, white lanate; rachis of the primary branches straight, white lanate, red; secondary bracts narrowly subtriangular-setiform or sublinear, 15–20 × 2–3 mm, exceeding the stipes but distinctly shorter than the branches, suberect to spreading, white sublanate to glabrous, stramineous, nerved; secondary branches 30–40 mm long, shortly stipitate to usually sessile, 2 to 6-flowered, usually verticillate, bearing 1 to 2 tertiary branches with one of it arising from the same basal node; rachis distinctly geniculate, white lanate, green, internodes 3–5 × 1.5–2 mm; floral bracts equaling the basal portion of the sepals, partially enfolding the ovary, with a subtriangular base 5–6 × 5.5–7 mm, and a spinescent apex, 4–4.5 mm long, green at the base and castaneous toward the apex, thin in texture, entire, nerved, ecarinate, white sublanate at the base to glabrescent. Flowers ca. 20 mm long, odorless, subdensely and distichously arranged, sessile; sepals subtrapeziform, 7 × 5–6 mm, distinctly asymmetrical with the lateral membranaceous rounded wing distinctly exceeding the midnerve but slightly exceeded by the apical spine, green with whitish spots, subdensely white lepidote, ecarinate, subfree, apical mucro 3–4 mm, castaneous, straight, erect to spreading, to irregularly curved; petals spathulate, 15–16 × 5–5.5 mm, acute, free, white at the base, greenish at the middle and lilac toward the apex, erect except for the slightly suberect apex, at the base bearing 2 narrowly spathulate, lacerate appendages of ca. 3.5 x 1 mm, as well as 2 conspicuous longitudinal callosities slightly shorter than the filaments; filaments 7.5–8 mm long, complanate, dilated toward the apex, white, the antesepalous ones free, the antepetalous ones basally adnate to the petal for ca. 3.5 mm; anthers narrowly oblong-ovate ca. 4 mm long, dorsifixed slightly below the middle, base shortly bilobed, apex narrowed, truncate to slightly bifid; stigma conduplicate-spiral, ellipsoidal, white, margins shortly crenulate-lacerate; ovary broadly obovoid, ca. 5.5 mm long, ca. 6 mm in diameter at the apex, glabrous, green with white spots, white lanate; epigynous tube inconspicuous; placentation apical; ovules caudate. Fruits unknown.</p><p>Distribution and habitat:–– Aechmea marcpignalii grows in a low-elevated area, about 300 m elevation, inside a deciduous forest in the Atlantic Forest biome (fig. 2 A), in the locality of Catolezinho, county of Itambé, Duas Barras, Bahia state, northeastern Brazil. The small forest fragment where it was found is located inside Santa Teresinha farm, which is owned by bromeliad and orchid lover, Raymundo Reis Junior. This new species grows as a terrestrial, forming large groups of plants, being sympatric with Cryptanthus bibarrensis Leme (2002: 86) and C. reisii Leme (2002: 87) which were originally discovered in the same region.</p><p>Etymology:––The name of this new species honors one of its collectors, the French botanist and taxonomist Marc Pignal, Curator of the National Herbarium of the Muséum National d´Histoire Naturelle of Paris, for his important contribution to the knowledge of Brazilian flora, mainly in Bahia state.</p><p>Distinctive characters:—This new species is a member of Aechmea subg. Aechmea, being close related to A. ramusculosa due to its size when flowering, the shape of the inflorescence, sepals, and petals, as well as the color of the corolla. Aechmea ramusculosa was originally described from a near sea level (i.e. coastal) Restinga vegetation in the locality of Copuva (or Copubá), in the county of Nova Viçosa, south region of Bahia. According to the label in the holotype [Hatschbach 48762 &amp; J.M. Silva (MBM!)], A. ramuscolosa has red bracts and lilac flowers, probably meaning that sepals and the well-preserved petals are lilac, giving an important clue on the general color appearance of the inflorescence of this species.</p><p>Aechmea marcpignalii morphologically differs from its close relative by its inflorescence 4-times branched (vs. 2 to 3-times branched), primary bracts stramineous (vs. red), the basal primary branches longer (15–26 cm vs. 10–12 cm long), with 10–15 secondary branches (vs. 2–5 in number), rachis distinctly geniculate (vs. slightly flexuous), floral bracts green at the base and castaneous toward the apex (vs. red), flowers smaller (ca. 20 mm vs. ca. 33 mm long), subdensely and distichously arranged (vs. laxly and polystichously arranged), sepals smaller (7 × 5–6 mm vs. ca. 12 × 9 mm), petals shorter (15–16 mm vs. ca. 27 mm long), basal appendages longer (ca. 3.5 mm vs. ca. 2 mm), lacerate (vs. crenulate), and the antepetalous filaments shorter adnate to the petals for ca. 3.5 mm (vs. higher adnate to the petals for ca. 10 mm).</p><p>On the other hand, the known populations of this new species are concentrated further inland and in higher areas about 150 km from the ocean and about 330 km in straight line from the known population of A. ramusculosa, at the seaside zone.</p></div>	https://treatment.plazi.org/id/039E87CD8629D06B95D1642BFEF7FDC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8626D06995D161E3FBF2FC5B.text	039E87CD8626D06995D161E3FBF2FC5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bromelia boqueiranensis Leme & J. A. Siqueira 2025	<div><p>Bromelia boqueiranensis Leme &amp; J.A. Siqueira, sp. nov. (Fig. 3 A–E)</p><p>Diagnosis:––This new species can be distinguished from the closest relative, Bromelia gurkeniana E. Pereira &amp; Moutinho (1983: 346) by the floral bracts glabrescent or glabrous (vs. densely brown lanate), sepals longer (22–24 mm vs. ca. 15 mm long), and free (vs. connate at the base for ca. 9 mm). It also differs from B. gurkeniana var. funchiana E. Pereira &amp; Leme (1985: 635) by the floral bracts obtuse (vs. acute), distinctly shorter than the sepals (vs. about equaling the sepals), and petals longer (38–40 mm vs. ca. 30 mm long).</p><p>Type: –– BRAZIL. Bahia: Santo Sé, Boqueirão da Onça, entre <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.419724&amp;materialsCitation.latitude=-10.334722" title="Search Plazi for locations around (long -41.419724/lat -10.334722)">Minas do Mimoso</a> e <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.419724&amp;materialsCitation.latitude=-10.334722" title="Search Plazi for locations around (long -41.419724/lat -10.334722)">Campo</a> dos Alegre, Serra da Embaúba, 1,177 m elev., 10º 20’ 05” S, 41º 25’ 11” W, 15 May 2010, E . Leme 8297, J. A . Siqueira Filho &amp; A. P . Fontana (holotype RB!) .</p><p>Description:–– Plant terrestrial in sandy soils, propagating by long rhizomes. Leaves ca. 7 in number, arcuate before anthesis and afterwards, thick coriaceous, green except for the inner ones becoming reddish toward the base at anthesis, forming at the base a thick, bulbous-like structure, 8–9 × 4.5 cm deeply sunk in the soil; sheath oblong-ovate, ca. 6 × 3 cm, thick coriaceous, abaxially densely and coarsely castaneous lepidote in the middle with ribbon-like, downwardly curved trichomes, white lepidote toward the distal end, adaxially glabrous; blade linear, attenuate, 75–120 × 0.8–1 cm, not narrowed at the base, strongly canaliculate, densely white lepidote and finely nerved abaxially, glabrous adaxially, apex caudate, margins densely to subdensely spinose at the base and entire for most of its length toward the apex; spines brown, strongly uncinate, complanate, 0.5–1.5 mm long, the proximal ones mostly retrorse, 0.2–0.5 mm apart, the distal ones retrorse, 1–2 mm apart. Peduncle inconspicuous, ca. 2 × 0.8 cm, deeply sunk in the rosette. Inflorescence shortly corymbose, the proximal half sunk in the rosette, densely once-branched, ca. 7 cm long (excluding the petals), ca. 2 cm in diameter in the middle; primary bracts oblong, 3.7–3.8 × 1.7 cm, glabrous except for the brown lepidote apex with fimbriate trichomes, subcoriaceous and stramineous toward the apex, whitish and membranaceous at the base, densely spinose toward the apex with mostly retrorse-uncinate spines, the basal ones with a narrowly subfoliaceous blade, ca. 7 cm long, entire, distinctly exceeding the branches, the upper ones apiculate, shorter than the sepals; fascicles inconspicuous, ca. 3 in number, ca. 2-flowered, inconspicuously stipitate; floral bracts shorter than the sepals, carinate, sublinear-oblong, whitish at the base and stramineous toward the apex, 30–32 × 4.5–5.5 mm, thin in texture, glabrescent or glabrous, nerved, apex obtuse, slightly cucullate, margins entire or inconspicuously denticulate. Flowers ca. 12 in number, 60–70 mm long, slightly fragrant, inconspicuously pedicellate; pedicel stout, ca. 5 × 3 mm, merging into the ovary, subdensely castaneous lepidote with fimbriate trichomes; sepals symmetrical, narrowly suboblong, apex obtuse, entire to inconspicuously denticulate, 22–24 × 5–6 mm, free, carinate toward the apex, cymbiform, nerved, corrugate, wine-purple toward the base and stramineous near the apex, subdensely castaneous lepidote at the base and along the keel, glabrous toward the apex; petals 38–45 mm long, glabrous, dark wine-purple (including the margins) at anthesis, reddish-wine to stramineous afterwards, connate for ca. 23 mm, without any callosities, lobes linear-oblong, rounded, ca. 4 mm wide, subspreading-arcuate at anthesis; filaments forming a common tube with the petals, the antesepalous ones free for 1–1.5 mm above the tube, the antepetalous ones almost completely adnate to the tube; anthers narrowly triangular, ca. 4 mm long, base distinctly bilobed, apex apiculate, dorsifixed at 1/5 to 1/6 of its length above the base; pollen ca. 50 μm, subglobose, sulcate, sulcus without exine elements, with margins well-defined, exine foveolate with tendency to reticulate, muri flat, lumina narrow to broad; stigma conduplicate-spiral, lobes white, crenulate-lacerate, ca. 2.5 mm long; ovary subclavate-cylindrical, nearly terete, ca. 17 × 6 mm, wine-purple, densely brown lepidote with fimbriate trichomes; epigynous tube funnelform, ca. 3 mm long; placentation from apical to near the base; ovules globulose, obtuse. Fruits unknown.</p><p>Distribution and habitat:–– Bromelia boqueiranensis is a terrestrial species living in sandy soils in areas covered by high altitude Caatinga vegetation (fig. 3 A), intermingled by rocky outcrops sheltering Rupestral Fields, in the county of Santo Sé, Bahia state, in a vast Caatinga area in northeastern Brazil known as Boqueirão da Onça, between the localities of Minas do Mimoso and Campo dos Alegre, in a hill called Serra da Embaúba, 1,177 m elevation. It grows deeply sunk in the sandy soil (fig. 3 B), being hard to remove, forming sparse groups of plants that spread vegetatively by means of long underground rhizomes.</p><p>Etymology:––The name of this new species is a reference to the area where it was discovered, in Boqueirão da Onça, the largest continuous area of Caatinga vegetation, recently transformed in a National Park. The area protects the largest population of jaguar (“onça” in Portuguese) in the Caatinga biome, besides many other endangered species of the fauna and flora.</p><p>Distinctive characters:— Bromelia boqueiranensis is morphologically related to B. gurkeniana, which is a species originally described from the Amazon region of Rio Gurupi, Pará state. This relationship arises from its linear leaf blades and the shortly corymbose inflorescence sunk in the rosette. However, this new species differs by its central leaves becoming reddish at anthesis (vs. green), floral bracts glabrescent or glabrous (vs. densely brown lanate), sepals longer (22–24 mm vs. ca. 15 mm long), free (vs. connate at the base for ca. 9 mm), and petals dark winish-purple (vs. white).</p><p>When compared to B. gurkeniana var. funchiana, this new species differs by the leaf blades with retrorse basal spines (vs. antrorse), floral bracts obtuse (vs. acute), distinctly shorter than the sepals (vs. about equaling the sepals), glabrescent or glabrous (vs. brown lanate near the apex), petals longer (38–40 mm vs. ca. 30 mm long), and the antepetalous filaments almost completely adnate to the petals (vs. free for ca. 3 mm above the common tube with the petals).</p><p>Bromelia gurkeniana var. funchiana inhabits the Caatinga vegetation of Chapada Diamantina, in the county of Palmeira, Bahia state, about 300 km distant in straight line from the county of Santo Sé, in the São Francisco valley, Bahia state, near the border with Pernambuco, where this new species was found.</p></div>	https://treatment.plazi.org/id/039E87CD8626D06995D161E3FBF2FC5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8624D06C95D16757FA40FB64.text	039E87CD8624D06C95D16757FA40FB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bromelia nordestina Leme & J. A. Siqueira 2025	<div><p>Bromelia nordestina Leme &amp; J.A. Siqueira, sp. nov. (Fig. 4 A–O)</p><p>Diagnosis:––This new species is closely related to Bromelia lagopus but differs from it by the shorter flowers (8–9 cm vs. ca. 5.7 cm long), sepals longer (30–35 mm vs. ca. 25 mm long), and petal longer (38–40 mm vs. ca. 32 mm long), lilac-purple (vs. rose), with obtuse apex (vs. subacute).</p><p>Type: –– BRAZIL. Alagoas, Chã Preta, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-36.34975&amp;materialsCitation.latitude=-9.216944" title="Search Plazi for locations around (long -36.34975/lat -9.216944)">Serra Lisa</a>, 815 m elev., 09º13’01” S, 36º20’59.1” W, 21 March 2005, E . Leme 6633, J. A . Siqueira-Filho &amp; M. S . Leite (holotype RB!) .</p><p>Description:–– Plant terrestrial, propagating by short rhizomes. Leaves ca. 32 in number, suberect before anthesis, subspreading at anthesis, coriaceous; sheath subreniform to subtrapeziform, 4–5 × 6–7 cm, thick, abaxially densely brown lepidote with irregularly stellate, lacerate trichomes with ribbon-like prolongations, adaxially glabrous and lustrous toward the base, margins spinose at the distal end; blade sublinear, attenuate toward the apex, 180–210 × 3.5–3.7 cm, not narrowed toward the base, strongly canaliculate toward the base, nearly flat near the apex, abaxially densely white lepidote, distinctly nerved, adaxially densely to sparsely white lepidote, green except for the inner ones being bright red near the base forming a large ring around the inflorescence, apex long caudate, pungent, recurved, margins laxly spinose, spines greenish to reddish near the base, castaneous toward the apex, strongly antrorse-uncinate, complanate, pungent, 4–8 mm long, 3–4 cm apart. Peduncle inconspicuous, 2–3 × 4.5 cm, thick; peduncle bracts the basal ones foliaceous, red toward the base and forming a large ring around the inflorescence, the median to upper ones with a basal sheath oblong, 8–10 × 4.5–5 cm, adaxially castaneous, glabrous and lustrous near the base, densely nerved, abaxially densely brown lepidote near the base with irregularly lacerate stellate trichomes with ribbon-like prolongation, the median to distal margins spinose, spines thin in texture to rigid, spreading to antrorse-uncinate, castaneous, 2.5–4 mm long, the distal blades narrowly lanceolate, red, abaxially subdensely white lepidote, adaxially inconspicuously and sparsely white lepidote, 15–30 × 1 cm (at the base), apex caudate-attenuate. Inflorescence once branched but resembling simple, capitate-corymbose, apex nearly flat, sunk in the center of the rosette, densely once-branched at the base, 13–14 cm long (not including the petals), 10.5–12 cm in diameter at the apex; primary bracts the outer ones resembling the upper peduncle bracts but with shorter blades, the inner ones oblong-spathulate, acute and apiculate, paleaceous, castaneous, erect, slightly shorter than the sepals, abaxially very densely brown lepidote toward the apex, margins spinose near the apex with spines thin in texture, 1–2 mm long; fascicles the outer ones complanate, flabellate, slightly elongate, 10–12 × 7 cm, 10–12-flowered, inconspicuously stipitate, stipes trapeziform, ca. 2.5 × 2 cm, the apical fascicles inconspicuous and the inflorescence appearing simple; floral bracts equaling the middle of the sepals, narrowly spathulate, 6–7 × 1 cm, nerved, abaxially densely brown lepidote, adaxially glabrous, apex acute, margins densely spinulose toward the apex, spines thin in texture, irregularly curved, the floral bracts of the fascicles carinate, cymbiform, those of the apical simple part of the inflorescence obtusely if at all carinate. Flowers diurnal, odorless, 8–9 cm long (including the petals), erect, inconspicuously pedicellate, pedicel merging with the ovary, 7–10 × 3–4 mm, densely brown lepidote; sepals symmetrical, sublinear, apex obtuse, slightly cucullate, 30–35 × 6–7 mm, subfree, carinate toward the apex, navicular, nerved, white-vinaceous near the base, castaneous toward the apex, adaxially glabrous, abaxially densely brown lepidote with trichomes irregularly lacerate with ribbon-like prolongations, margins entire; petals erect to slightly suberect at anthesis and forming a tubular corolla, distinctly exceeding the sepals, lilac-purple (including the margins) at anthesis, lilac-rose afterward, glabrous, 38–44 mm long, connate at base for 15–17 mm in a common tube with the filaments, blades lingulate, obtuse, 6–7 mm wide; filaments terete, white, ca. 32 mm long, equaly free above the common tube with the petals; anthers sublinear, 9–11 mm long, base, bilobed, apex long apiculate, apiculus strongly incurved mainly before anthesis, dorsifixed near the base; stigma conduplicate-spiral, lilac with white margins, blades ca. 5 mm long, margins scalloped, papillose; ovary subcylindrical, trigonous, subangulose, 32–37 × 5–6 mm, densely brown lepidote with trichomes irregularly lacerate, stellate with ribbon-like prolongation; epigynous tube distinctly but narrow, ca. 4 mm long; placentation basal; ovules globose, obtuse, sparsely arranged. Fruits narrowly ellipsoid, yellowish, 6.5 × 2–2.5 cm, not including the persistent sepals and pedicel, densely brown lepidote with trichomes irregularly lacerate with ribbon-like prolongations. Seeds 7–15 in number, suborbicular to broadly elliptic, 6–6.5 × 5–6 mm, complanate, dark brown.</p><p>Distribution and habitat:––This new species is a terrestrial inhabitant of the Atlantic Forest biome of northeastern Brazil, where it grows in partially shaded sites. It occurs in the region of Serra Lisa, county of Chã Preta, Alagoas state, in areas of 200-815 m elevation. Probably it also grows in the neighbouring state of Pernambuco. For a precise indication of its geographical range it is mandatory to perform a carefull revision of herbarium material to point out eventual misnamed specimens of potentially close related species, like Bromelia grandiflora Mez (1919: 3), B. lagopus Mez (1891: 188), B. magnifica Esteves &amp; Gouda (2017: 228), and B. villosa Mez (1901: 3), which is far beyond the scope of this work.</p><p>Etymology:––The name of Bromelia nordestina is a reference to the region related to its geographical range, in northeastern Brazil, based on the the Brazilian word “nordestino”, which means “from northeast”.</p><p>Additional specimens examined (paratypes):–– BRAZIL. Alagoas: Chã Preta, Serra Lisa, 21 March 2005, J. A . Siqueira-Filho et al. 1483 (UFP!); Murici, Est. Ecol. de Murici, 23 July 2008, R . Moura 624 (R!); Murici, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-35.844444&amp;materialsCitation.latitude=-9.248889" title="Search Plazi for locations around (long -35.844444/lat -9.248889)">Serra do Ouro</a>, 200‒500 m elev., 09º14’56” S, 35º50’40” W, 3 May 1999, G . Martinelli et al. 15335 (RB!) .</p><p>Distinctive characters:—This new species was circumscribed, illustrated, and provisionally identified by Leme &amp; Siqueira-Filho (2006) as Bromelia karatas sensu Jacquin (1770: pl. 31, 32) because at that time it was not known for sure which plant material would typify B. karatas, and no name could be safely attributed to it without first carrying out an ample revision of the genus. It was also verified by them that more than one taxon in northeast Brazil could be included in the rather imprecise concept of B. karatas then used.</p><p>The solution to the problem was proposed 10 years later by the typification of Bromelia karatas Linnaeus (1753: 285) by Monteiro &amp; Forzza (2016) who designated a lectotype for the species ‒ the type species of Bromelia Linnaeus (1753: 285) ‒ and narrowed its concept to be only applicable to the populations of Caribbean, Mexico, Central America, and northern South America. Monteiro &amp; Forzza (2016) also concluded that the specimens of Bromelia collected in northeastern and central-western Brazil contradict the concept of B. karatas and are likely to belong to a new species.</p><p>Subsequently, Pereira &amp; Gouda (2017) described Bromelia magnifica Esteves &amp; Gouda (2017: 228) as a new species from the Cerrado vegetation of the northeastern state of Bahia, near the São Francisco river, belonging to the so called “ Bromelia karatas complex”. According to the authors, B. grandiflora is its closest morphological relative. However, the precise identity of Bromelia species with inconspicuously pedunculate, nidular, and capitate inflorescence, like B. grandiflora, as well as B. lagopus, and B. villosa, remains challenging because the protologues and type-specimens do not provide all needed morphological information to accurately circumscribe them, e.g., the color of the corolla as currently attributed to these species.</p><p>Despite the morphological obscurities that involve this complex of species, it is possible to recognize similarities and distinguish Bromelia nordestina from B. lagopus, its putative closer relative due to its leaf blades with large marginal spines, the inner leaf blades being bright red near the base forming a large ring around the inflorescence, which is capitate-corymbose and sunk in the rosette, and the flowers densely brown lepidote except for the glabrous petals. This new species, however, differs from it by the longer flowers (8–9 cm vs. ca. 5.7 cm long), longer sepals (30–35 mm vs. ca. 25 mm long), petal longer (38–40 mm vs. ca. 32 mm long), lilac-purple (vs. rose), with apex obtuse (vs. subacute), and ovary longer (32–37 mm vs. ca. 20 mm long). Since there is no precise data in its protologue, the geographical range of B. lagopus is still not clearly known.</p><p>This new species can be also compared to Bromelia grandiflora, which does not have any geographical data associated to its protologue, since it was described from cultivated material of Victoria Hortus, in Cameroon, Africa. Based on the protologue and the analysis of the type specimen, it is possible to distinguish B. nordestina from B. glandiflora by its smaller floral bracts (6–7 × 1 cm vs. ca. 8 × 2 cm), which are brown lepidote (vs. villose), flowers brown-lepidote (except for the petals) with lacerate trichomes with ribbon-like prolongations (vs. long villose, trichomes more or less subspreading), shorter pedicel (7–10 mm vs. ca. 15 mm), and longer sepals (30–35 cm vs. ca. 26 mm long).</p><p>In relation to Bromelia magnifica, this new species is distinctly different due to its longer leaf blades (180–210 cm vs. ca. 120 cm long) with antrorse spines (vs. the basal ones retrorse), floral fascicles with more numerous flowers (10–12 vs. 2–4 in number), floral bracts longer (6–7 cm vs. ca. 4.5 cm long), narrowly spathulate (vs. sublinear), and acute (vs. obtuse and cucullate), flowers longer (8–9 cm vs. 6.2–7 cm long), sepals longer (30–35 mm vs. ca. 22 mm), entire (vs. minutely serrulate), and petals obtuse, not cucullate (vs. slightly cucullate), longer ovary (32–37 mm vs. ca. 15 mm), and fruits narrowly ellipsoidal (vs. ellipsoidal to obovate) and yellowish (vs. bright yellow). According to the protologue of B. magnifica, it is a typical species of the Cerrado vegetation of the northeastern state of Bahia, along the São Francisco river, while this new species is known from the Atlantic Forest of the states of Alagoas.</p><p>Despite more distantly related, Bromelia nordestina can be compared to B. karatas differing by its spathulate floral bracts (vs. narrowly oblong to linear-lanceolate), sepals obtuse (vs. subacute), petals shorter (38–40 mm vs. 45–55 mm long), lilac-purple (vs. pinkish to magenta), not cucullate (vs. cucullate), and fruits yellowish (vs. rose to dark red). The non-Brazilian geographical range of B. karatas (i.e., Caribbean, Mexico, Central America and northern South America), greatly contrast with the exclusive Brazilian Atlantic Forest range of B. nordestina .</p><p>Identification key for the species of the Bromelia karatas complex</p><p>1. Flowers 6.2–9 cm long .......................................................................................................................................................................2</p><p>- Flowers up to 6 cm long .....................................................................................................................................................................5</p><p>2. Sepals 30–35 mm long .......................................................................................................................................................................3</p><p>- Sepals 22–26 mm long .......................................................................................................................................................................4</p><p>3. Petals lilac purple; fruits yellowish ................................................................................................................................ B. nordestina</p><p>- Petals pinkish to magenta; fruits rose to dark red................................................................................................................ B. karatas</p><p>4. Floral bracts ca. 4.5 × 1.5 cm, much exceeded by the sepals.......................................................................................... B. magnifica</p><p>- Floral bracts ca. 8 × 2 cm, about equaling the sepals.................................................................................................... B. grandiflora</p><p>5. Leaves up to 1 m long; sepals ca. 17 mm long; petals purple............................................................................................... B. villosa</p><p>- Leaves longer than 1 m; sepals 25–27 mm long; petals rose .............................................................................................. B. lagopus</p></div>	https://treatment.plazi.org/id/039E87CD8624D06C95D16757FA40FB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8621D07095D1666BFD6BFE8B.text	039E87CD8621D07095D1666BFD6BFE8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bromelia saracataquerensis Leme, J. B. F. da Silva, J. A. Siqueira & E. H. Souza 2025	<div><p>Bromelia saracataquerensis Leme, J.B.F. da Silva, J.A. Siqueira &amp; E.H. Souza, sp. nov. (Figs. 5 A–E, 6 A–M)</p><p>Diagnosis:––This new species is closely related to Bromelia granvillei L.B. Smith &amp; Gouda (1996: 9), differing from it by the shorter rhizomes (ca. 5 cm vs. ca. 25 cm long), broader leaf blades (2.4–3 cm vs. 1.5–2.2 cm wide), sepals with longer free lobes (5–11 mm vs. 3–3.5 mm long), and by the longer ovary (20–25 mm vs. ca. 10 mm long).</p><p>Type: –– BRAZIL. Pará, Oriximiná, Porto Trombetas, FLONA Saracá-Taquera, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.380108&amp;materialsCitation.latitude=-1.6953611" title="Search Plazi for locations around (long -56.380108/lat -1.6953611)">Mineração Rio do Norte</a>, 103 m elev., 01º41’43.3” S, 56º22’48.4” W, 28 April 2018, E . Leme 9392, J . B . F. da Silva, J . A . Siqueira Filho &amp; E . Hilo de Souza (holotype RB!, isotype SEL) .</p><p>Description:–– Plant terrestrial, propagating by short rhizomes to ca. 5 cm long. Leaves 25–30 in number, suberect before anthesis, more or less spreading at anthesis, coriaceous, green, often cracked; sheath subreniform to subtrapeziform, 2–3 × 4.5–5 cm, thick, abaxially glabrous at the proximal end, densely covered with a dense coat of long sublinear ferrugineous trichomes downwardly oriented toward the distal end, adaxially glabrous at the proximal end to inconspicuously and sparsely to densely ferruginous lepidote with long filamentous trichomes upwardly oriented and coarsely corrugate toward the distal end, margins spinose at the apex with antrorse-uncinate spines; blade sublinear, 200–400 × 2.4–3 cm, distinctly narrowed toward the base but not petiolate, strongly canaliculate toward the base, abaxially sparsely to subdensely but inconspicuously white lepidote with trichomes concentrated along the intercostal areas, distinctly nerved, adaxially glabrous, attenuate toward the apex, apex long caudate, pungent, margins laxly spinose, spines castaneous near the apex, strongly retrorse-uncinate, complanate, pungent, 2–3 mm long, 1.5–2.5 cm apart. Peduncle inconspicuous, 2–3 cm long, thick; peduncle bracts foliaceous, pale rose at the base at anthesis. Inflorescence (fertile part) once branched, capitate-corymbose, sunk in the center of the rosette, congested at the base, ca. 6 cm long not including the petals, to ca. 10 cm long including the petals, 4–5 cm in diameter in the middle; primary bracts distinctly exceeding the fascicles, suberect, with an obovate base, 3.3–4 × 1.8–2.5 cm, densely castaneous lepidote mainly toward the base abaxially with peltate, irregularly dentate trichomes, glabrescent adaxially, coarsely and densely spinose toward the apex, with irregular curved, flat and soft spines, not pungent, the blades of the primary bracts narrowly and long subtriangular to long caudate, thin in texture, 1–17 × 0.3–0.8 cm, glabrous, pale rose, densely spinulose, spines antrorse, ca. 0.5 mm long; fascicles 5–6 in number, inconspicuous, complanate, flabellate, inconspicuously stipitate, 2–5 flowered; floral bracts slightly shorter to exceeding the ovary, thin in texture to membranaceous, densely nerved, abaxially densely castaneous lepidote with peltate, irregularly lacerate trichomes, adaxially glabrescent, margins densely spinulose toward the apex, spines thin in texture, irregularly curved, those of the fascicles sublinear to narrowly subspathulate, 20–40 × 3–9 mm, carinate mainly toward the apex, apex acute and apiculate, those of the inner simple part of the inflorescence suboblong, 35–45 × 14–19 mm, ecarinate, rounded and apiculate. Flowers ca. 25 in number, diurnal, sweetly fragrant, 80–88 mm long (including the petals), erect, inconspicuously pedicellate, pedicel merging with the ovary; sepals symmetrical, linear, apex obtuse-cucullate to emarginate, 25–27 × 3.5–5 mm, unequally connate at the base for 14–22 mm, carinate toward the apex to ecarinate, navicular, densely brown lanate mainly toward the base and the apex, with subpeltate, irregularly lacerate, long filamentous trichomes, inconspicuously spinulose at the apex with irregularly curved, minute spines, distinctly nerved; petals narrowly spathulate, apex obtuse to rounded, erect at anthesis except for the recurved apical portion, white, abaxially sparsely to subdensely pale brown lepidote toward the apex, 55–66 × 7–8 mm, connate at the base for 22–27 mm in a common tube with the filaments; filaments terete, white, the antesepalous ones equally adnate to the petals for 22–27 mm and free above it, the antepetalous ones adnate to the petals for 29–33 mm; anthers sublinear, 9– 11 mm long, base, bilobed, apex apiculate, dorsifixed near the base; stigma conduplicate-spiral, white, blades ca. 5 mm long, margins distinctly lacerate-scalloped, papillose; ovary narrowly subclavate to narrowly obovate, subangulose, 20–25 × 6–7 mm, whitish, densely brown lanate with subpeltate, irregularly lacerate, long filamentous trichomes; epigynous tube distinct but narrow, 3–3.5 mm long; placentation from central to apical; ovules globose, obtuse, sparsely arranged. Fruits ovoid to narrowly ellipsoid, whitish near the base and green toward the distal end, 3–3.7 × 1.8–2.5 cm, not including the persistent sepals and pedicel, with a distinct fruit-like fragrance, sparsely and irregularly brown lanate. Seeds ca. 10 in number, orbicular or nearly so, 5.5–6 × 5–5.5 mm, complanate, dark brown.</p><p>Distribution and habitat:––This new species is a typical terrestrial in Terra Firme forests, in the counties of Oriximiná (fig. 5 A) and Terra Santa, in the Amazonian state of Pará, northern region of Brazil. In the shaded and humid areas where it grows (fig. 5 E), their narrow leaves may reach 4 m length and are propped up by the surrounding bushes (fig. 5 B–C). It was observed living sympatrically with other terrestrial bromeliads like Disteganthus lateralis (L.B. Smith 1954: 525) Gouda (1994: 134) and Bromelia morreniana (Regel 1888: 157) Mez (1891: 186), to name a few.</p><p>Etymology:––The name of Bromelia saracataquerensis is a reference to the place where it was originally found, in the Floresta Nacional (FLONA) Saracá-Taquera in Porto Trombetas, Oriximiná, in the Amazonian state of Pará.</p><p>Additional specimen examined (paratypes): –– BRAZIL. Pará: Terra Santa, Plateau Aramã, terrestre em Mata de Terra Firme, April 2018, J. B . Fernandes da Silva s.n., cult. E . Leme 9374 (RB!); Santarém, BR 163, KM 67, June 2019, L. O. A . Teixeira s.n., cult. E . Leme 9697 (RB!) .</p><p>Distinctive characters:— Bromelia saracataquerensis (fig. 5 B–E, 6 A–M) belongs to the complex of Amazonian species formed by B. granvillei from French Guiana, B. gurkeniana, from the Gurupi river, in the state of Pará, near the border to the state of Maranhão, B. morreniana (fig. 6 N–O), and B. tubulosa L.B. Smith (1964: 40) (fig. 6 P–Q), both from the states of Amazonas and Pará, Brazil, except for the last one, which was originally described from Venezuela, not far from the border with Brazil. These species are characterized by the small and compact, capitate-corymbose inflorescence sunk in the center of the rosette due to the inconspicuous peduncle, sepals highly connate and with very short free lobes, and the white or whitish petals, which are inconspicuously castaneous lepidote abaxially and toward the apex, forming a prevailingly tubular corolla.</p><p>Despite Bromelia morreniana having been registered in the same area where this new species was collected, and although B. granvillei ‒ known from the northeast of French Guiana only ‒ being described in the fruit stage, without data on petals and stamens, it is possible to recognize that B. granvillei has a closer morphological affinity with B. saracataquerensis, both being characterized by very long, narrow, and often cracked leaf blades. However, this new species differs from B. granvillei by the propagation by means of much shorter rhizomes (ca. 5 cm vs. ca. 25 cm long), broader leaf blades (2.4–3 cm vs. 1.5–2.2 cm wide), with shorter spines along the margins (2–3 mm vs. 3–7 mm long), sepals with longer free lobes (5–11 mm vs. 3–3.5 mm long), longer ovary (20–25 mm vs. ca. 10 mm long), and by the larger fruit (3–3.7 × 1.8–2.5 cm vs. ca. 2.5 × 1 cm).</p><p>While Bromelia saracataquerensis is known as an inhabitant of the inland Terra Firme forest, ca. 100 m elevation, about 700 km from the Atlantic coast, B. granvillei is reported from 100 to 200 m elevated areas, about 30 to 50 km from the Atlantic coast, living in open forest on rocky soils with a thin layer of humus (Smith &amp; Gouda 1996).</p><p>In relation to Bromelia gurkeniana, this new species differs by the much longer leaf blades (200–400 cm vs. ca. 70 cm long), flowers longer (80–88 mm vs. ca. 55 mm long), longer sepals (25–27 mm vs. ca. 15 mm long), which are higher connate at the base for 14–22 mm (vs. ca. 9 mm), longer petals (55–66 mm vs. ca. 35 mm long), and by the longer anthers (9–11 mm vs. ca. 5 mm long).</p></div>	https://treatment.plazi.org/id/039E87CD8621D07095D1666BFD6BFE8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD863DD07695D16227FE69F822.text	039E87CD863DD07695D16227FE69F822.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptanthus cajuitensis Leme & L. Kollmann 2025	<div><p>Cryptanthus cajuitensis Leme &amp; L. Kollmann, sp. nov. (Figs. 7 A–I, 8 A–B)</p><p>Diagnosis:––This new species is closely related to Cryptanthus ubairensis I. Ramirez (1998: 221), but can be distinguished from it by the shorter caulescent habit (stem 7–19 cm long vs. 15–123 cm long), leaf blades smaller (7–10 × 1.9–2.4 cm vs. 13–24 × 2.5–4 cm), sepals shorter (11–14 mm vs. 15–16 mm long) and shorter connate (4–5 mm vs. 7–8 mm), and petals shorter (18–25 mm vs. ca. 33 mm long).</p><p>Type: –– BRAZIL. Bahia: Guaratinga, road Guaratinga-Cajuíta via <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.79767&amp;materialsCitation.latitude=-16.641333" title="Search Plazi for locations around (long -39.79767/lat -16.641333)">Jacutinga</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.79767&amp;materialsCitation.latitude=-16.641333" title="Search Plazi for locations around (long -39.79767/lat -16.641333)">Mata de Licuri</a>, em vertente muito íngreme, 285–350 m elev., 16º 38.48’ S, 39º 47.86’ W, 23 April 2009, E . Leme 7788, L . Kollmann, A. P . Fontana &amp; C . Esgario (holotype RB!, isotype SEL!, CEPEC) .</p><p>Description:–– Plant terrestrial, distinctly caulescent, stem 7–19 cm long, ascending toward the apex, propagating by elongate axillar basal shoots. Leaves 7 to 12 in number, spreading-recurved before anthesis and afterwards, laxly to subdensely and subequally disposed along the stem; sheath inconspicuous, subreniform, 0.8–1 × 1.5–1.7 cm, pale, glabrous or nearly so, rugulose abaxially, apical margins densely spinulose; blade lanceolate, 7–10 × 1.7–2.4 cm, distinctly narrowed toward the base but not petiolate, thinly coriaceous, pliable, without any thicker central zone, green to bronze colored, inconspicuously canaliculate mainly toward the base, densely and coarsely white-lepidote abaxially, trichomes obscuring the leaf color, adaxially sparsely white-lepidote at the base and glabrous toward the apex, apex acuminate-caudate, margins undulate, subdensely to densely spinulose, spines antrorse, green to bronze colored toward the base, 0.5–1 mm long, 2–5 mm apart. Inflorescence (fertile part) 2–2.5 cm long, 1–2 cm in diameter, sessile; primary bracts foliaceous; fascicles 1 to 4 in number, 20 × 7–8 mm (excluding the petals), 2–3-flowered; floral bracts equaling 1/3 to 1/2 of the sepal length, narrowly triangular-lanceolate to linear, hyaline toward the base, greenish near the apex, 10–14 × 1.5–3 mm, inconspicuously and sparsely lepidote to glabrous, acuminate, margins remotely denticulate, those of the fascicles carinate; Flowers 25–32 mm long (with extended petals), sessile, fragrance not detected, those of the upper central part of the inflorescence staminate, those of the fascicles both staminate and perfect; sepals 11–14 mm long, connate for 4–5 mm, glabrous, greenish-white, hyaline along the margins, lobes narrowly suboblong-lanceolate, acuminate, 6.5–10 × 2.5–3 mm, symmetrical, obtusely if at all carinate, margins remotely denticulate; petals sublinear, apex subacute, 18–25 × 3–4.5 mm, white except for the pale greenish apex mainly before anthesis, exceeding the stamens but suberect-recurved at anthesis and exposing the stamens, connate for 2–4 mm, with conspicuous ca. 5 mm long callosities at the base of the blades; filaments 16–18 mm long, the antepetalous ones adnate to the petals for ca. 6 mm, the antesepalous ones adnate to the petal tube; anthers 1.7–3 mm long, dorsifixed near the base, base distinctly bilobed, apex obtuse; pollen spherical or nearly so, ca. 60 μm in diameter, sulcate, sulcus large, densely covered by distinctly reticulate exine elements forming a net, exine reticulate, lumina irregularly polygonal, muri thick; stigma conduplicate-spreading, blades 3–3.5 × 0.8 mm, densely and shortely scalloped-lacerate, white. Ovary 6–8 × 3–4 mm, subtrigonous, white, glabrous; epigynous tube lacking; placentation apical; ovules few, obtuse. Fruits unknown.</p><p>Distribution and habitat:–– Cryptanthus cajuitensis is a terrestrial inhabitant of the Atlantic Forest biome of the southern part of Bahia state (fig. 7 A–B), northeastern Brazil, where it was found in the county of Guaratinga in the way to the locality of Cajuíta. It grows in a steep whatershed characterized by the presence of a large formation of “licuri”, an Arecaceae species of the genus Syagrus .</p><p>The area shelters other important bromelioid species, like Hohenbergia kollmannii, described in this study, as well as the paratype population of Neoregelia retrorsa Leme &amp; Kollmann (2011: 19), to name a few.</p><p>Etymology:—Since this new species was collected in the neighborhood of the locality of Cajuíta, its name is a reference to its geographical origin.</p><p>Distinctive characters:— Cryptanthus cajuitensis is morphologically related to C. ubairensis which includes its general vegetative appearance resulting from its caulescence and the lanceolate leaf blades distinctly narrowed toward the base but not petiolate. However, it differs from C. ubairensis by its shorter caulescent habit (stem 7–19 cm long vs. 15–123 cm long), leaf blades smaller (7–10 × 1.9–2.4 cm vs. 13–24 × 2.5–4 cm), floral bracts smaller (10–14 × 1.5–3 mm vs. 17–19 × 8–10 mm), flowers smaller (25–32 mm vs. ca. 42 mm long), sepals shorter (11–14 mm vs. 15–16 mm long) and shorter connate at the base for 4–5 mm (vs. 7–8 mm), petals shorter (18–25 mm vs. ca. 33 mm long) and connate at the base for 2–4 mm only (vs. ca. 8 mm).</p><p>Cryptanthus venecianus Leme &amp; L. Kollmann (Leme et al. 2010: 31), from Nova Venécia, Espírito Santo state, is another morphologically related species due to some shared features like caulescent habit, leaf blades narrowed toward the base, and flowers of similar size. However, C. cajuitensis can be distinguished from it by the less numerous leaves (7–12 vs. 14–18), leaf blades distinctly narrowed toward the base (vs. slightly narrowed toward the base), floral bracts narrowly triangular-lanceolate (vs. triangular), much narrower (1.5–3 mm vs. 7–8 mm), sepals with lobes narrowly suboblong-lanceolate (vs. broadly ovate), narrower (2.5–3 mm vs. ca. 4 mm), petals smaller (18–25 × 3–4.5 mm vs. 28–29 × 4.5–5 mm), anthers dorsifixed near the base (vs. dorsifixed in the middle), and pollen (fig. 8 A–B) with sulcus densely covered by distinctly reticulate exine elements (vs. sparsely covered by inconspicuously reticulate exine elements; fig. 8 C–D).</p></div>	https://treatment.plazi.org/id/039E87CD863DD07695D16227FE69F822	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD863BD07595D164CFFE59FD1F.text	039E87CD863BD07595D164CFFE59FD1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptanthus spathulatifolius Leme & E. H. Souza 2025	<div><p>Cryptanthus spathulatifolius Leme &amp; E.H. Souza, sp. nov. (Fig. 9 A–K)</p><p>Diagnosis:––This new species can be distinguished from the morphologically closest relative, Cryptanthus pseudopetiolatus Philcox (1992: 265), by its shorter leaf blades (30–48 cm vs. 50–75 cm long), the basal ones subpetiolate with the subpetioles to equaling the length of the broader distal portion (vs. the basal ones distinctly narrowed toward the base but not subpetiolate), floral bracts and sepals white lepidote (vs. brown lepidote).</p><p>Type: –– BRAZIL. Bahia: Guaratinga, ca. 6 km from the city in the road Guaratinga to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.8&amp;materialsCitation.latitude=-16.65" title="Search Plazi for locations around (long -39.8/lat -16.65)">Cajuíta</a>, ca. 250 m elev., 16º39’ S, 39º48’W, leg. J . E. dos Santos s.n., fl. cult. December 2020, E . Leme 9815 (holotype RB!, isotype HURB!) .</p><p>Description:–– Plants stemless, propagating by short basal shoots. Leaves 7 to 12 in number, subspreading at anthesis, laxly arranged and forming an open rosette; sheath broadly subtrapeziform to subtriangular-reniform, greenish toward the base, reddish near the apex, densely lepidote, corrugated, nerved abaxially, adaxially glabrescent; blade narrowly spathulate to narrowly lanceolate, distinctly narrowed toward the base and the basal ones subpetiolate, 30–48 × 3.6–6.4 cm (subpetiole included), coriaceous, reddish-ferruginous or sometimes green toward the apex, abaxially densely and coarsely white lepidote, adaxially glabrous toward the apex and subdensely to densely white lepidote near the base, apex caudate, margins slightly undulate, canaliculate mainly toward the base, flat toward the apex, densely spinulose; spines triangular, reddish-ferruginous, antrorsely uncinate, 0.3–0.5 mm long, 1–4 mm apart; subpetiole 8–25 × 1.5–2.2 cm, gradually merging into the much broader distal portion, distinctly shorter than to equaling the length of the broader distal portion. Inflorescence (fertile part) sessile, shortly corymbose, ca. 4.5 cm long, ca. 3.5 cm in diameter, once branched at the base with prevailingly perfect flowers and bearing a simple central head of densely arranged staminate flowers; primary bracts resembling the leaves; fascicles ca. 6 in number, subpulvinate, sessile, the basal ones ca. 35 × 17 mm (petals excluded), with 3–6 perfect and staminate flowers; floral bracts ovate-lanceolate, greenish-white to reddish along the keel, 20–22 × 7–11 mm, membranaceous, sparsely white lepidote near the apex to glabrescent, acute, margins densely spinulose towards the apex, those of the fascicles slightly exceeding the ovary, carinate, those of apical simple head of flowers to equaling the middle of the sepals, ecarinate. Flowers sessile, odorless, the perfect ones ca. 52 mm long (with the petals extended), the staminate ones ca. 45 mm long; sepals 16–19 mm long, connate for 8–10 mm, greenish-white, sparsely white lepidote, lobes broadly ovate-lanceolate, long acuminate, 7–8 × 3.5–4 mm, symmetrical, obtusely carinate, margins densely and irregularly spinulose, spines membranaceous; petals narrowly spathulate, apex acute to subrounded, white, exceeding the stamens but spreading-recurved at anthesis and exposing them, 34–36 × 7–8 mm, weakly connate at the base for ca. 12 mm, bearing 2 longitudinal callosities at the base of the free lobes; filaments ca. 28 mm long, adnate to the petal tube and free above it; anthers 3.5–6 mm long, dorsifixed at 2/5 of its length above the base, base long bilobed, apex apiculate; stigma conduplicate-patent, lobes ca. 3 mm long, white, margins scalloped, without papillae; ovary narrowly subclavate, trigonous, ca. 15 × 5 mm, white, glabrous; epigynous tube lacking; placentation subapical; ovules few, obtuse. Fruits not seen.</p><p>Distribution and habitat:–– Cryptanthus spathulatifolius is known from its type locality only, which is situated about 6 km from the city of Guaratinga along the road to the locality of Cajuíta, southern Bahia state, northeastern Brazil. It grows as a terrestrial (fig. 9 A) in a low-elevated hill (about 250 m elevation) covered by a humid Atlantic Forest fragment, nearby a small farm community.</p><p>Etymology:––The name of this new species is a reference to its narrowly spathulate leaf blades, which are distinctly narrowed toward the base and much broadened in their distal portion.</p><p>Additional specimen examined (paratype): –– BRAZIL. Bahia: Guaratinga, ca. 6 km from the city in the road Guaratinga to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.8&amp;materialsCitation.latitude=-16.65" title="Search Plazi for locations around (long -39.8/lat -16.65)">Cajuíta</a>, ca. 250 m elev., 16º39’ S, 39º48’W, leg. J . E. dos Santos s.n, fl. cult. February 2021, E . Leme 9944 (RB!)</p><p>Distinctive characters:— Cryptanthus spathulatifolius is morphologically closely related to C. pseudopetiolatus due to plant size, inflorescence conformation, and the shape of the floral bracts, sepals, and petals. However, C. pseudopetiolatus is a confusing species with floral morphology poorly known due to its partially inaccurate original description and the broad species concept adopted by Ramírez (1998) including three or more taxa (Leme et al. 2020), making it challenging to accurately establish morphological differences.</p><p>Cryptanthus pseudopetiolatus is an inhabitant of low-elevated area of the coastal Atlantic Forest in the southern region of Bahia State, about 15 km from the ocean in straight line, while C. spathulatifolius has a more inland occurance about 70 km in straight line from the ocean, in areas at about 250 m elevation. The new species differs from its likely closest relative by the shorter leaf blades (30–48 cm vs. 50–75 cm long), the basal ones subpetiolate with the subpetioles to equaling the length of the broader distal portion (vs. the basal ones distinctly narrowed toward the base but not subpetiolate), floral bracts equaling the middle of the sepals (vs. exceeding the middle of the sepals), smaller (20–22 × 7–11 mm vs. 25–26 × 10–25 mm), white lepidote (vs. brown lepidote), and sepals white lepidote (vs. brown lepidote).</p><p>In comparison to Cryptanthus ruthae Philcox (1992: 268), which also has distinctly subpetiolate leaf blades and grows in the lowland, coastal region of Maraú and neighbourhood, south Bahia state, this new species differs by the much broader leaf blades (3.6–6.4 cm vs. 2.5–3 cm wide), margins densely spinulose throughout (partially entire, mainly toward the base), fascicles with more numerous flowers (3–6 vs. 2–3 in number), floral bracts densely spinulose towards the apex (vs. entire or nearly so), sepals with lobes broadly ovate-lanceolate (vs. narrowly ovate to elliptic), larger (7–8 × 3.5–4 mm vs. 6 × 3–3.5 mm) and margins densely and irregularly spinulose (vs. entire to laxly and inconspicuously spinulose), and petals larger (34–36 × 7–8 mm vs. 28–30 × 4–4.5 mm).</p><p>Another species with petiolate leaves and similar floral morphology is Cryptanthus capitellatus Leme &amp; L. Kollmann (Leme et al. 2010: 29), which growns in the mountainous region of Santa Teresa, Espírito Santo state. In relation to it, the distinctive characteristics of C. spathulatifolius are the larger leaf blades (30–48 × 3.6–6.4 cm vs. 9–31 × 1.5–4.5 cm), the basal fascicles with more numerous flowers (3–6 vs. ca. 2 in number), floral bracts ovate-lanceolate (vs. sublinear-lanceolate to narrowly triangular), larger (20–22 × 7–11 cm vs. 14–17 × 3–7 cm), flowers longer (ca. 52 cm vs. 32–40 cm long), sepals with lobes broadly ovate-lanceolate (vs. narrowly ovate) and larger (7–8 × 3.5–4 mm vs. 6 × 2–2.5 mm), and petals larger (34–36 × 7–8 mm vs. 24–30 × 4–5 mm) and completely white (vs. white with greenish apex).</p></div>	https://treatment.plazi.org/id/039E87CD863BD07595D164CFFE59FD1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8638D07B95D161ABFB80FD1F.text	039E87CD8638D07B95D161ABFB80FD1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dyckia albolutea Leme & A. S. Farias-Castro 2025	<div><p>Dyckia albolutea Leme &amp; A.S. Farias-Castro, sp. nov. (Fig. 10 A–N)</p><p>Diagnosis:––This new species can be distinguished from the morphologically closest relative, Dyckia tubifilamentosa Wanderley &amp; G. Souza (2014: 315), by the sepals free (vs. connate at the base) and shorter (6–7 mm vs. 8–11 cm), petals yellow (vs. green), and the filament tube white at anthesis and afterwards (vs. white at anthesis becoming winish or purplish colored afterwards).</p><p>Type: –– BRAZIL. Ceará: Granja, Serra da Timbaúba, distrito de Pessoa Anta, São Miguel, próximo <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.02837&amp;materialsCitation.latitude=-3.36255" title="Search Plazi for locations around (long -41.02837/lat -3.36255)">Cachoeira de São Miguel</a>, 104 m elev., 03º21’45.18” S, 41º01’42.13” W, 26 May 2022, A. S . Farias-Castro 3109, cult. E. Leme 10168 (holotype RB!, isotype SEL!) .</p><p>Description:–– Plant saxicolous, clustering, flowering 27–45 cm tall; Leaves 14–18 in number, densely rosulate, strongly thick coriaceous; sheath much broader than the blades, subreniform, ca. 1.8 × 2.5 cm, whitish toward the base, yellowish-green to yellowish and corrugate near the distal end, glabrescent; blade suberect-arcuate to subspreading, narrowly triangular, attenuate toward the apex, 8–10 cm long, 1.2–1.5 cm wide at the base, ca. 4 mm thick in the middle, abaxially inconspicuously and sparsely white lepidote to glabrescent, distinctly nerved, adaxially densely and coarsely white lepidote near the base and inconspicuously and sparsely white lepidote to glabrescent toward the apex, slightly canaliculate mainly toward the base, nearly flat toward the apex, apex long-caudate, margins subdensely (at the base) to laxly (toward the apex) spinose; spines narrowly triangular, 1–2 mm long, 0.5–0.8 mm wide at the base, 3–12 mm apart, flat, glabrous, dark castaneous toward the apex, antrorse to sometimes nearly straight; peduncle lateral, 18– 30 cm × 1.5–2.5 mm, erect, rigid, glabrous, green; peduncle bracts with a subtriangular base, 3–4 × 2.5–3 mm, and a filiform distal blade, 10–15 × 0.3–0.5 mm, distinctly shorter than the internodes, erect or nearly so, entire, stramineous, finely nerved, ecarinate but bearing a protruded central nerve. Inflorescence (fertile part) racemose, simple, 5–10 cm long, erect, subspirally contorted, rachis slender, nearly straight, green, glabrous, smooth, internodes 5–10 × 1–1.5 mm; floral bracts the basal ones resembling the peduncle bracts, with a subtriangular ovate base, 3–4 × 2–2.5 mm, and a slenderly filiform distal blade, 3–6 mm, about equaling to slightly shorter than the sepals, the upper ones shorter than the sepals, suberect except for the abruptly recurved distal end, ecarinate but sometimes bearing a protruded central nerve, subtriangular-ovate, stramineous, 3–5 × 2–2.5 mm, glabrous, finely acuminate-caudate to acute and apiculate, margins entire. Flowers 5–12 in number, upwardly secund forming an unilaterally oriented row, subdensely to laxly arranged, completely exposing the rachis, 27–35 mm long (including the stamens), erect, inconspicuously pedicellate, pedicel stout, ca. 2 × 2.5 mm, green; sepals symmetrical, free, erect, suboblong to subtriangular-ovate, apex acute to obtuse, ecarinate, 6–7 × 3–3.5 mm, yellowish, glabrous, thin in texture, finely nerved, entire, slightly covering each other at the base; petals symmetrical, narrowly suboblong-elliptic to narrowly subspathulate, acute to narrowly obtuse and apiculate, exappendiculate, free, 18–21 × 5–6 mm, yellow, glabrous, slightly covering each other at the base and distinctly shorter than the stamens, erect, forming a tubular calyx-like corolla; stamens distinctly exceeding the petals; filaments flat, membranaceous, white at anthesis and afterwards, completely connate to each other and forming a tube distinctly exceeding the petals, not adnate to the petals, 24–30 mm long; anthers sublinear-attenuate, 5.5–6 mm long, base bilobed to obtuse, apex acuminate, dorsifixed near the base, erect and coalescent-convergent toward the apex before anthesis, distinctly spreading-recurved at anthesis and afterwards, radially arranged, forming a stellate structure. Pistil 17–21 mm long, deeply included in the filament tube; ovary totally superior, narrowly ovate, 4–6 mm long; placentation basal; ovules obtuse; style 9–10 mm long, greenish-yellow, terete; stigma conduplicate-spiral, subcylindrical, blades 4–5 mm long, margins scalloped. Capsules unknown.</p><p>Distribution and habitat:–– Dyckia albolutea is known from the type locality only, inhabiting the county of Granja, the westernmost municipality in the state of Ceará, northeastern Brazil. It is a heliophyte and lives as a saxicole on karstic soils in metacarbonate rocky outcrops in Cerrado vegetation (fig. 10 A–D). It shares the habitat with Encholirium spectabile Martius ex Schultes &amp; Schultes f. (1830: 1233), as well as with the Euphorbiaceae Euphorbia lycioides Boissier (1860: 29), the Velloziaceae Vellozia tubiflora (A. Richard 1822: 300) Kunth (1824: 155), and Melastomataceae Comolia villosa (Aublet 1775: 334) Triana (1871 [1872]: 37).</p><p>Etymology:––The name of this new species is a reference to its flowers with the white colored, well-developed filament tube at anthesis and afterwards, in contrast with the yellow color of its corolla (fig. 10 F, G), on the basis of the Latin words albus (= white) combined with luteus (= yellow).</p><p>Distinctive characters:— Dyckia albolutea is a very unusual species by its filament tube distinctly exceeding the corolla and with a contrasting color, sharing this characteristic only with D. tubifilamentosa . It can be distinguished from the morphologically closest relative, D. tubifilamentosa, by the leaf blades inconspicuously and sparsely white-lepidote to glabrescent abaxially (vs. cinereous lepidote), sepals free (vs. connate at the base), shorter (6–7 mm vs. 8–11 cm), and yellow (vs. green), petals yellow (vs. green), filament tube longer (24–30 mm vs. 20–23 mm), white at anthesis and afterwards (vs. white at anthesis and winish or purplish colored afterwards).</p></div>	https://treatment.plazi.org/id/039E87CD8638D07B95D161ABFB80FD1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8636D07995D161ABFD2AF9D2.text	039E87CD8636D07995D161ABFD2AF9D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dyckia flabellata Leme 2025	<div><p>Dyckia flabellata Leme, sp. nov. (Fig. 11 A–M)</p><p>Diagnosis:––This new species is closely related to Dyckia estevesii Rauh (1987: 918), but can be distinguished from it by the inflorescence simple to once-branched at the base (vs. once to twice-branched), flowers longer (17–18 mm long vs. ca. 10 mm long), sepals broader (8–10 mm vs. 2–3 mm wide), and petals broadly obcordate (vs. narrowly lingulate).</p><p>Type:— BRAZIL. Mato Grosso: Alto Garças, estrada do meio para o Rio das Garças, ca. 10 km de Alto Garças, próximo da ponte, 631 m elev., 16º 52’ 17.7” S, 53º 24’ 4.0” W, 15 July 2018, W. M . Kranz B-637, fl. cult. E. Leme 9913 (holotype RB!) .</p><p>Description:–– Plant terrestrial or saxicolous, flowering ca. 1.65 m tall, propagating by basal shoots. Leaves 12– 17 in number, distichously arranged, thick-coriaceous; sheath subreniform, ca. 4 × 7 cm, castaneous near the base, green toward the distal portion, densely white lepidote, margins spinulose; blade narrowly subtriangular-attenuate, inconspicuously if at all canaliculate, suberect-arcuate, 40–50 × 2.3–3 cm at the base, ca. 4 mm thick near the base, green, opaque, abaxially distinctly and finely nerved, densely white lepidote with trichomes arranged along the nerves, adaxially densely and inconspicuously white lepidote with trichomes arranged along the nerves and not obscuring the leaf color, abaxial and abaxial surfaces slightly if at all contrasting in color, apex long spinescent-caudate, terminating in a pungent spine, margins inconspicuously white lepidote, very laxly spinose; spines 2.5–6 × 2–3 mm at the base, 2–12 cm apart, narrowly triangular, complanate, subdensely and inconspicuously white lepidote toward the base, pale castaneous near the apex, retrorse-uncinate. Peduncle lateral, erect, ca. 115 cm long, 0.6–0.7 cm in diameter, with very inconspicuously and sparsely unifilamentous white trichomes, but appearing glabrous, and a thin inconspicuous layer of white wax, greenish; peduncle bracts inconspicuously white lepidote, distinctly nerved, distinctly shorter than the internodes, the basal ones with a membranaceous, subtriangular-ovate base and a long subtriangular-lanceolate blade, acuminate and spinescent, 30–60 mm long, greenish except for the reddish margins, margins minutely spinulose at the base to crenulate or entire, the upper ones broadly subtriangular, acuminate to acute, stramineous at anthesis, erect, 15–20 × 13–17 mm, inconspicuously crenulate, ecarinate. Inflorescence (fertile part) simple to compound at the base, ca. 50 cm long; primary bracts similar to the upper peduncle bracts, distinctly shorter than the stipes of the branches; primary branches ca. 4 in number (not including the terminal one), laxly arranged, the basal ones with late development, 7–12 cm long, bearing 11–13 laxly arranged flowers, stipes distinct, 2–3 × 0.3–0.4 cm, subterete, orange-red, sparsely and inconspicuous lepidote with unifilamentous whitish to pale castaneous trichomes, but appearing glabrous; terminal branch suberect, ca. 30 cm long, with ca. 35 flowers laxly arranged toward the base and subdensely arranged near the apex, rachis 3–4 mm in diameter, straight, terete, orange-red, sparsely and inconspicuously lepidote with unifilamentous whitish to pale castaneous trichomes, but appearing glabrous; floral bracts contiguous with the flowers, equaling (basal ones) to slightly shorter than the sepals (upper ones), broadly triangular, reddish at the base and stramineous toward the apex at anthesis, 5–7.5 × 8–10 mm, nerved, sparsely and inconspicuously lepidote with unifilamentous whitish to pale castaneous trichomes but soon glabrous, acute to acuminate, margins remotely denticulate-crenulate to entire. Flowers laxly and polystichously arranged, 17–18 mm long, suberect at anthesis, odorless; pedicels inconspicuous, pale orange-yellow, glabrous, 2–2.5 mm long, ca. 6.5 mm in diameter at the apex; sepals suborbicular to subreniform, apex rounded to obtuse-emarginate, ecarinate, convex, 5–6 × 8–10 mm, reddish-orange, sparsely and inconspicuously lepidote with unifilamentous whitish to pale castaneous trichomes, margins entire but with sparsely unifilamentous inconspicuous trichomes; petals symmetric, broadly obcordate, apex obtuse-emarginate, connate at the base for ca. 2 mm in a common tube with the filaments, 13–14 × 13–14 mm, ecarinate, orange, glabrous, margins entire, glabrous, erect at anthesis and forming a tubular corolla 6–8 mm in diameter at the apex. Stamens equaling to slightly exceeding the petals by a fraction of the anthers; filaments complanate, connate for 2–3 mm above the common tube with the petals, 11–12 × 1.5–2.2 mm, pale orange; anthers narrowly ovate-lanceolate, base distinctly bilobed, apex acute, 4–5.5 mm long, strongly recurved at anthesis, slightly exceeding the petals, fixed near the base; pistil ca. 11 mm long, slightly shorter than the anthers; ovary narrowly suboblong, ca. 8 mm long; stigma conduplicate-spiral, blades ca. 2 mm long, orange, margins scalloped; style ca. 1 mm long, pale yellow. Capsules unknown.</p><p>Distribution and habitat:–– Dyckia flabellata is a saxicolous species found on more or less flat rock outcrops in fragments of Cerrado vegetation (fig. 11 A), about 630 m elevation, near the city of Garças, Mato Grosso state, central Brazil. It forms small populations composed by densely grouped individuals, being visually characterized by the flabellate leaf rosette (fig. 11 B–C).</p><p>The region where this new species was found is dominated by intense agricultural and cattle raising activities that, besides mineral exploration, constitute a human pressure that makes the future survival of the current known population of Dyckia flabellata uncertain.</p><p>Etymology:––The name of this new species is a reference to the distichous arrangement of its leaves, based on the Latin adjective flabellatus, meaning “fan-shaped”.</p><p>Distinctive characters:—The genus Dyckia Schultes &amp; Schultes f. (1830: 194) comprises 181 species (Gouda et al. cont. updated) but only four of them share leaves totally or in part distichously arranged and forming somewhat laterally flat “rosettes”, with plants being more or less prostrate.They are D. estevesii, D. mauriziae Esteves &amp; Hofacker (Pereira &amp; Hofacker, 2011: 39), D. mirandiana Leme &amp; Z.J.G. Miranda (2009: 75), and D. pottiorum Leme (Leme et al. 2012: 26) .</p><p>Dyckia flabellata is the fifth known species of the genus with distichous leaf arrangement. Most of these distichous-leaved species are found in the state of Goiás ( D. estevesii, D. mauriziae, and D. mirandiana), except for D. pottiorum from Mato Grosso do Sul state and this new species from the state of Mato Grosso, all in central Brazil. Their populations are distant from each other, in straight line, from 380 km to 1,000 km, with exception of D. flabellata and D. estevesii, with a geographically closer range, being apart from each other by 180 km in straight line.</p><p>Despite sharing the same leaf conformation, these species are not necessarily close relatives, and so the evaluation of potential new species requires an extra-carefull study in order to avoid the almost irresistible tendency to only compare it with known species that share such a distinctive attribute (Leme et al. 2012). Even considering this fact, the morphological relative of Dyckia flabellata is D. estevesii due to its size when in bloom, its tubular corolla, and the stamens equaling to slightly exceeding the petals by a fraction of the anthers. However, D. flabellata differs from it by the thick-coriaceous leaves (vs. slightly succulent), inflorescence simple to once-branched (vs. once to twice-branched), flowers longer (17–18 mm long vs. ca. 10 mm long), floral bracts larger (5–7.5 × 8–10 mm vs. ca. 2 × 2 mm), flowers more densely arranged, sepals broader (8–10 mm vs. 2–3 mm wide), petals broadly obcordate (vs. narrowly lingulate), and filaments connate for 2–3 mm above the common tube with the petals (vs. free above it).</p><p>The erect petals of Dyckia flabellata forming a tubular corolla, with exserted stamens and anthers distinctly visible, makes it clearly distinct from D. mauriziae and D. pottiorum which have suberect petals forming a subcampanulate corolla, with included stamens not distinctly visible over the petals. On the other hand, despite having tubular corolla and exserted stamens, D. mirandiana presents striking morphological differences in relation to D. flabellata, like its distinctly shorter size when in bloom (20–25 cm high), the leaf blades with much smaller marginal spines (0.5–1 mm long), the inflorescence strongly curved, flowers distinctly fragrant, sepals dark purplish-wine with an acuminate apex, and petals yellow with apiculate to subacute apex.</p></div>	https://treatment.plazi.org/id/039E87CD8636D07995D161ABFD2AF9D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8634D07C95D165DFFC88FC27.text	039E87CD8634D07C95D165DFFC88FC27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Encholirium inhaiense Leme 2025	<div><p>Encholirium inhaiense Leme, sp. nov. (Fig. 12 A–J)</p><p>Diagnosis:––This new species is closely related to Encholirium spectabile, but can be distinguished from it by the longer inflorescence (ca. 125 cm vs. 35–88 cm long), larger floral bracts (to 40 × 13 mm vs. to 22 × 6 mm), and petals smaller (8–10 × 2–2.5 mm vs. 9–22 × 4–10 mm).</p><p>Type: –– BRAZIL. Minas Gerais: Diamantina, 7.5 km a sudeste de Inhaí, 794 m elev., 17º59’13” S, 43º36’10” W, 7 August 2010, E . Leme 8436, E . Guarçoni, B . Paixão &amp; R . Oliveira (holotype RB!) .</p><p>Description:–– Plant rupiculous, clustering, flowering 2.10 m tall; Leaves ca. 35 in number, densely rosulate, strongly coriaceous, thick mainly toward the base, white-cinereous throughout; sheath much broader than the blades; blade subspreading to strongly recurved, narrowly triangular, attenuate toward the apex, ca. 75 cm long, ca. 5 cm wide at the base (not including the spines), densely white lepidote on both surfaces with trichomes obscuring the color of the blade, finely nerved mainly abaxially, nearly flat, apex long-caudate, margins subdense to laxly and coarsely spinose; spines triangular, 17–25 mm apart, flat, coarsely white lepidote except for the castaneous apex, the basal straight, 10– 12 mm long, 8–10 mm wide at the base, the upper ones distinctly antrorse-uncinate, 4–7 mm long, 4–7 mm wide at the base; peduncle 85 × 1.8–2 cm, terminal, erect, rigid, glabrous, glaucous; peduncle bracts the proximal ones foliaceous to subfoliaceous, distinctly exceeding the internodes, the median ones with a broadly ovate to subtrapeziform, entire base contrasting with a long sublinear-attenuate, spreading to recurved, laxly spinose blade, exceeding to equaling the internodes, stramineous, the distal ones ovate, long acuminate, suberect, spinose near the apex, shorter than the internodes. Inflorescence (fertile part) racemose, simple, ca. 125 cm long, ca. 6 cm in diameter at the base, ca. 3 cm in diameter near the apex, erect, rachis stout, 0.4–1.8 cm in diameter, straight, green, glabrous, strongly sulcate at anthesis; floral bracts spreading to recurved near the apex, the basal ones equaling the flowers, the upper ones shorter than the pedicels, narrowly lanceolate, green near the base, stramineous toward the apex, 9–40 × 3–13 mm, finely nerved, ecarinate, glabrous, finely acuminate, margins entire except for few remote minute spines and the inconspicuously crenulate distal portion. Flowers many, polystichously, subdensely to laxly arranged, exposing the rachis in most part, 22–30 mm long (including the stamens), subverticillate, suberect, distinctly pedicellate, pedicels clavate, 8–11 mm long, 3–5 mm in diameter at the apex; sepals symmetrical, oblong to ovate, apiculate to obtuse, ecarinate, 6–11 × 3–4 mm, green toward the base and nigrescent at the apex, soon stramineous, glabrous, distinctly nerved, irregularly crenulate mainly near the apex, not covering each other; petals symmetrical, narrowly oblong to lingulate, free, obtuse-emarginate, 8–10 × 2–2.5 mm, green except for the nigrescent-purple apex, glabrous, erect at anthesis, not covering each other and distinctly exposing the stamens, crenulate near the apex; stamens distinctly exceeding the petals; filaments terete, subfree, 11–12 × 0.8 mm; anthers sublinear, ca. 5 mm long, base distinctly bilobed, apex obtuse, dorsifixed near the base; ovary ca. 7 mm long; style ca. 8 mm long, slightly shorter than the anthers, green; stigma conduplicate-spiral, subcapitate, yellow, blades ca. 1.5 mm long, margins deeply scalloped. Capsules narrowly ovate, beaked, green, lustrous, 25–26 × 8–9 mm. Seeds orbicular, obtuse, flat, 2–2.5 mm in diameter.</p><p>Distribution and habitat:––This new species is known from the type collection only, near the district of Inhaí, county of Diamantina, Minas Gerais state, southeastern Brazil. It forms a large group of plants on more or less horizontal, blackish quartzitic rocky outcrops (fig. 12 A) in the Campos Rupestres domain, about 800 m elevation, under conditions exposed to full sun, associated to large groups of Euphorbia sp. (fig. 12 A).</p><p>Etymology:––The chosen specific epithet of this new species is a reference to the place where it was originally collected, in the district of Inhaí, located in the county of Diamantina, Minas Gerais state.</p><p>Distinctive characters:—The classical concept of Encholirium Martius ex Schultes &amp; Schultes f. (1830: 1233) adopted here follows Smith &amp; Downs (1974), Smith &amp; Read (1989) and Forzza (2005), not the proposed synonymy of Deuterocohnia Mez (1894: 506) and Encholirium under Dyckia by Gomes-da-Silva et al. (2019).</p><p>In the recent revision of the Dyckia selloa (Koch, 1873: 7) Baker (1889: 136) complex, Büneker (2021) also adopted the concept of Dyckia sensu stricto, excluding Encholirium and Deuterocohnia as synonyms. The author highlights the divergence of the results of Gomes-da-Silva et al. (2019) with the results previously obtained by Krapp et al. (2014) and Schütz et al. (2016). Büneker (2021) found that Gomes-da Silva et al. (2019) did not take into consideration the strong evidence for plastid sequestration through ancient hybridization, which should plausibly be verified through individual analyses of the plastid and nuclear marker datasets, as done by Krapp et al. (2014) and Schütz et al. (2016). These analyses could highlight incongruences in phylogenies based on nuclear and plastid data, which consequently would not allow concatenated analyses, and the verification of the possible monophyly of Deuterocohnia from these and from morphological data. Based on the considerations summarized here, Büneker (2021) assumed that there are no strong arguments supporting the proposed inclusion of Deuterocohnia – and also Encholirium – in Dyckia . In the case of Encholirium, he pointed the high evidence of paraphyly, suggesting the need of a more advanced molecular analysis to express the known outstanding morphological differences among them, which were poorly or sometimes imprecisely explored by Gomes-da-Silva et al. (2019).</p><p>The generic concept adopted here, following Gouda et al. (cont. updated) and Büneker (2021), is based on the outstanding morphological differences that clearly distinguishes these genera from each other in a simple and practical way. In addition, the classical treatment based on convincing morphological evidence fulfill the backbone of the International Code of Botanical Nomenclature expressed in its preamble, that aims at the provision of a stable method of naming taxonomic groups, avoiding and rejecting the use of names that may cause error or ambiguity or throw science into confusion. The nomenclatural confusion is already happening among researchers and can be exemplified by the implicit rejection of the synonymy proposed by Gomes-da-Silva et al. (2019) in recent articles that described new species of Encholirium (Braun 2021) or proposed news combinations, transferring to Encholirium species originally described as Dyckia (Braun &amp; Tank 2021) . A detailed analysis of the Gomes-da-Silva et al. (2019) proposition and its conflict with the current taxonomical knowledge of Bromeliaceae will be addressed in an upcoming study.</p><p>According to the identification key elaborated by Forzza (2005) in the taxonomical revision of Encholirium, this new species is related to E. spectabile (fig. 12 K–M) ‒ considering the broader concept of the species adopted by Forzza (2005) ‒ differing by the longer inflorescence (ca. 125 cm vs. 35–88 cm long), the larger floral bracts (to 40 × 13 mm vs. to 22 × 6 mm), petals smaller (8–10 × 2–2.5 mm vs. 9–22 × 4–10 mm) and narrowly oblong to lingulate (vs. elliptic). On the other hand, E. inhaiense (fig. 12 A–J) is similar to E. subsecundum (Baker, 1889: 135) Mez (1896: 540) due to inflorescence and flower conformation, but it differs from it by the much longer inflorescence (ca. 125 cm vs. 15–48 cm long), floral bracts shorter than to equaling the flowers (vs. exceeding the flowers), sepals smaller (6–11 × 3–4 mm vs. 12–18 × 5–7 mm, and petals smaller (8–10 × 2–2.5 mm vs. 15–22 × 2–4 mm), green with nigrescent apex (vs. green throughout).</p><p>Encholirium inhaiense can be also morphologically associated to E. luxor L.B. Smith &amp; Read (1989: 229) due to the rosette characteristics and inflorescence conformation, but it clearly differs by the leaf blades with longer basal spines (10–12 mm vs. 3–6 mm long), longer inflorescence (ca. 125 cm vs. 45–85 cm long), basal floral bracts equaling the flowers (vs. shorter than the pedicels), larger (9–40 × 3–13 mm vs. 5–16 × 2–4 mm), pedicels shorter (8–11 mm vs. 11–25 mm long), and petals smaller (8–10 × 2–2.5 mm vs. 19–29 × 5–9 mm) and narrowly oblong to lingulate (vs. elliptic).</p><p>Despite its distinctly larger size, this new species somewhat resembles Encholirium nibertii P.J. Braun &amp; Gastaldi (Braun 2021: 85), another inhabitant of Minas Gerais state, due to its cinereous leaves with pronounced marginal spines. However, E. inhaiense can be distinguished from it by its larger size when in bloom (ca. 2.10 m vs. 0.80–1.30 m high), larger leaf blades (ca. 75 × 5 cm vs. 13–30 × 1–2.5 cm), flowers with shorther pedicels (8–11 mm vs. 12–25 mm long), sepals narrower (3–4 mm vs. 5–6 mm wide), petals narrower (2–2.5 mm vs. 3–5 mm), green with nigrescent apex (vs. green), and longer anthers (ca. 5 mm vs. 2–3 mm long).</p></div>	https://treatment.plazi.org/id/039E87CD8634D07C95D165DFFC88FC27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8631D04295D160C3FDB2FA0B.text	039E87CD8631D04295D160C3FDB2FA0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hohenbergia kollmannii Leme & A. P. Fontana 2025	<div><p>Hohenbergia kollmannii Leme &amp; A.P. Fontana, sp. nov. (Fig. 13 A–L)</p><p>Diagnosis:—This new species is morphologically related to Hohenbergia loredanoana Leme &amp; L. Kollmann (2011: 14), but differs from it by the primary branches shorter (14–16 cm vs. 25–40 cm long), floral bracts larger (15–17 × 12–13 mm vs. 6–7 × 8–9 mm), exceeding the sepals (vs. distinctly shorter than the sepals), and by the longer petals (ca. 15 mm vs. 10–11 mm long).</p><p>Type:— BRAZIL. Bahia: Guaratinga, road Guaratinga to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.79767&amp;materialsCitation.latitude=-16.641333" title="Search Plazi for locations around (long -39.79767/lat -16.641333)">Cajuíta</a>, via <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.79767&amp;materialsCitation.latitude=-16.641333" title="Search Plazi for locations around (long -39.79767/lat -16.641333)">Jacutinga</a>, 285-350 m elev., 16º 38.48’ S, 39º 47.86’ W, 23 April 2009, E . Leme 7791, L . Kollmann, A. P . Fontana &amp; C . Esgario (holotype RB!) .</p><p>Discription:— Plants terrestrial, flowering ca. 115 cm tall. Leaves ca. 15 in number, coriaceous, forming a broadly crateriform rosette; sheath elliptic, 119–20 × 11–11.5 cm, densely pale castaneous lepidote on both sides, dark castaneous toward the base, entire; blade linear, suberect-arcuate, 60–120 × 6–7 cm, densely to subdensely and inconspicuously white lepidote on both sides but mainly abaxially with trichomes not obscuring the color of the blade, green, apex acuminate and terminating in a long somewhat pungent point, margins sparsely spinose; spines the basal ones 2–4 × 1–2 mm, narrowly triangular, dark castaneous, spreading to slightly retrorse, 7–20 mm apart, sparsely and inconspicuously spinulose toward the apex, ca. 0.5 mm long, 30–45 mm apart. Peduncle erect, stout, ca. 40 cm long, ca. 1.2 cm in diameter, greenish to red, densely white lanate; peduncle bracts narrowly lanceolate, acuminate, erect, 9–10 × 3.5 cm, exceeding the internodes, stramineous, nerved, white lanate at the base and glabrous toward the apex, entire to inconspicuously and sparsely spinulose near the apex. Inflorescence (fertile part) broadly subpyramidal, twice to 3-times branched, ca. 48 cm long, ca. 29 cm in diameter at the base, erect, rachis 0.6–1.1 cm in diameter, straight, red, subdensely white lanate; primary bracts resembling the upper peduncle bracts, spreading, distinctly exceeding the stipes; primary branches ca. 6 in number (excluding the apical portion of the inflorescence which resembles a basal primary branch), spreading, the basal ones 14–16 cm long, distinctly stipitate with stipes 2–3 × 0.8–1 cm, slightly complanate, dark red, sparsely white lanate, bearing 10 to 12 shortly stipitate to sessile secondary branches, the basal to median primary branches laxly arranged, 4–4.5 cm apart, the upper ones subdensely arranged, 0.5–1.5 cm apart, resembling the secondary branches, 2.5–5 cm long, shortly stipitate to sessile; secondary bracts subtriangular-lanceolate, narrowly acuminate, red but soon drying, 1.5–2 × 0.8–0.9 cm, equaling to shorter than the secondary fascicles, papyraceous, distinctly nerved, ecarinate, sparsely white sublanate to glabrous, subspreading to spreading with the secondary branches; secondary branches narrowly ovate, shortly stipitate to sessile, 2.5–3 × 1.2–1.7 cm, the basal ones sometimes bearing at the base 1 tertiary branches, the upper ones narrowly ellipsoid to subcylindrical, sessile, bearing 6 to 10 densely arranged flowers; tertiary bracts resembling the basal floral bracts, shorter than the branches; tertiary branches resembling the upper secondary branches but smaller, bearing 3 to 5 densely arranged flowers; floral bracts erect to suberect, exceeding the sepals, broadly subtriangular to ovate-lanceolate, dark red, 15–17 × 12–13 mm, including the apical spines, thinly coriaceous, distinctly nerved, glabrous except for the inconspicuously white sublanate margins, lustrous, carinate toward the apex (basal ones) to ecarinate (upper ones), acute with a 2–3 mm long spinescent apex, margins entire. Flowers 20–22 mm long, sessile, densely and polystichously arranged, suberect, odorless; sepals distinctly asymmetric with the membranaceous lateral wing shorter than the apex, 9 × 3–3.5 mm, including the 1.5–2 mm long apical mucro, connate at the base for ca. 3 mm, glabrous, lustrous, red, ecarinate; petals spathulate, apex acute, ca. 15 × 3 mm, free, purple, bearing 2 sublinear, dentate-digitate appendages ca. 3.5 mm above the base, without any callosities. Stamens included; filaments slightly complanate, the antepetalous ones adnate to the petals for ca. 3 mm, the antesepalous ones free; anthers oblong, base bilobed, apex apiculate, dorsifixed slightly below the middle, ca. 2.5 mm at anthesis, ca. 4 mm long before anthesis; stigma conduplicate-spiral, subcapitate, light blue, margins crenulate, ca. 2 mm long; ovary subglobose, terete, ca. 4 mm long, ca. 4.5 mm in diameter, whitish, glabrous; placentation apical; ovules long apiculate; epigynous tube infundibuliform, ca. 1 mm long. Fruits unknown.</p><p>Distribution and habitat:— Hohenbergia kollmannii shares the same type locality with Cryptanthus cajuitensis described above, as well as with Neoregelia retrorsa, in the county of Guaratinga, along the way to the locality of Cajuíta. It is a typical Atlantic Forest inhabitant in the southern region of Bahia state, northeastern Brazil, where sparsely distributed individuals were found as a terrestrial, in the shaded forest floor of steep whatersheds (fig. 13 A–B). The forest at the type spot is characterized by the presence of a large formation of Syagrus sp. ( Arecaceae), known as “licuri”.</p><p>Etymology:—The epithet for this new species honors the botanist Ludovic Jean Charles Kollmann, a specialist in Begoniaceae, Bromeliaceae, and Orchidaceae, of the National Institute of the Atlantic Forest, in Santa Teresa, Espírito Santo state, who is responsible for the discovery and description of numerous new species of the Atlantic Forest biome.</p><p>Distinctive characters:— Hohenbergia kollmanii (fig. 13 A–L) is morphologically related to H. loredanoana (fig. 13 M–N) due to its large size when in bloom and the structure and red color of the inflorescence, but differs from it by the primary branches shorter (14–16 cm vs. 25–40 cm long), floral bracts broadly subriangular to ovate-lanceolate (vs. suborbicular), larger (15–17 × 12–13 mm vs. 6–7 × 8–9 mm), exceeding the sepals (vs. distinctly shorter than the sepals), the basal ones carinate toward the apex (vs. ecarinate), flowers longer (20–22 mm vs. 14–15 mm long), sepals longer (ca. 9 mm vs. 4.5–5 mm long) with the lateral wing distinctly shorter than the apex (vs. exceeding the apex) and with a longer apical mucro (1.5–2 mm vs. ca. 0.5 mm long), petals longer (ca. 15 mm vs. 10–11 mm long), and stigma light blue (vs. dark blue).</p><p>Based on its large size when in bloom and the structure and red color of the inflorescence, this new species is also related to Hohenbergia sandrae Leme (2003: 174) (fig. 13 O–P), but can be distinguished from it by the leaf blades narrower (6–7 cm vs. 9–10 cm long), primary branches shorter (14–16 cm vs. 22–32 cm long), floral bracts broadly subriangular to ovate-lanceolate (vs. subreniform to suborbicular), larger (15–17 × 12–13 mm vs. ca. 5 × 8 mm), exceeding the sepals (vs. equaling to slightly exceeding the ovary), the basal ones carinate toward the apex (vs. ecarinate), flowers longer (20–22 mm vs. ca. 14 mm long), sepals longer (ca. 9 mm vs. ca. 5 mm long) with the apex distinctly spinescent (vs. muticous to remotely apiculate), petals longer (ca. 15 mm vs. ca. 10 mm long), acute (vs. obtuse), and stigma light blue (vs. white).</p></div>	https://treatment.plazi.org/id/039E87CD8631D04295D160C3FDB2FA0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD860FD04095D166A7FD96F9F7.text	039E87CD860FD04095D166A7FD96F9F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthophytum afonsoclaudense Leme, D. R. Couto & Fraga 2025	<div><p>Orthophytum afonsoclaudense Leme, D.R. Couto &amp; Fraga, sp. nov. (Fig. 14 A–I)</p><p>Diagnosis:—This new species is morphologically related to Orthophytum estevesii (Rauh [Rauh &amp; Gross, 1991: 29]) Leme (2004: 37), but differs by its larger size when in bloom (45–48 cm vs. 12–30 cm tall), leaf blades distinctly white lepidote (vs. mostly glabrous), longer peduncle (38–39 cm vs. 6–17 cm), petals greenish (vs. white toward the apex), and green stigma (vs. white).</p><p>Type:— Brazil. Espírito Santo: Afonso Claudio, Serra Pelada, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.075027&amp;materialsCitation.latitude=-20.01" title="Search Plazi for locations around (long -41.075027/lat -20.01)">Palmital</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.075027&amp;materialsCitation.latitude=-20.01" title="Search Plazi for locations around (long -41.075027/lat -20.01)">Pedra do Sol</a>, propriedade da família Brandemburg, 760 m elev., 20°00’36” S, 41°04’30.1” W, 20 October 2019, D. R . Couto 4892 &amp; C . N . Fraga, fl. cult. E . Leme 9740 (holotype RB!) .</p><p>Discription:— Plants saxicolous, stemless, 45–48 cm tall when flowering, propagating by a shoot originated from the inflorescence apex. Leaves 12–15 in number, subspreading recurved at anthesis, subdensely disposed; sheath subtrapeziform, ca. 1.7 × 4 cm, densely and coarsely white lepidote toward the distal end; blade narrowly lanceolate, 20–23 × 3–3.2 cm, thickly coriaceous, ca. 2.5 mm thick near the base, slightly canaliculate, green, abaxially completely covered by a dense layer of coarse white trichomes completely obscuring blade color, distinctly nerved, adaxially densely to subdensely and coarsely white lepidote toward the base with trichomes disposed in longitudinal rows not obscuring blade color, glabrescent toward the apex, apex attenuate-caudate, margins subdensely to laxly spinose; spines triangular, straight or antrorsely uncinate, glabrous, the basal ones 2.5–3 × 1.5 mm, 3–4 mm apart, the upper ones 1–1.5 × 1 mm, 6–8 mm apart. Inflorescence (fertile part) pseudosimple, densely subcapitate-rosulate, 5–5.5 cm long, 4.5–5.5 cm in diameter, with 12–16 densely arranged flowers; peduncle 38–39 × 0.6–0.9 cm, green, densely white lanate mainly toward the base, nearly straight to flexuous at the distal end; peduncle bracts foliaceous, laxly arranged, not concealing the peduncle, spreading-recurved; primary bracts resembling the floral bracts, distinctly exceeding the inconspicuous fascicles; fascicles inconspicuous or abortive, 1–2 in number, nearly sessile, ca. 2-flowered; floral bracts strongly recurved, exceeding the sepals, ecarinate to obtusely carinate mainly the apical ones, o v ate to ovate-lanceolate, yellowish-green, 30–35 × 15–22 mm, thinly coriaceous, finely nerved, glabrous, acuminate, margins densely spinose, spines 1–1.2 mm long, uncinate, irregularly curved. Flowers sessile, 35–37 mm long (with extended petals), odorless; sepals narrowly triangular - lanceolate, attenuate-caudate, erect, thin in texture, 20–22 × 4 mm, free, pale green, glabrous, entire or the adaxial ones inconspicuously crenulate at the apex, the abaxial one ecarinate, the adaxial ones distinctly carinate with the keel decurrent on the ovary; petals narrowly spathulate, free, apex obtuse-emarginate and slightly cucullate, 26–29 × 4–5 mm, greenish throughout, erect at anthesis, bearing 2 longitudinal callosities slightly exceeding the antepetalous filaments; petal appendages echinatiform, ca. 4 mm above the base, with multi-oriented, irregularly fimbriate-digitate prolongations; filaments unequal in length, the antesepalous ones ca. 20 mm long, free, the antepetalous ones ca. 15 mm long, adnate to the petals for ca. 10 mm; anthers 2–2.5 mm long, dorsifixed at the middle, base and apex obtuse, slightly complanate at anthesis; ovary 6–7 × 7 mm, whitish, glabrous except for the inconspicuous long fimbriate white trichomes at the base, trigonous, sharply carinate laterally; epigynous tube ca. 1 mm long, crateriform; stigma simple-dilated, lobes broadened, lip-shaped, spreading, ca. 0.5 mm long, green, margins densely papillate; placentation apical; ovules many, obtuse. Fruits unknown.</p><p>Distribution and habitat:— Orthophytum afonsoclaudense is known from the type collection only, within a private farm, in Afonso Claudio municipality, in the locality of Palmital, Serra Pelada district, in the central region of Espírito Santo state, southeastern Brazil. This new species grows on bare, slightly inclined granitic/gneiss rocky outcrops, on Humic Litholic Neosols accumulated in shallow depressions, which are surrounded by exposed rocks and inselberg vegetation, forming islands of dense groups of plants under full sunlight.</p><p>The known populations of this new species are located between 650 and 760 m elevation and are greatly degraded by agricultural activities. Despite the presence of some small, disturbed fragments of Atlantic Forest, the inselbergs where it grows are mostly surrounded by small farms and coffee plantations and are under pressure from the rock mining industry.</p><p>Etymology:—The name chosen for this new species is a reference to the county of Afonso Claudio, state of Espírito Santo, southeastern Brazil, where this new species was discovered.</p><p>Distinctive characters:— Orthophytum afonsoclaudense is morphologically related to O. estevesii because of its general appearance when in bloom and the conformation and color of the inflorescence. However, it differs by its larger size when in bloom (45–48 cm vs. 12–30 cm), longer leaf blades (20–23 cm vs. 9–20 cm), which are broader (3–3.2 cm vs. 1.5–2.2 cm) and distinctly white lepidote (vs. mostly glabrous). The trichomes on the adaxial surface and toward the base are disposed in longitudinal rows not obscuring the blade color. This new species also presents a longer peduncle (38–39 cm vs. 6–17 cm), petals greenish (vs. white toward the apex) with obtuse-emarginate and slightly cucullate apex (vs. rounded), as well as a green stigma (vs. white).</p><p>The peculiar disposition in rows of the white trichomes on the adaxial surface of the leaf blades of Orthophytum afonsoclaudense, despite less pronounced, is similar to the pattern observed in O. striatifolium Leme &amp; L. Kollmann (2007: 152) . However, this species can be easily distinguished from it mainly by its larger size when in bloom (45–48 cm vs. 10–13 cm tall), larger leaf blades (20–23 × 3–3.2 cm vs. 8–11 × 1.4–1.5 cm), inflorescence with more flowers (12–16 vs. 6–9 in number), longer sepals (20–22 mm vs. 15–16 mm long), and petals greenish (vs. white) and erect at anthesis (vs. spreading-recurved at anthesis).</p></div>	https://treatment.plazi.org/id/039E87CD860FD04095D166A7FD96F9F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD860DD04695D165F3FCF1F9F7.text	039E87CD860DD04695D165F3FCF1F9F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orthophytum pseudoestevesii Leme 2025	<div><p>Orthophytum pseudoestevesii Leme, sp. nov. (Fig. 15 A–L)</p><p>Diagnosis:—This new species is morphologically close to Orthophytum estevesii but differs by its larger size when in bloom (ca. 70 cm long vs. 12–30 cm long), long caulescent habit (vs. stemless), inflorescence densely subcylindrical-rosulate (vs. capitate-rosulate), and by the shorter sepals (14–15 mm vs. 18–20 mm long).</p><p>Type:— Brazil. Bahia: Guaratinga, entre Buranhém e a divisa com Minas Gerais, ca. 300 m elev., August 2019, J . E. dos Santos s.n., fl. cult. E. Leme 9712 (holotype RB!) .</p><p>Description:— Plants saxicolous, long caulescent, ca. 70 cm tall when flowering, stem prostrate, ca. 30 cm long, 0.7–0.8 cm in diameter, propagating by shoots at the base of the stem and near the proximal portion of the peduncle. Leaves ca. 28 in number, equally arranged along the stem, subspreading at anthesis, subdensely disposed; sheath inconspicuous, subtrapeziform, ca. 1.2 × 2.1 cm, sparsely and inconspicuous white lepidote toward the distal end to glabrescent; blade narrowly subtriangular-lanceolate, 15–20 × 1.4–1.7 cm, soft in texture but slightly thick, ca. 1.5 mm thick near the base, flat or nearly so, green, abaxially near the base inconspicuously white lepidote, glabrous toward the apex, distinctly nerved, adaxially glabrous, apex attenuate-caudate, margins subdensely spinose; spines triangular, antrorsely uncinate, glabrous, 0.3–0.5 mm long, 3–7 mm apart. Inflorescence (fertile part) pseudosimple, elongate mainly at the end of anthesis, densely subcylindrical-rosulate, ca. 18 cm long, 4.5–5 cm in diameter, with ca. 35 densely arranged flowers; peduncle ca. 17 × 0.4 cm, green, glabrous, nearly straight, suberect but becoming prostrate with the increasing height during the development of the inflorescence; peduncle bracts foliaceous, laxly arranged, not concealing the peduncle, spreading-recurved, the upper ones resembling the basal floral bracts; primary bracts resembling the basal floral bracts; fascicles abortive, rarely 1 in number; floral bracts strongly recurved, exceeding the sepals, ecarinate to obtusely carinate mainly the apical ones, subtriangular-ovate to ovate-lanceolate, green to light green, 27–32 × 15–19 mm, thinly coriaceous, finely nerved, glabrous, acuminate, margins densely spinose, spines 0.5–1 mm long, straight to antrorsely uncinate. Flowers sessile, ca. 23 mm long (with extended petals), odorless; sepals narrowly lanceolate, attenuate-caudate, erect, thin in texture, 14–15 × 3.5 mm, free, pale green, glabrous, entire, the abaxial one ecarinate, the adaxial ones distinctly carinate with the keel decurrent on the ovary; petals narrowly spathulate, free, apex obtuse and slightly cucullate, ca. 20 × 4 mm, white, apical portion subspreading at anthesis, bearing 2 longitudinal callosities slightly exceeding the antepetalous filaments; petal appendages echinatiform, ca. 1 mm above the base, with spreading or nearly so irregularly fimbriate-digitate prolongations; filaments unequal in length, the antesepalous ones ca. 14 mm long, free, the antepetalous ones ca. 11 mm long, adnate to the petals for 7–8 mm; anthers 2–2.5 mm long, dorsifixed in the middle, base obtuse, apex obtuse and inconspicuously apiculate, slightly complanate at anthesis; ovary ca. 4 × 4.5 mm, white, glabrous, trigonous, carinate laterally; epigynous tube ca. 0.5 mm long, crateriform; stigma simple-dilated, lobes broadened, lip-shaped, spreading, ca. 0.7 mm long, white, margins sparsely long-papillate; placentation apical; ovules many, obtuse. Fruits unknown.</p><p>Distribution and habitat:–– Orthophytum pseudoestevesii grows as a saxicole in shallow soils on horizontal rocky outcrops at the top of inselbegs, or on its nearly bare steep rocky slopes, in the district of Buranhém, county of Guaratinga, Bahia state, northeastern Brazil, near the border with Minas Gerais state, in southeastern Brazil. It is a heliophile that forms small group of plants scattered in the area, under full exposed condition or sometimes partially protected under thin bushes.</p><p>Despite the area where this new species was found is a difficult-to-reach one, it suffers periodical fires, which represents an important threat to its survival. On the other hand, the absence of data on its occurrence does not allow to accurately point out its current conservation status.</p><p>Etymology:––The Greek prefix “pseudo” is used to mark something that appears to be one thing but is something else. It indicates here that this new species is a “false” Orthophytum estevesii, its likely closest morphological relative, and can be easily confused with it, mainly at the beginning of the anthesis, when its inflorescence is not at all distinctly elongate. These species share very similar general color pattern mainly of the bracts and petals, despite the presence of important morphological differences.</p><p>Distinctive characters:— Orthophytum pseudoestevesii is morphologically related to O. estevesii and can be confused with it due to the similar color pattern of the inflorescence and flower conformation. However, this new species can be distinguished by its larger size when in bloom (ca. 70 cm long vs. 12–30 cm tall), the long caulescent habit (vs. stemless), and greater number of leaves (ca. 28 vs. 7–10 in number). On the other hand, its inflorescence is elongate and densely subcylindrical-rosulate mainly at the end of anthesis (vs. capitate-rosulate), being distinctly longer (ca. 18 cm vs. 2.5–3 cm long). The floral bracts have shorter marginal spines (0.5–1 mm vs. 1.5–2.5 mm long), and the sepals are shorter (14–15 mm vs. 18–20 mm long).</p></div>	https://treatment.plazi.org/id/039E87CD860DD04695D165F3FCF1F9F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD860BD04495D165F3FC83F947.text	039E87CD860BD04495D165F3FC83F947.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pitcairnia kranziana Leme 2025	<div><p>Pitcairnia kranziana Leme, sp. nov. (Fig. 16 A–N)</p><p>Diagnosis:––This new species is closely related to Pitcairnia torresiana L.B. Smith (1952: 4), but differs from it by leaves monomorphic (vs. dimorphic), the outer ones narrowly triangular-lanceolate (vs. filiform), and leaf blades broader (12–18 mm vs. 5–6 mm wide), appearing glabrous but with sparse and inconspicuous glandulose trichomes (vs. vestite beneath with linear twisted trichomes).</p><p>Type: –– BRAZIL. Tocantins: Rio dos Bois, Paredão do Urubu, em afloramento no <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.5445&amp;materialsCitation.latitude=-9.225249" title="Search Plazi for locations around (long -48.5445/lat -9.225249)">Cerrado</a>, 272 m elev., 9°13’30.9” S, 48°32’40.2” W, 9 July 2021, W. M . Kranz B-740, cult. E . Leme 10046 (holotype RB!) .</p><p>Description:–– Plant saxicolous, forming sparse groups, propagating by short shoots, flowering ca. 40 cm tall; Leaves ca. 8 in number, fasciculate, suberect-arcuate, monomorphic, thin in texture, at least the basal ones deciduous along a straight, transverse line resulting in an enrolled leaf base; sheath inconspicuous, entire; blade the basal ones very short and contrasting with the much longer inner ones, narrowly triangular-lanceolate, acuminate, castaneous toward the base abaxially, green toward the apex, entire, the inner ones sublinear-lanceolate, V-shaped with a narrow central channel, narrowed toward the base but not petiolate, green, distinctly nerved mainly abaxially, 30–60 × 1.2–1.8 cm, with subdense to sparse and inconspicuous glandulose white trichomes mainly abaxially and including the margins, but appearing glabrous, apex attenuate-caudate, margins slightly undulate; peduncle erect, ca. 30 cm long, 0.3-0.4 cm in diameter, glabrous, green or greenish; peduncle bracts the basal ones foliaceous to subfoliaceous and distinctly exceeding the internodes, the upper ones narrowly lanceolate, attenuate-caudate, erect, exceeding to shorter than the internodes, glabrous, entire, green. Inflorescence (fertile part) racemose, simple, erect, shorter than the leaves, ca. 5 cm long, rachis straight, 2–3 mm in diameter, subangulose, greenish to rose, glabrous. floral bracts suberect, shorter to slightly exceeding the pedicels, narrowly lanceolate, green, 6–15 × 1.5–2.5 mm, thin in texture, with sparse and inconspicuous uniseriate white trichomes and sparse and inconspicuous white glandulose trichomes mainly near the margins and at the base, acuminate-caudate, margins entire. Flowers ca. 20 in number, 58–60 mm long (including the pedicels), subdensely to laxly arranged, suberect to nearly erect at anthesis, slightly if at all secund, diurnal, odorless; pedicel 10–18 mm long, 1–1.5 mm in diameter, terete, orange, with inconspicuous and sparse white glandulose trichomes but appearing glabrous, suberect to nearly spreading at anthesis, forming an internal angle of ca. 135° with the axis; sepals narrowly ovate-lanceolate or lanceolate, apex narrowly acute, ecarinate or inconspicuously if at all carinate, 15 × 4.5–5.5 mm, yellow at the base and orange toward the apex, with inconspicuous and sparse white glandulose trichomes but appearing glabrous; petals lanceolate, apex acute, 43 × 10.5–11.5 mm, orange, erect except for the slightly suberect apex at anthesis, exposing the anthers, convergent over the stamens forming a zygomorphic corolla, with very sparse and inconspicuous glandulose white trichomes on both surfaces but appearing glabrous, bearing at the base a single appendix, ca. 4 × 2.5 mm, dentate or subdentate, adnate to the petals for ca. 3 mm; stamens shorter than the petals; anthers linear, ca. 6 mm long, fixed near the base, base distinctly bilobed, apex acute and inconspicuously apiculate; filaments terete, yellow, free, with inconspicuous and sparse glandulose trichomes, 31–33 mm long, ca. 0.6 mm in diameter; stigma subcapitate, conduplicate-spiral, slightly exceeding the petals, completely exposed at anthesis, yellow, margins densely papillose; ovary 1/2 superior; ovules many, caudate.</p><p>Distribution and habitat:–– Pitcairnia kranziana is known from the type locality only. It was discovered in the county of Rio dos Bois, in the central-north region of the state of Tocantins, central Brazil, where the Cerrado vegetation prevails. It was found growing as a saxicole on a more or less vertical rock outcrop, in a shaded site in the Cerrado vegetation (fig. 16 A).</p><p>Etymology:––The name of this new species honors its collector, the Brazilian agronomist Walter Miguel Kranz from Paraná state, who has contributed significantly to the knowledge of Bromeliaceae by the introduction into cultivation of countless bromeliad species, most of them of Pitcairnioideae, which includes many new species.</p><p>Distinctive characters:— Pitcairnia kranziana (fig. 16 A–N) is morphologically related to P. torresiana (fig. 16 O–Q) due to its similar stature, color and structure of the inflorescence, and flower conformation. Nonetheless, it differs from it by its monomorphic leaves (vs. dimorphic), the outer ones narrowly triangular-lanceolate (vs. filiform), leaf blades broader (12–18 mm vs. 5–6 mm wide), appearing glabrous but with sparse and inconspicuous glandulose trichomes (vs. vestite beneath with linear twisted trichomes), peduncle glabrous (vs. flocculose-lepidote), petals orange (vs. red), and ovary 1/2 superior (vs. 2/3 superior).</p><p>This new species also resembles the enigmatic Pitcairnia ensifolia Mez (1894: 436), which is typified by its original description and the informative plate since its type specimen deposited in Wien (W) was destroyed during the World War II. However, P. kranziana can be distinguished from it by its monomorphic leaves (vs. dimorphic), leaf blades entire (vs. serrulate), appearing glabrous but with sparse and inconspicuous glandulose trichomes (vs. furfuraceous), sepals ecarinate or slightly if at all carinate (vs. alate-carinate), petals orange (vs. red), and ovules caudate (vs. ecaudate). On the other hand, P. kranziana can be also morphologically associated to P. irwiniana L.B. Smith (1966: 153), differing by its monomorphic leaves (vs. dimorphic), leaf blades entire (vs. serrulate), appearing glabrous but with sparse and inconspicuous glandulose trichomes (vs. white lepidote), sepals ecarinate or slightly if at all carinate (vs. alate-carinate), and ovules caudate (vs. apiculate).</p></div>	https://treatment.plazi.org/id/039E87CD860BD04495D165F3FC83F947	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8609D04B95D16462FA79FD1F.text	039E87CD8609D04B95D16462FA79FD1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Portea santosiana Leme & E. H. Souza 2025	<div><p>Portea santosiana Leme &amp; E.H. Souza, sp. nov. (Fig. 17 A–R)</p><p>Diagnosis:––This new species differs from its close relative, Portea grandiflora Philcox (1992: 276), by the upper peduncle bracts involucral and densely aggregated below the inflorescence (vs. not involucral), inflorescence subcorymbose-capitate with the rachis short and not visible (vs. elongate, axis visible at least in part), and sepals with a shorter apical mucro (ca. 2.5 mm vs. 4–5 mm).</p><p>Type: –– BRAZIL. Bahia: Guaratinga, BA-686, ca. 7 km from Pedra do Oratório to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.784416&amp;materialsCitation.latitude=-16.645971" title="Search Plazi for locations around (long -39.784416/lat -16.645971)">Cajuíta</a>, locality of “Serra”, 361 m elev., 16º38’45.5” S, 39º47’03.9” W, August 2023, leg. J . E. dos Santos s.n., cult. E. Leme 10300 (holotype RB!) .</p><p>Description:–– Plant terrestrial, flowering 70–80 cm tall. Leaves ca. 20 in number, rosulate, suberect-arcuate, coriaceous, forming a broadly funnelform rosette; sheath broadly elliptic-ovate, 17–19 × 12–13 cm, densely and minutely castaneous lepidote on both sides, abaxially castaneous toward the base, adaxially purple near the distal end; blade ligulate, not narrowed toward the base, 40–50 × 5.8–6.7 cm, completely green or bronze colored near the apex, densely white lepidote abaxially, inconspicuously white lepidote adaxially, apex acute to subrounded and apiculate, margins subdensely spinose; spines dark brown to nigrescent, triangular, flat, straight to slightly antrorse, 1–2 mm long, 0.7–1 mm wide at the base, 5–10 mm apart. Peduncle erect, ca. 50 cm long, 1–1.2 cm in diameter, rose to red, white sublanate; peduncle bracts the basal ones subfoliaceous, covering the peduncle, the upper ones involucral and densely aggregated below the inflorescence, elliptic to obovate, subacute and distinctly apiculate, exceeding the outer branches, 5.5–7.5 × 3–4 cm, erect to suberect except for the suspreading distal part, nerved, densely and minutely spinulose near the apex, entire toward the base, subdensely white lepidote, distinctly exceeding the internodes, red, papyraceous. Inflorescence (fertile part) 2 to 3-times branched, densely subcorymbose-capitate, shorter to equaling the leaves, erect, ca. 8 cm long (fertile part, including the petals), ca. 7 cm in diameter (excluding the involucral bracts); rachis ca. 1 cm in diameter, short and covered by the branches, white sublanate, rose; primary bracts resembling the involucral bracts but smaller, narrowly obovate to oblanceolate, acute and apiculate, inconspicuously spinulose at the apex to subentire, nerved, subdensely white lepidote with fimbriate trichomes mainly abaxially and along the apical margins, red, thin in texture, suberect with the branches, abruptly decreasing in the inner branches, 2–5 × 0.7–2.5 cm, to equaling the sepals; primary branches ca. 10 in number, densely polystichously arranged, inconspicuous, erect, the basal ones 3.5–4 × 3.5–4.5 cm (excluding the petals), ca. 10-flowered, the basal ones bearing 1–2 secondary branches densely arranged; stipes very short but stout, ca. 3 × 7 mm, ebracteate, complanate, rose, subdensely white sublanate with fimbriate trichomes; rachis inconspicuous, rose, white sublanate; secondary bracts resembling the floral bracts; secondary branches ca. 3 × 2 cm (excluding the petals), inconspicuous, subsessile, ca. 3-flowered; rachis not visible, white sublanate, rose, ca. 3 mm in diameter, slightly complanate; floral bracts inconspicuous, shorter than to equaling the pedicels, ecarinate, triangular, whitish except for the castaneous spinescent apex, 2–4 × 3–4 mm, thin in texture, white sublanate, apex spinescent, margins entire. Flowers 47–50 mm long, odorless, densely and subdistichously arranged, inconspicuously pedicellate; pedicels stout, indistinct, merging into the ovary, 3–4 × 2.5–3 mm; sepals ca. 16 mm long, strongly asymmetrical with the lateral membranaceous rounded wing ca. 9 mm wide, exceeding the apical spine and distinctly projected laterally, lilac toward the apex, sparsely and inconspicuously white lepidote to glabrescent, ecarinate, connate at the base for ca. 8 mm, apical mucro ca. 2.5 mm long, pale castaneous, erect to slightly curved; petals narrowly spathulate, 32–37 × 7 mm, obtuse, slightly cucullate, free, white at the base and lilac toward the apex, erect except for the slightly suberect apex, at the base bearing 2 cuneate, denticulate appendages of 2– 2.5 × 1.5 mm, as well as 2 conspicuous longitudinal callosities slightly shorter than the filaments; filaments ca. 27 mm long, slightly complanate, slightly dilated and lilac toward the distal end, the antesepalous ones free, the antepetalous ones basally adnate to the petals for ca. 20 mm; anthers narrowly oblong ca. 5.5 mm long, dorsifixed slightly above the middle, base and apex obtuse, pale lilac; stigma conduplicate-spiral, subglobose, ca. 2.4 × 2.2 mm, lilac, margins entire, with an inconspicuous but dense fringe of white filamentous trichome-like structure before anthesis, at anthesis the fringe collapses and is melting becoming a translucent substance; ovary clavate, ca. 14 mm long, ca. 5 mm in diameter at the apex, sparsely and inconspicuously white lepidote to glabrescent, whitish; epigynous tube crateriform, ca. 2 mm long; placentation from median to apical; ovules caudate. Fruits unknown.</p><p>Distribution and habitat:–– Portea santosiana is known from its type locality only, which is situated about 7 km from Pedra do Oratório, situated nearby the city of Guaratinga, along the road toward Cajuíta, in the locality known as “Serra”, southern Bahia state, northeastern Brazil. It grows as a terrestrial, forming large and dense groups of plants (fig. 17 B–C) in a low-elevated area (about 360 m elevation) covered by disturbed Atlantic Forest. This forest fragment is situated at the foothill (fig. 17 A) of a granitic inselberg alongside the road, which represents an additional threat to the survival of the known population of this apparently endemic species.</p><p>Etymology:––The name chosen for this new species honors its collector, the bromeliad collector João Eduardo dos Santos, responsible for the preservation of an important fragment of Atlantic Forest in the county of Itapebi, Bahia state, from where he presented new species of different botanical families to the scientific community.</p><p>Distinctive characters:— Portea Brongniart ex K. Koch (1856: 368) is a small bromelioid genus endemic to Brazil that comprises eight recognized species mostly inhabiting Atlantic Forest habitats, but also encountered in Restinga and Campos Rupestres vegetation. Five of them grow in northeastern state of Bahia ‒ its center of diversity ‒, two in Rio de Janeiro state, two in Espírito Santo state, and one in Minas Gerais state, southeastern Brazil .</p><p>The genus is characterized by small to large-sized plants with branched inflorescence, large and colorful peduncle and primary bracts, short to long-pedicellate flowers, distinctly connate and strongly asymmetric sepals with a very large lateral wing, sometimes larger than the main erect part of the sepal itself. The free petals are lilac or purple, with short basal appendages. The large, capitate, conduplicate-spiral stigma presents entire margins bearing an inconspicuous to distinct, dense, white, and very delicate filamentous trichome-like fringe only before anthesis (about two days before anthesis or earlier; see fig. 17 Q; see also fig. 17 O, in Leme et al. 2021), which collapses and melts, becoming a translucent or sometimes milky substance at anthesis (fig. 17 P). The pollen is polyporate (see fig. 19 L, in Leme et al. 2021) and the lilac to purple large fruits (see fig. 21 U, in Leme et al. 2021) hold long caudate seeds (see fig. 25 Q, in Leme et al. 2021). Portea santosiana presents all typical features listed for the genus, except for the unusual, dense subcorymbose, head-like inflorescence ornamented by showy involucral bracts and the stout and indistinct pedicels, merging into the ovary, which distinguishes it from all known species (fig. 17 D).</p><p>Portea grandiflora, another species endemic to Bahia state, is its close morphological relative due to the rosette conformation and the color pattern of the inflorescence and flowers. However, P. santosiana differs from it by the upper peduncle bracts involucral and densely aggregated below the inflorescence (vs. not involucral), inflorescence subcorymbose-capitate with the rachis short and not visible before and after anthesis (vs. elongate, axis visible at least in part), sepals with a shorter apical mucro (ca. 2.5 mm vs. 4–5 mm), petals shorter and broader (32–37 × 7 mm vs. 35– 45 × 5 mm), and ovary sparsely and inconspicuously white lepidote (vs. densely white floccose), whitish (vs. rose).</p></div>	https://treatment.plazi.org/id/039E87CD8609D04B95D16462FA79FD1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8606D04995D161ABFE42FB2A.text	039E87CD8606D04995D161ABFE42FB2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tillandsia montezumensis Leme & W. Till 2025	<div><p>Tillandsia montezumensis Leme &amp; W. Till, sp. nov. (Fig. 18 A–F)</p><p>Diagnosis:––This new species can be distinguished from its likely closest relative, Tillandsia tenuifolia Linnaeus (1759: 286), by the leaf blades completely covered by a dense coat of white trichomes obscuring the color of the blades (vs. densely and minutely lepidote with trichomes not obscuring the color of the blades) and longer (14–17 cm vs. 5–10 cm long), and sepals longer (ca. 16 mm vs. ca. 10 mm long).</p><p>Type: –– BRAZIL. Minas Gerais: Montezuma, Brejinho, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.567497&amp;materialsCitation.latitude=-15.045" title="Search Plazi for locations around (long -42.567497/lat -15.045)">Fazenda Brejinho</a> (prox. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.567497&amp;materialsCitation.latitude=-15.045" title="Search Plazi for locations around (long -42.567497/lat -15.045)">Faz. Alge da Savana</a>), 980 m elev., 15º02’42” S, 42º34’03” W, 27 May 2010, E . Leme 8332 &amp; R . Oliveira (holotype RB!) .</p><p>Description:–– Plants epilithic, flowering 20–25 cm tall, stemless to shortly caulescent, propagating by elongate shoots developed at the base of the basal portion of the rosette, forming dense groups of plants. Leaves ca. 30 in number, densely subfasciculate, without impounding capacity; sheath inconspicuous, subtrapeziform, corrugate abaxially, 1 × 0.7–0.9 cm, completely covered on both surfaces by a dense coat of white, appressed trichomes; blade narrowly triangular, 14–17 × 0.3–0.5 cm (near the base), suberect, not secund, not narrowed at the base, subrigid toward the base, pliable toward the apex, completely covered on both sides by a dense layer of white appressed trichomes obscuring the blade color, apex filiform-caudate. Peduncle slender, ca. 8 cm long, ca. 0.2 cm in diameter, erect, glabrous, whitish; peduncle bracts the basal ones foliaceous, the upper ones with a narrowly ovate-lanceolate base 15–20 × 4 mm, partially enfolding the peduncle, reddish-rose, glabrous toward the base, and a filiform distal blade 12–30 mm long, erect, densely white lepidote, distinctly exceeding the internodes. Inflorescence (fertile part) simple, 30–35 mm long, erect, 3–5-flowered, slightly shorter than to slightly exceeding the leaves; rachis slender, flexuous, pale rose, glabrous, internodes ca. 5 mm long; floral bracts equaling to slightly shorter than the sepals, ecarinate, narrowly ovate-lanceolate, reddish-rose, 15–17 × 5–6 mm, membranaceous, thinly nerved, glabrous except for the densely white lepidote apex, apex shortly caudate (basal ones) to acute (apical ones). Flowers diurnal, odorless, subpolystichous, densely to subdensely arranged, erect, 23–24 mm long with extended petals, pedicels inconspicuous, ca. 1.5 mm long greenish, glabrous; sepals narrowly lanceolate, symmetrical, erect, apex long acuminate, 16 × 3.5–4.5 mm, glabrous, rose, thin in texture, the abaxial one free, ecarinate, the adaxial ones connate at the base for 3.5–5 mm, carinate toward the apex; petals narrowly spathulate with narrow proximal portion in contrast with the broader distal portion, apex rounded, 21 × 3–3.5 mm, white, free, erect and forming basally a subtubular corolla except for the strongly recurved distal portion, exappendiculate; stamens distinctly shorter than the petals; filaments complanate, plicate in the middle, white, free, ca. 0.8 mm wide; anthers narrowly oblong, base distinctly bilobed, apex subacute, ca. 2 mm long, dorsifixed near the base; style distinctly exceeding the anthers, ca. 12 mm long; ovary totally superior; stigma apparently conduplicate, blades ca. 0.5 mm long; ovules caudate. Capsules unknown.</p><p>Distribution and habitat:––The type of Tillandsia montezumensis was found in the locality of Brejinho, county of Montezuma, near the border with Bahia, in the microregion of Salinas, north of Minas Gerais state, in the Espinhaço range, southeastern Brazil. It grows as an epilithic species on quartzitic outcrops (fig. 18 B), forming small group of plants scattered in the Campos Rupestres that prevails in the area (fig. 18 A), which is in most part well conserved due to the difficult access and the existing Montezuma State Park. It is also present in the county of Presidente Kubitschek, in Minas Gerais state.</p><p>However, the absence of data on its occurrence does not allow to accurately point out its current conservation status.</p><p>Etymology:––The name chosen for this new species is a explicit reference to the county of Montezuma, Minas Gerais state, where the type was found.</p><p>Additional specimen examined (paratype): –– BRAZIL. Minas Gerais: Presidente Kubitschek, along the road from Serro to Datas, 1 km before the bridge over the Ribeirão Andrequicé, 1,200 m elev., epilithic, 30 January 1995, W . Till et al. 11082, cult. in hort. bot. Vindob. B32 /95 (WU).</p><p>Distinctive characters:— Tillandsia montezumensis belongs to the widespread T. tenuifolia complex due to its habit, leaf, inflorescence, and flower conformation. In relation to T. tenuifolia, this new species can be distinguished by its leaf blades completely covered by a dense coat of white trichomes obscuring the color of the blades (vs. densely and minutely lepidote with trichomes not obscuring the color of the blades) and longer (14–17 cm vs. 5–10 cm long), floral bracts equaling to slightly shorter than the sepals (vs. exceeding the sepals), and by the longer sepals (ca. 16 mm vs. ca. 10 mm long).</p><p>Tillandsia iassuensis L. Hromadnik &amp; H. Hromadnik (2021: 76), from Bahia state, is another morphological relative. However, T. montezumensis differs from it by the stemless to shortly caulescent habit (vs. long caulescent), shorter when in bloom (20–25 cm vs. to 40 cm tall), inflorescence with fewer flowers (3–5 vs. 6–12 in number), floral bracts longer (15–17 mm vs. 10–14 mm long), sepals longer (ca. 16 mm vs. 6–12 mm long), the posterior ones connate at the base to 1/3 of its length (vs. connate for more than 2/3 of its length).</p><p>Tillandsia milagrensis Leme(1993:243) can be also considered morphologically closely related to T.montezumensis, but this new species differs by the stemless to shortly caulescent habit (vs. distinctly caulescent), shorter when in bloom (20–25 cm vs. ca. 50 cm tall), inflorescence shorter (30–35 mm vs. ca. 50 mm long), with fewer flowers (3–5 vs. ca. 10 in number), floral bracts equaling to slightly shorter than the sepals (vs. exceeding the sepals), and the posterior sepals connate at the base for 3.5–5 mm (vs. 9–11 mm).</p><p>Both Tillandsia iassuensis and T. milagrensis grows in the same region of the state of Bahia, and their populations are separated from each other by about 40 km in straight line, in the Caatinga biome. They form large population on vertical granitic outcrops. In contrast, T. montezumensis is know from Minas Gerais state, near the border with Bahia, about 380 km distant in straight line, and was observed in small groups of plants on quartizitic outcrops in the domain of the Campos Rupestres.</p></div>	https://treatment.plazi.org/id/039E87CD8606D04995D161ABFE42FB2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8604D04F95D167C7FA87FCCB.text	039E87CD8604D04F95D167C7FA87FCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tillandsia tephrosa Leme & W. Till 2025	<div><p>Tillandsia tephrosa Leme &amp; W. Till, sp. nov. (Fig. 19 A–E)</p><p>Diagnosis:––This new species is closely related to Tillandsia hofackeri Ehlers (2013: 22), but can be distinguished from it by the leaf blades longer (4–6 cm vs. 2–3 cm long), inflorescence slightly exceeding the leaves (vs. distinctly exceeding the leaves), flowers shorter (ca. 20 mm vs. ca. 30 mm long), and petals shorter (ca. 19 mm vs. 24–33 mm long).</p><p>Type: –– BRAZIL. Minas Gerais: Lagoa Santa, Parque Estadual do Sumidouro, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.95861&amp;materialsCitation.latitude=-19.56111" title="Search Plazi for locations around (long -43.95861/lat -19.56111)">Gruta da Lapinha</a>, limestone outcrop above the cave, 722 m elev., 19º33’40” S, 43º57’31” W, 25 May 2010, E . Leme 8305 &amp; R. Oliveira (holotype RB!) .</p><p>Description:–– Plants epilithic, flowering 11–15 cm tall, distinctly caulescent, propagating by elongate shoots developed near the base of the peduncle, forming dense groups. Leaves 25–30 in number, densely and equally arranged along the stem, without impounding capacity; sheath inconspicuous, subtrapeziform, corrugate abaxially, ca. 0.5 × 0.6 cm, completely covered on both sides by a dense coat of cinereous, appressed trichomes; blade narrowly triangular-attenuate, 4–6 × 0.3–0.4 cm (near the base), spreading, not secund, not narrowed at the base, subrigid toward the base, pliable toward the apex, completely covered on both sides by a dense layer of cinereous appressed trichomes obscuring the blade color, apex filiform-caudate. Peduncle slender, ca. 3 cm long, 1.5–2 mm in diameter, erect, glabrescent, green; peduncle bracts the basal ones foliaceous, the upper ones with a narrowly ovate-lanceolate base 12–15 × 4 mm, partially enfolding the peduncle, reddish-rose, cinereous lepidote, and with a filiform blade 7–15 mm long, erect, densely cinereous lepidote, distinctly exceeding the internodes. Inflorescence (fertile part) simple, 20–30 mm long, erect, densely 3–4-flowered, slightly exceeding the leaves; rachis slender, slightly flexuous, greenish, glabrous, internodes 3–4 × 1–1.5 mm long; floral bracts distinctly exceeding the sepals, ecarinate, narrowly elliptic-lanceolate, reddish-rose, 12–17 × 6 mm, membranaceous, thinly nerved, glabrous or the apex sometimes densely white lepidote, apex shortly caudate to apiculate (basal ones) or acute (apical ones). Flowers diurnal, odorless, polystichously and densely arranged, erect, ca. 20 mm long with extended petals, pedicels inconspicuous, ca. 1 mm long, green, glabrous; sepals narrowly lanceolate, symmetrical, erect, apex acute, 10–10.5 × 3–4 mm, glabrous, rose, thin in texture, the abaxial one free, ecarinate, the adaxial ones highly connate for ca. 7.5 mm, carinate toward the apex; petals narrowly spathulate with a narrow proximal portion in contrast to the broader distal portion, apex rounded and emarginate, 19 × 3.5–4 mm, violet-blue, free, its basal portion erect and forming a subtubular corolla except for the subspreading distal portion, exappendiculate; stamens distinctly shorter than the petals; filaments complanate, strongly plicate in the middle, white, free, ca. 0.8 mm wide; anthers narrowly oblong, at the base distinctly bilobed, apex subacute, ca. 2 mm long, dorsifixed near the base, style distinctly exceeding the anthers, ca. 12.5 mm long; ovary totally superior; stigma conduplicate-patent, lobes ca. 0.5 mm long, sparsely papillose; ovules apiculate. Capsules unknown.</p><p>Distribution and habitat:–– Tillandsia tephrosa is known from the type locality only, the state park of Sumidouro, in the county of Lagoa Santa, Minas Gerais state, southeastern Brazil. It is an epilithic species, forming small clumps on the vertical parts of the limestone outcrops above the Lapinha Cave, which is a touristic attraction in the region. Besides situated inside a state park, the area where T. tephrosa grows is located outside the permitted access to tourists, which provides additional protection for the species.</p><p>Etymology:––The name of this new species is a reference to the somewhat ash color of its leaves, based on the ancient Greek word “tephra” meaning “ashes”, which currently also means “ash and debris” ejected by a volcanic eruption.</p><p>Distinctive characters:— Tillandsia tephrosa is a member of Tillandsia subg. Anoplophytum (Beer 1854: 346) Baker (1887: 212), belonging to the species complex formed by T. tenuifolia . It is morphologically closely related to T. hofackeri because of its leaf blades completely covered on both sides by a dense layer of cinereous appressed trichomes obscuring the blade color and the inflorescence conformation. However, it can be distinguished from it by the larger size when in bloom (11–15 cm vs. ca. 9 cm tall), leaf blades longer (4–6 cm vs. 2–3 cm long), inflorescence slightly exceeding the leaves (vs. distinctly exceeding the leaves), floral bracts narrower (ca. 6 mm vs. 8–9 mm), flowers shorter (ca. 20 vs. ca. 30 mm long), and petals shorter (ca. 19 mm vs. 24–33 mm long) and lilac (vs. light-blue).</p></div>	https://treatment.plazi.org/id/039E87CD8604D04F95D167C7FA87FCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8602D04D95D160E7FC55FC27.text	039E87CD8602D04D95D160E7FC55FC27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vriesea ilhagrandensis Leme & W. Till 2025	<div><p>Vriesea ilhagrandensis Leme &amp; W. Till, sp. nov. (Fig. 20 A–H)</p><p>Diagnosis:—This new species is morphologically closely related to Vriesea rubyae E. Pereira (1971: 115), differing from it by the leaf blades narrowly subriangular and acuminate (vs. narrowly lingulate, rounded to slightly emarginate with a slender apiculus), narrower inflorescence (ca. 2 cm vs. 3–3.5 cm), floral bracts elliptic (vs. suborbicular), and sepals elliptic-lanceolate, acuminate (vs. oblong-ovate, acute).</p><p>Type:— BRAZIL. Rio de Janeiro: Angra dos Reis, Ilha Grande, trilha para o <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.192238&amp;materialsCitation.latitude=-23.155787" title="Search Plazi for locations around (long -44.192238/lat -23.155787)">Pico do Papagaio</a>, ca. 828 m elev., 23º 09’ 20.83” S, 44º 11’ 32.05” W, 2 March 2002, E . Leme 5427, B. R . Silva, F . Tatagiba &amp; L . Souza (holotype RB!) .</p><p>Description:— Plants epiphytic, flowering ca. 35 cm tall, propagating by spreading stolons ca. 6 cm long, ca. 0.7 cm in diameter, covered by densely arranged, stramineous cataphylls. Leaves ca. 15, suberect, forming at the base a narrow funnelform rosette; sheath broadly elliptic, 5–5.5 × 3.5–4 cm, subdensely brown lepidote, dark winish-purple, finely nerved; blade narrowly subriangular, 10–11 × 1.5–2 cm, not narrowed toward the base, dark winish-purple near the base, green toward the apex, sparsely and inconspicuously white lepidote mainly abaxially to glabrescent, apex acuminate. Peduncle erect, ca. 18 cm long, ca. 0.4 cm in diameter, rigid, greenish, glabrous; peduncle bracts lanceolate, acute to acuminate and shortly caudate, exceeding the internodes, erect, imbricate, completely covering the peduncle, red except for the greenish apex. Inflorescence (fertile part) simple, densely flowered, decurved but not pendulous, ca. 15 × 2 cm, sublinear in outline, distinctly complanate, exceeding the leaf blades, rachis completely covered by the floral bracts, subangulose, glabrous, ca. 3 mm in diameter, internodes 8–10 mm long; floral bracts slightly exceeded by the sepals, base truncate, slightly incurved toward the apex, erect with the flowers, densely imbricate, laterally compressed, obtusely if at all carinate, elliptic, red except for the pale yellowish apex, ca. 27 × 18 mm, thin in texture, glabrous, acute. Flowers ca. 14 in number, anthesis diurnal, odorless, distichous, not secund, ca. 45 mm long (not including the stamens), pedicels ca. 8 × 4 mm (at the distal end), green, glabrous; sepals elliptic-lanceolate, acuminate, ca. 25 × 8 mm, glabrous, reddish near the base, pale yellowish toward the apex, free, ecarinate, cymbiform, thin in texture; petals sublinear-spathulate, ca. 38 × 5.5–6 mm, connate at the base for ca. 4 mm, apex narrowly rounded and inconspicuously emarginate, forming a tubular corolla except for the suberect distal portion, yellow, bearing at the base 2 narrowly obovate, obtuse to subacute, entire, ca. 11 × 1.5–2 mm appendages adnate to the petals for ca. 8 mm. Stamens distinctly exceeding the petals at anthesis; anthers linear, ca. 5 mm long, base bilobed, apex obtuse, dorsifixed ca. 1 mm above the base; stigma convolute-blade II (vrieseoid), densely papillose, ca. 1.5 mm in diameter, exceeding the anthers; ovules caudate. Capsules unknown.</p><p>Distribution and habitat:–– Vriesea ilhagrandensis is only known from the type locality, where it grows as an epiphyte on the higher tree branches, in an a 830 m elevated area covered by well-preserved Atlantic Forest vegetation (fig. 20 A). It was found in the trail to Pico do Papagaio, the second highest peak on the Ilha Grande, a large oceanic island situated at the so called “Green Coast”, in the county of Angra dos Reis, Rio the Janeiro state, southeastern Brazil. Ilha Grande is well-known for its wild beaches of clear water and well-preserved forest, which attracts a lot of tourists every year.</p><p>Besides its close morphological parent, V. rubyae, other epiphytic Vriesea species were documented in the same area, such as V. jonghei (K. Koch 1868: 91) E. Morren (1878: 257), V. unilateralis (Baker 1888: 105) Mez (1894: 545), and V. vagans (L.B. Smith 1943: 121) L.B. Smith (1966: 118), as well as a rupicolous V. aff. regnelli treated by Moura (2011) as a new species, but not yet officially published.</p><p>Etymology:—The name chosen for this new species is a direct reference to Ilha Grande, the place where it was discovered.</p><p>Distinctive characters:— Vriesea ilhagrandensis is closely related to V. rubyae, another inhabitant of the Atlantic Forest of Rio de Janeiro state, which also lives in the same area where this new species was found. Both species are small-sized and stoloniferous, despite their rosette conformation and leaf blades color clearly distinguish them from each other. They also share the complanate inflorescence bearing floral bracts and flowers of similar colors. In addition, V. ilhagrandensis clearly differs from V. rubyae by its leaf blades narrowly subriangular and acuminate (vs. narrowly lingulate, rounded to slightly emarginate with a slender apiculus), narrower (1.5–2 cm vs. 3–4 cm wide), green toward the apex (vs. green-glaucous toward the apex), narrower inflorescence (ca. 2 cm vs. 3–3.5 cm), sublinear in outline (vs. oblong-lanceolate to narrowly elliptic-ovate), floral bracts elliptic (vs. suborbicular), narrower (ca. 18 mm vs. 30 mm wide), and sepals elliptic-lanceolate, acuminate (vs. oblong-ovate, acute).</p></div>	https://treatment.plazi.org/id/039E87CD8602D04D95D160E7FC55FC27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD8600D05395D160C3FE8BFA2F.text	039E87CD8600D05395D160C3FE8BFA2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vriesea punctatissima Leme & E. H. Souza 2025	<div><p>Vriesea punctatissima Leme &amp; E.H. Souza, sp. nov. (Fig. 21 A–J)</p><p>Diagnosis: ___ This new species is closely related to Vriesea pulchra Leme &amp; L. Kollmann (2011: 29), differing from it by the leaf blades abaxially green and ornamented with dense and minute dark red spots (vs. greenish to dark purple, without red spots), adaxially without noticeable longitudinal lines and cross-veins (vs. with dark green or dark purple longitudinal lines and irregularly wavy, distinct, dark green cross-veins), and by sepals greenish with large dark purplish-wine spots toward the apex and margins (vs. green toward the base and yellowish toward the apex, except for the dark purple margins and apex, without conspicuous spots).</p><p>Type: –– BRAZIL. Bahia Camacã, Fazenda Serra Bonita, L . F . N. Carvalho 2386, fl. cult. July 2009, E . Leme 6144 (holotype RB!) .</p><p>Description:— Plants epiphytic, flowering 38–50 cm tall, propagating by short basal shoots. Leaves 15–18 in number, suberect-arcuate, forming a funnelform rosette, chartaceous; sheath ovate, ca. 10 × 8 cm, densely brown-lepidote on both sides, abaxially and toward the base dark castaneous, greenish and ornamented with dense and minute dark red spots, adaxially pale castaneous toward the base; blade linear, not narrowed at the base, 20–23 × 4.3–6 cm, inconspicuously and sparsely white lepidote and inconspicuously covered by a thin layer of epicuticular white wax on both sides, nerved, abaxially green and ornamented with dense and minute dark red spots, adaxially green or sometimes with an apical dark red, small spot, apex rounded and shortly apiculate. Peduncle erect, 25–35 cm long, ca. 0.7 cm in diameter, greenish to purple-castaneous, glabrous; peduncle bracts erect, broadly ovate to suborbicular, broadly acute to obtuse and minutely apiculate, imbricate, equalling to slightly exceeding the internodes, green with dense and large purple-wine spots, glabrous or nearly so. Inflorescence (fertile part) simple, erect, broadly ellipsoidic or nearly so in outline and inconspicuously spirally-twisted at anthesis, 8–12 × 6–8 cm (excluding the petals), laxly flowered toward the base at anthesis, slightly covered by a weak layer of glutinous substance, rachis 4–7 mm in diameter, smooth, flexuous, slightly angled, green to greenish-purple or greenish-castaneous, glabrous; floral bracts not imbricate, equaling ca. 1/3 of the sepals length, bearing decurrent auricles at the base, strongly convex, ecarinate, suborbicular, margins and apex greenish, centrally castaneous, ornamented with large, dark purple-wine spots, 20–28 × 20–27 mm, thinly coriaceous, lustrous, glabrous abaxialy, inconspicuously white-lepidote adaxially, holding inside a translucent mucilagenous substance, subacute and minutely apiculate (basal) to obtuse (apical). Flowers 6–12 in number, 43–55 mm long, nocturnal, distichous, producing a strong garlic smell, laxly arranged at anthesis, subspreading, divergent to slightly secund, pedicels stout, 9–15 mm long, 8–9 mm in diameter at the apex and ca. 7 mm in diameter at the base, pale yellowish-castaneous, glabrous; sepals elliptic, obtuse-emarginate, 21–25 × 13–14 mm, greenish with large dark purplish-wine spots toward the apex and the margins, ecarinate, glabrous abaxially and inconspicuously white-lepidote adaxially, coriaceous near the base and membranaceous toward the margins and the apex, convex; petals obovate to subelliptic, 35–38 × 18–22 mm, apex distinctly emarginate, suberect to subspreading toward the apex at anthesis and forming a campanulate corolla ca. 25 mm in diameter, connate at the base for ca. 3 mm, pale yellow with irregular wine-colored nerves near the apex, bearing at the base 2 appendages; appendages 11× 2.5–4 mm, basally adnate for 3– 4 mm, blades ovate-lanceolate, long acuminate-caudate to long-bifid. Stamens shorter than the petals, positioned side by side in the lower part of the corolla at anthesis; filaments adnate to the petals for ca. 3 mm, slightly dilated toward the apex; anthers ca. 9 mm long, dorsifixed near the base, base bilobed, apex obtuse; pollen ellipsoid, sulcate, exine broadly rediculate; style elongate, yellowish, positioned above the anthers and about equaling them; stigma convolutebladed, ca. 2 mm in diameter, blades papillose, yellow; ovary pale yellow; ovules caudate. Capsules unknown.</p><p>Distribution and habitat:–– Vriesea punctatissima is known from the neighboring municipalities of Arataca and Camacã, in the southern region of Bahia state, located in the northeast of Brazil. It inhabits submontane to montane Atlantic Forest, at 350 to 1,000 m elevation (fig. 19 A), as an epiphyte of the canopy (fig. 21 A–B). This new species was documented inside two conservation units: a private one, RPPN Serra Bonita, in Camacã, and in the Parque Nacional da Serra das Lontras, situated in Arataca, a public conservation unit.</p><p>Etymology:––The name of this new species is based on the Latin word “ punctatus ”, meaning “with spots”, with the ending “ issumus ” to express a superlative condition, as a direct reference to its prominently spotted leaf blades.</p><p>Additional specimen examined (paratypes): –– BRAZIL. Bahia: Camacã, RPPN Serra Bonita, Núcleo Serra Bonita, Trilha das Bromélias, 14 April 2009, R . Moura 793 (R!); ibidem, 9.7 km de Camacã na estrada para Jacareci, 6 km SW na estrada para a RPPN Serra Bonita e torre da Embratel, 900–1,000 m elev., 15º23’30” S, 39º33’55” W, 9 December 2006, R. A . X. Borges et al. 369 (RB!, CEPEC); Arataca, Parque Nacional da Serra das <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.38111&amp;materialsCitation.latitude=-15.201388" title="Search Plazi for locations around (long -39.38111/lat -15.201388)">Lontras</a>, 15º12’05” S, 39º22’52” W, July 2011, P . Leitman 253 (RB!); ibidem, trilha da Serra das <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.435&amp;materialsCitation.latitude=-15.189167" title="Search Plazi for locations around (long -39.435/lat -15.189167)">Lontras</a>, 15º12’21” – 15º11’21” S, 39º26’07” – 39º26’06” W, 5 May 2011, P . Leitman et al. 218 (RB!); ibidem, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.351387&amp;materialsCitation.latitude=-15.181389" title="Search Plazi for locations around (long -39.351387/lat -15.181389)">Trilha do Talhão</a>, 348–583 m elev., 15º10’53” S, 39º20’53” – 39º21’05” W, 21 November 2011, P . Leitman et al. 442 (RB!); ibidem, Pratas, entrada na BR 101 em frente a entrada para Jussari, Fazenda da Dra. Karitas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.393333&amp;materialsCitation.latitude=-15.186111" title="Search Plazi for locations around (long -39.393333/lat -15.186111)">Serra do Mangue</a> (complexo Serra das Lontras), 361–516 m elev., 15º12’10” – 15º11’10” S, 39º24’31” – 39º23’36” W, 25 November 2011, P . Leitman et al. 508 (RB!, CEPEC) .</p><p>Distinctive characters:— Vriesea punctatissima (fig. 20 C–J) is morphologically closely related to V. pulchra (fig. 20 K–M) because of its similar stature, inflorescence structure, and night flowers. However, it can be visually distinguished from the close relative by its more compact leaf rosette due to the broader leaf blades (4.3–6 cm vs. 3.5–3.7 cm wide), which are abaxially green and ornamented with dense and minute dark red spots (vs. greenish to dark purple, without red spots), which is its most striking characteristic. Additionally, the adaxial surface of leaf blades of V. punctatissima lacks noticeable longitudinal lines and cross-veins, contrasting with the dark green or dark purple longitudinal lines and irregularly wavy, distinct, dark green cross-veins of the adaxial surface of the leaf blades of V. pulchra (fig. 21 K), which also presents sepals without conspicuous spots (fig. 21 M), while the new species has sepals with large dark purplish-wine spots (fig. 21 E, G).</p><p>While Vriesea punctatissima is only known from the Bahian counties of Arataca and Camacã, its close morphological relative, V. pulchra, was observed at its type locality only, over a distance of 500 km in a straight line, in the municipality of Santa Teresa, Espírito Santo state, in similar Atlantic forest sector, altitude, and distance from the Atlantic Ocean.</p></div>	https://treatment.plazi.org/id/039E87CD8600D05395D160C3FE8BFA2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
039E87CD861ED05195D166DBFCC2F90F.text	039E87CD861ED05195D166DBFCC2F90F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wittrockia organensis Leme & Barfuss 2025	<div><p>Wittrockia organensis Leme &amp; Barfuss, sp. nov. (Fig. 22 A–G)</p><p>Diagnosis:— This new species can be distinguished from its likely closest relative, Wittrockia cyathiformis (Vellozo 1831: pl. 144) Leme (1997: 67), by the inflorescence narrower (5–8 cm vs. 8–15 cm in diameter in the distal portion), flowers shorter (35–36 mm vs. 55– 60 mm long), and petals smaller (21–22 × 5 mm vs. ca. 43 × 11 mm), green toward the apex (vs. golden yellow toward the apex).</p><p>Type:— Brazil. Rio de Janeiro: Teresópolis, Serra dos Órgãos, 14 March 2015, E. Leme 8988, M. Barfuss &amp; M. de La Harpe (holotype RB!) .</p><p>Description:— Plants propagating by basal shoots. Leaves 16–20 in number, thinly subcoriaceous in texture, forming a crateriform rosette; sheath elliptic-ovate, 16 × 9–10.5 cm, pale green, densely and coarsely brown lepidote toward the base adaxially, densely and minutely brown lepidote abaxially, strongly nerved; blade lingulate, narrowed toward the base, 20–40 × 5–6.5 cm, glabrescent, green, distinctly nerved toward the base and along the margins, apex acute and apiculate, margins densely spinose, spines retrorse (basal ones) to spreading (distal ones), 1–2 mm long, 2–4 mm, apart; peduncle suberect, 37–38 × 0.7–1 cm, dark red, glabrous, sulcate; peduncle bracts narrowly ovate-lanceolate, acute and apiculate, 6–9 × 2.5–3 cm, glabrescent, dark red, densely spinulose, spines 0.5–1 mm long, one near the middle of the peduncle or at 2/3 of its length above the base, the remaining concentrated around the inflorescence and involucrate, suberect, equalling to exceeding the petals. Inflorescence (fertile part) shortly corymbose, obconic, apex stellate, twice branched, distinctly elevated above the rosette, 6–7 cm long, 5–8 cm in diameter at the apex; primary bracts resembling the involucral bracts, but gradually smaller, the outer ones narrowly ovate-lanceolate, apex acute and apiculate, suberect, 5–6 × 2–2.5 cm, glabrescent, dark red, densely spinose, spines irregularly curved, 0.5–1.5 mm long; primary fascicles 5–6 in number, 40–45 × 20–28 mm, flabellate, subcomplanate, 5–8-flowered, distinctly pedunculate, peduncle 5–7 × 11–14 mm; floral bracts equalling about 3/5 of the sepals length, carinate, narrowly triangular-lanceolate, hyaline at the base and red at the apex, 26–28 × 10–11 mm, membranaceous, finely nervate, glabrous, apex long acuminate, margins entire to remotely spinulose. Flowers 35–36 mm long, sessile, diurnal, odorless; sepals narrowly lanceolate, subsymmetrical, apex long attenuate-caudate, 24–25 × 5–5.5 mm, connate at the base for 1–1.5 mm, glabrous, green, ecarinate to obtusely if at all carinate, thick at the base and thin in texture toward the base and margins; petals subspathulate, 21–22 × 5 mm, free, membranaceous, apex acute, green toward the apex, erect except for the spreading apex at anthesis, without callosities, bearing 2 suboblong appendages at the base, ca. 4 × 1.5 mm, with irregular and long laciniate apex; filaments ca. 17 mm long, free; anthers sublinear, ca. 5 mm long, base bilobed, apex acute, fixed at ¼ of its length above the base; stigma conduplicate-spiral, obovoid, ca. 3 × 2 mm, lobes with minutely crenulate margins, without papillae, greenish-yellow; ovary subclavate to rectangular, trigonous, ca. 10 × 7 mm, white, glabrous; epigynous tube 0.5–1 mm long; placentation from median to apical; ovules many, subcylindrical, obtuse. Fruits unknown.</p><p>Distribution and habitat:––This new species was found growing terrestrially in nebular Atlantic Forest above 1,500 m elevation in the county of Teresópolis, in the Orgão mountain, Rio de Janeiro state, southeastern Brazil (fig. 22 A). It forms large and dense groups of plants on the forest floor, being associated with a Vriesea sp. (fig. 22 B).</p><p>According to the additional specimens examined below, Wittrockia organensis is also encountered inside the Serra dos Órgãos National Park, in the neighbourhood of Pedra do Sino and Agulha do Diabo, about 2,000 m elevation, which represents an effective preservation of its local populations.</p><p>Etymology:––The name of this new species is a reference to the Orgãos montains, which is a portion of the Serra do Mar range in Rio de Janeiro state.</p><p>Additional specimens examined (paratypes):–– BRAZIL. Rio de Janeiro: Cachoeira do Rancho Frio, 1,400 m elev., 23 August 1940, A.C. Brade 16627 (RB!); Teresópolis, Serra dos Órgãos, 14 March 2015, E. Leme et al. 8989 (RB!) ; Parque Nacional da Serra dos Órgãos, arredores da Pedra do Sino , 2,100 m elev., 26 September 2007, G. Heiden 892 (RB!) ; ibidem, trilha para o alojamento 4 e a Pedra do Sino, 1,190-2,130 m elev., 22º 25-32’ S, 42º 59’– 43º 07’W, 12–13 April 2011, J.A. Lombardi 8231 &amp; Taxonomia de Campo 2011 (caderno de campo 154) (RB!) ; ibidem, Agulha do Diabo, 21 February 2009, C.R. França 41 &amp; R. Moura (RB!) .</p><p>Distinctive characters:— Wittrockia organensis has been misidentified in Brazilian herbaria as W. cyathiformis, which is its close morphological relative [see material examined by Leme (1997) for Rio de Janeiro state concerning this species], due to its similar stature and leaf conformation, as well as the similar inflorescence shape and structure. Smith (1955) highlighted that in Nidularium classification based only on herbarium sheets is difficult and ambiguous, and this situation would only improve if much live material where examined. This is true for all members of the “Nidularioid complex”, including Wittrockia . It was for no other reason that the discovery of this new species was only possible due to the analysis of living species in bloom.</p><p>This new species can be distinguished from Wittrockia cyathiformis by the basal portion of the leaf blades with shorter marginal spines (1–2 mm vs. 2–5 mm long), primary bracts smaller (5–6 × 2–2.5 cm vs. 7–12 × 3–5 cm), suberect at anthesis (vs. spreading-recurved to strongly reflexed), with shorter spines along its basal portion (0.5–1.5 mm vs. 2–3 mm), inflorescence narrower (5–8 cm vs. 8–15 cm in diameter in the distal portion), flowers shorter (35–36 mm vs. 55–60 mm long), sepals shorter (24–25 mm vs. ca. 30 mm), connate at the base for 1–1.5 mm (vs. free), and petals smaller (21–22 × 5 mm vs. ca. 43 × 11 mm), green toward the apex (vs. golden yellow toward the apex), with basal appendages long laciniate (vs. irregularly denticulate).</p></div>	https://treatment.plazi.org/id/039E87CD861ED05195D166DBFCC2F90F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leme, Elton M. C.;Souza, Everton Hilo De;Till, Walter;Barfuss, Michael H. J.;Filho, José Alves Siqueira;Kollmann, Ludovic J. C.;Couto, Dayvid R.;Fraga, Claudio Nicoletti De;Fontana, André P.;Farias-Castro, Antônio S.;Fernandes, João B.;Silva, Da	Leme, Elton M. C., Souza, Everton Hilo De, Till, Walter, Barfuss, Michael H. J., Filho, José Alves Siqueira, Kollmann, Ludovic J. C., Couto, Dayvid R., Fraga, Claudio Nicoletti De, Fontana, André P., Farias-Castro, Antônio S., Fernandes, João B., Silva, Da (2025): Twenty Miscellaneous New Species and One New Nothogenus and Nothospecies in Brazilian Bromeliaceae. Phytotaxa 692 (1): 1-60, DOI: 10.11646/phytotaxa.692.1.1, URL: https://doi.org/10.11646/phytotaxa.692.1.1
