taxonID	type	description	language	source
039B87B7B22B3A69FFC4FA34D88B4CBD.taxon	materials_examined	Type species. Notoatherix antiqua Oberprieler & Yeates, sp. nov.	en	Oberprieler, Stefanie K., Yeates, David K. (2014): Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae). Zootaxa 3866 (1): 138-144, DOI: 10.11646/zootaxa.3866.1.8
039B87B7B22B3A69FFC4FA34D88B4CBD.taxon	diagnosis	Diagnosis. Only a single wing blade is preserved, with the anterior base obscured. R 1 and R 2 + 3 relatively straight, R 2 + 3 meeting R 1 at costa. R 4 + 5 proximally straight, then weakly curved upwards posteriorly, meeting costa of wing just before apex. Crossvein r-m almost in middle of cell d, but tending slightly towards its basal end. Vein M 1 sinuous at base, remainder almost straight, meeting wing margin just behind wing apex. M 2 relatively straight, meeting wing margin. Crossvein m-m closing cell d at level of origin of M 1. Species included. Type species only.	en	Oberprieler, Stefanie K., Yeates, David K. (2014): Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae). Zootaxa 3866 (1): 138-144, DOI: 10.11646/zootaxa.3866.1.8
039B87B7B22B3A69FFC4FA34D88B4CBD.taxon	discussion	Remarks. Venational features characteristic of the Athericidae were defined by Stuckenberg (1973). The diagnostic feature of the family, the close terminal convergence of veins R 1 and R 2 + 3, occurs in Notoatherix. In addition, crossvein r-m meets the discal cell further from its basal end, another feature to distinguish rhagionids and athericids used by Stuckenberg (1973). Two other features of many athericids, veins R 4 and R 5 diverging at a far smaller angle than in rhagionids and tabanids, and R 4 ending before the tip of the wing (Stuckenberg 1973), do not occur in Notoatherix because R 4 and R 5 are fused throughout their length. Mostovski et al. (2003) reported that the conformation of R 4 and R 5 is more variable in the fossil record of the three families than in extant forms. Notoatherix dates to an age when these three families are believed to have diverged (Wiegmann et al. 2011) and could possess a venation different from that of the current Athericidae crown group. FIGURE 2. Line drawing of wing of venation of Notoatherix antiqua gen. et sp. nov., holotype; composite, from part and counterpart. Abbreviations: C = costa; Sc = subcosta; R = radius; R 2 + 3 = anterior branch of the radial sector; R 4 + 5 = posterior branch of the radial sector; M 1, M 2 = anterior (sectoral) branches of media; M 3 = posterior (sectoral) branch of media; CuA 1, CuA 2 = anterior branches of cubitus; A = anal vein; r-m = radial to medial cross-vein; m-m = medial to medial cross-vein; m-cu = medial to cubital cross-vein; d = discal cell; br = first basal cell; bm = second basal cell; m 1 - 3 = first to fourth medial cells; cua 1 = anterior cubital cell; cup = posterior cubital cell.	en	Oberprieler, Stefanie K., Yeates, David K. (2014): Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae). Zootaxa 3866 (1): 138-144, DOI: 10.11646/zootaxa.3866.1.8
039B87B7B2293A69FFC4FBD8DB0349E2.taxon	materials_examined	Type material. Holotype: Talbragar Fish Bed (Late Jurassic: Kimmeridgian (151 ± 4 Ma), Gulgong, New South Wales, Australia, collector Jack Hinde. Part and counterpart impressions of wing; incomplete preservation with portion of anterior base obscured.	en	Oberprieler, Stefanie K., Yeates, David K. (2014): Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae). Zootaxa 3866 (1): 138-144, DOI: 10.11646/zootaxa.3866.1.8
039B87B7B2293A69FFC4FBD8DB0349E2.taxon	etymology	Etymology. The species is named for its great age, being the oldest known adult representative of its family. Occurrence. Only known from the Talbragar Fish Bed.	en	Oberprieler, Stefanie K., Yeates, David K. (2014): Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae). Zootaxa 3866 (1): 138-144, DOI: 10.11646/zootaxa.3866.1.8
039B87B7B2293A69FFC4FBD8DB0349E2.taxon	description	Description. Wing relatively large and broad, 12 mm long, 4.8 mm at widest point. R 1 and R 2 + 3 relatively straight, R 2 + 3 meeting R 1 at costa. R 4 + 5 proximally straight, distally weakly curved anteriad, meeting costa before apex. R-m crossvein almost at middle of cell d but tending slightly towards its basal end. Vein M 1 sinuous at base, remainder almost straight, meeting wing margin just behind apex. M 2 gently evenly curved, meeting wing margin. M-m crossvein closing cell d at level of origin of M 1. M 3 and CuA 1 gently evenly curved, ending before wing margin. Cell br longer than bm, base of M 3 and CuA 1 arising close together at apex of cell bm. CuA 2 slightly curved posteriorly, CuP visible as a concave vein just behind CuA 2. A 1 well developed, with similar curvature to CuA 2, ending before wing margin. Stigma faint, with indistinct margins, more prominent behind R 1 but extending in front of this vein.	en	Oberprieler, Stefanie K., Yeates, David K. (2014): Notoatherix antiqua gen. et sp. nov., first fossil water snipe fly from the Late Jurassic of Australia (Diptera: Athericidae). Zootaxa 3866 (1): 138-144, DOI: 10.11646/zootaxa.3866.1.8
