taxonID	type	description	language	source
03A77E503542FFA0A6A0FE75FBEBF75D.taxon	description	Most of the species of the tribe Anacolini are small and the genus Hystatoderes stands out for the large size of itsspecies. They exhibita stocky habitus rather like other Prionini, a transverse pronotum, swollen on the discal area, more or less wrinkled with lateral spines and a rather clear pilosity. The elytra present punctation and wrinkles on the basal and periscutellar zone, the costae being not very apparent. The antennae are 11 - articulated in both sexes, relatively short, not exceeding 2 / 3 or 3 / 4 of the elytral length with antennomeres 4 to 10 of about the same length. Meso- and metatibiae are strongly recessed on the inner side. The lower eye-lobe is not much wider than the upper one and does not protrude below the insertion of the antennae. Sexual dimorphism - All the species of this genus are characterized by a very important sexual dimorphism to the point that it is difficult to make a species link between the two sexes. - The male shows much smaller body size and elytra elongated, narrowed at apex and with more angular humeral corners, whereas the female shows an oblong elytral shape, which is slightly enlarged between the middle and two thirds of the elytra, and rounded humeral angles. - The elytra of the male are covered with sparse, fairly fine hairs, with fairly sharp ridges and a satiny appearance, while those of female are hairless, with barely visible ridges and a shiny appearance. In addition to the elytra, golden hairs are present on forehead, vertex, pronotum and scutellum of males. - The male has a narrower head with a closer upper interocular space than that of the female and longer antennae with strongly punctate antennomeres and sharp or angular apical angles, whereas the antennomeres are smooth with rounded angles at the apex in the female. - The pronotum of the male is trapezoidal, entirely punctate, with spines towards the base of the lateral margins and effaced apical angles. The pronotum of the female is sub-quadrangular, with a smooth swollen disc, strongly crenellated or spiny sides and very angular apical angles. - The femora and tibiae are spotted and scattered with hairs in the male, whereas they are almost smooth and hairless in the female. Finally, the tarsi of the male are slightly longer and expanded compared to the female. FAUNITAXYS - ISSN (Print): 2269 - 6016 - ISSN (Online): 2970 - 4960	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E503543FFA1A683FF45FA55FA4D.taxon	description	(Fig. 1, 2 A-B, 6 A & 6 D)	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E503543FFA1A683FF45FA55FA4D.taxon	materials_examined	Holotype, ♀ (Fig. 2 B): Vietnam, Da Nang City, Ba Na Mt., 1450 m, V. 2015, local collector leg. (will be deposited in NSMT). Allotype, ♂ (Fig. 2 A): same locality as holotype, VI. 2014, local collector leg. (will be deposited in RBINS, I. G.: 34.629). Paratypes	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E503543FFA1A683FF45FA55FA4D.taxon	description	1 ♂, same locality and date as holotype (ZKC); 1 ♂, same locality as holotype but IV. 2015 (ZKC); 1 ♂, Vietnam, Quang Nam province, Tay Giang, Axan Mt., 1300 m, V. 2017, local collector leg. (CRC); 1 ♂, same locality but IV. 2018, local collector leg. (ZKC); 1 ♂, same locality and date, local collector leg. (SIC); 1 ♂, same locality but III. 2020, local collector leg. (AKC); 2 ♂, same locality but IV. 2020, local collector leg. (AKC); 2 ♂, same locality but IV. 2021, local collector leg. (AKC); 1 ♂, same locality but III. 2022, local collector leg. (AKC); 1 ♂, same locality but III. 2022, local collector leg. (NDC); 1 ♂, same locality but III. 2022, local collector leg. (ZCDTU); 2 ♂, same locality, local collector leg. (ADC); 1 ♂, same locality, local collector leg. (BSC); 1 ♀, Vietnam, Gia Lai province, Kon Chu Rang Nature Reserve, V. 2021, local collector leg. (AKC); 1 ♀, Vietnam, Kon Tum province, Ngoc Linh Nature Reserve, 1900 m, III. 2022, local collector leg. (AKC).	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E503543FFA1A683FF45FA55FA4D.taxon	distribution	Distribution Only recorded from China, Laos and Vietnam. In Vietnam, Hystatoderes occurs in several provinces of northern (formerly Tonkin) and central parts and is represented by the species weissi Lameere, 1915 in Ha Giang province, H. vitalisi Lameere, 1917 in Hoa Binh, Lao Cai and Yen Bai provinces, and H. antonkozlovi sp. nov., which exhibits a more southern distribution and occupies Da Nang, Gia Lai, Kon Tum and Quang Nam provinces.	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E503543FFA1A683FF45FA55FA4D.taxon	description	Description Male (Fig. 2 A, 6 A) Dimensions. – Body length: 25 - 36 mm (allotype ♂: 30 mm). Habitus. – Oblong, rather slender in appearance, with long legs and chestnut-brown colour with a slightly satin finish. Head. – Broader than long, practically hairless except for the presence of disparate hairs behind the eyes, strongly granular, granulation becoming coarse between and in front of the eyes; longitudinal furrow starting between the antennal tubercles and extending behind the vertex; antennal tubercles with deep foveoli and bordered downwards with a groove; clypeus granular with a few scattered golden setae; mandibles strongly punctate, with a slightly curved outer margin in the basal half and then regularly closing	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	description	(Fig. 1, 4 A-C, 6 C, F & H)	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	materials_examined	Material examined Type material studied Holotype ♀, 41 mm (Fig. 4 B-C, 6 F & H): Museum Paris, Tonkin Sept., Frontière de Chine, Ha Giang, A. Weiss 1901 / oct. - dec. / Emphiesmenus WeissiA. Lameere vid. 1915 / TYPE (MNHN) / Holotype Emphiesmenus weissi Lameere, 1915, vid. Drumont et al., 2023. FAUNITAXYS - ISSN (Print): 2269 - 6016 - ISSN (Online): 2970 - 4960 G	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	description	C, F, H. Hystatoderes weissi (Lameere, 1915). C. ♂ (ADC), forebody. F. ♀ (MNHN), forebody. H. ♀, basal part of elytra.	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	materials_examined	Other material studied 1 ♂, Vietnam, Ha Giang province, env. Lung Cu, 1200 m, local collector leg. (ADC) (Fig. 4 A).	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	description	Description Male (Fig. 4 A, 6 C) Dimensions. – Body length: 31 mm. Habitus. – With a stocky appearance (like H. vitalisi), oblong, concolor dark brown, less than twice as long as wide at the basal third of elytra. Pronotum, elytra, scutellum, head and antennae concolor, pronotum with a golden dense pilosity near apical and basal margins. Legs brown with elongated tarsi. Antennae barely exceeding the elytral half. Head. – Robust, slightly wider than long; frons and vertex with a scattered punctation evenly distributed over the surface; median longitudinal groove on the top running between the antennal tubercles towards the rear and bordered by a strong carina; a small other carina bordering the upper part of ocular lobe joining the first one near the middle of the vertex. Eyes moderately indented; lower lobe barely wider than the upper lobe; inter-ocular space equal to the width of the upper lobe. Mandibles regularly rounded and strongly punctate, wider from base to central part, bifid with an intermediate tooth in the basal third. Antennae. – Rather short, just reaching the apical third of the elytra. Scape strongly vermiculated and wrinkled, slightly flattened on dorsal side, enlarged at apex with rounded outer corner, antennomere 3 as long as scape, with irregularly spaced strong punctation; antennomere 4 punctate or with few longitudinal groove; antennomeres 5 – 10 equal in length and expanding bell-shaped towards the apex, with longitudinal grooves; last antennomere longer with bevelled apex. Pronotum. – Transverse, hexagonal, swollen on the disc; surface strongly vermiculate with huge wrinkles, partially hairless except along the apical and basal margin, which have a fringe of long golden hairs; apical margin much wider than head; apical angles irregularly shaped with a fairly strong spine offset downward; a weaker tooth about halfway up the pronotum followed by a strong, slightly downwardfacing spine. Under this last tooth, the pronotum narrows to the basal border without teeth. Scutellum: large, rounded, granular and covered with sparse golden setae. Elytra. – Almost smooth and hairless, showing only areas with weak fine wrinkles and a small punctuation on the basal part around the scutellum, punctuation becoming stronger on the humeral angles; costae inconspicuous; lateral margins relatively wide; apical angles weakly sharp, without spine. Legs. – Dark brown; femora straight, punctate and covered with a scattered pilosity; tibiae punctate, strongly concave inner face and finely crenellated on the lower side, widening towards the apex and with few scattered hairs; apex toothed at both sides; tarsi narrow and elongated; onychium as long as tarsomeres 1 - 3 combined. Female (Fig. 4 B-C, 6 F & H) Dimensions. – Body length holotype ♀: 41 mm. Habitus. – Body moderately large, stout, thicken dorsal ventral side, and brighten red. Head. – Remarkable shorter than wide. Eyes narrow in dorsal view, distantly separated, distance between eyes about haft of wide of the head and wider than dorsal lobe of the eyes. Labrum with some short setae. Mandible strong, robust, curved inward and downward, right mandible with inner apical teeth forming a bifid apex. Underside of head concave and not smooth. Antennae. – Short, about 0.6 as long as body, 11 - segmented; scape the thickest, as long as antennomere 3, round at sides, somehow little pressed in dorsal-ventral direction, expanded apically; antennomere 2 visible; antennomere 3 about as long as scape, mostly rounded but somehow flattened at ventral side, narrow at base and expanded apically; antennomeres 4 - 5 similar in shape and size, antennomeres 4 and 5 combined about as long as antennomere 3; antennomeres 6 - 11 combined about as long as 3 - 5 combined; antennomeres 1 - 9 somehow smooth with punctuation; antennomeres 10 - 11 with wrinkles in the length. Pronotum. – Short, about 0.6 as long as wide (measured between 2 tips of basal spines); apical margin concave; basal margin convex, making a basal lobe; surface smooth on the middle, disc convex and two lateral parts much more punctate, reaching the lateral margins; lateral margin with three evident spines and one fold forming a small spine, the basal one longest and more pointed that others. Except the smallest tooth, all spines are curved upward. Scutellum: short and rounded, making a semicircle with basal margin of pronotum, not smooth as the middle part of pronotum but less punctate than the elytra. Elytra. – Reddish and shining, strong punctuation at base and reduced from the apical fourth; lateral margin curves up strongly from base and nearly to flatten at apex; apex rounded with a short angle. Legs. – Flattened, same colour as the body, tarsi segment with lateral apical somehow point. Diagnostic characters. – Hystatoderes weissi is easily distinguished from the other two species composing the genus by the following combination of characters: - For the males (only one specimen studied): remarkable for the very transverse pronotum, marked with deep wrinkles and provided with three lateral spines, the vertex showing a strong longitudinal carina, and the blackish brown elytra, mat and with basal angles rounded and without tooth. - For the females (only one specimen studied): by the relatively square pronotum with three evident spines and a fold of margin forming a small spine, and the reddish brown elytra with basal angles rounded and without tooth.	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	etymology	Etymology. – Described in honour of the collector A. Weiss. Flight period / biotope altitude. – For H. weissi, only a few data are available, for the altitude the male was recorded at 1200 m and the flight period for the holotype female is October-December.	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
03A77E50354CFFA8A684FA02FAF1FAF3.taxon	distribution	Distribution. – Extreme northern part of Vietnam (Fig. 1)	en	Drumont, Alain, Komiya, Ziro, Do, Cuong, Phan, Quoc Toan, Constant, Jérôme, Ripaille, Claude, -, O. D. (2023): Revision of the genus Hystatoderes Lameere, 1917 (Part I): species from Vietnam (Coleoptera, Cerambycidae, Prioninae, Anacolini). Faunitaxys 11 (43): 1-11, DOI: 10.57800/faunitaxys-11(43), URL: http://dx.doi.org/10.5281/zenodo.15376488
