taxonID	type	description	language	source
03A31D382841066304E1FA20C49BFD7D.taxon	materials_examined	Type material: male holotype, 6 males and 4 females paratypes (MZSP 31022) from Vale Verde, Parque Nacional do Caparaó, Alto Caparaó, Minas Gerais. 20 ° 25 ’ 10 ” S 41 º 50 ’ 45 ” W. 06 / IV / 2009; Pinto-da-Rocha, DaSilva & Bragagnolo; 2 males and 1 female paratypes (MZSP 21732) from Parque Nacional do Caparaó, Alto Caparaó, Minas Gerais. 01 – 05 / V / 2002; Equipe Biota; 4 males, 1 female and 1 immature paratypes (MZSP 29995) from Parque Estadual Serra do Brigadeiro, Fervedouro, Minas Gerais. 21 – 24 / III / 2005; DaSilva, Yamaguti & Nogueira; 1 male paratype (MZSP 31024) from Cachoeira Bonita, Parque Nacional do Caparaó, Alto Caparaó, Minas Gerais. 20 ° 24 ’ 24 ” S 41 ° 50 ’ 20 ” W. 07 / IV / 2009; Pinto-da-Rocha, DaSilva & Bragagnolo; 1 male paratype (MZSP 31023) from Cachoeira do Aurélio, Parque Nacional do Caparaó, Espera Feliz, Minas Gerais. 1,820 m. 20 ° 28 ’ 43 ’’ S 41 ° 49 ’ 45 ’’ W. 08 / IV / 2009; Pinto-da-Rocha, DaSilva & Bragagnolo; 5 males and 2 females paratypes (MZSP 77541) from Parque Nacional do Caparaó, Alto Caparaó, Minas Gerais. 01 – 05 / V / 2009; Equipe Biota.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382841066304E1FA20C49BFD7D.taxon	diagnosis	Diagnosis: The new species is distinguished from others in the genus by the presence of a pair of small tubercles on the ocularium, and on mesotergal area III (instead of large spines, similar to G. macracanthum), presence of a complex dry mark resembling a mask on the dorsal scutum (similar to G. varium), and a ventral process with a flabellum on the penis (similar to G. varium and G. carum).	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382841066304E1FA20C49BFD7D.taxon	etymology	Etymology: In honor of the opilionologist, author of many species of Gonyleptidae, as well as friend, Cibele Bragagnolo, who collected some of the type-specimens of the new species.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382841066304E1FA20C49BFD7D.taxon	distribution	Distribution: slopes of the eastern end of the Mantiqueira range, in the Caparaó and Brigadeiro mountains.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382841066304E1FA20C49BFD7D.taxon	description	Description (values in parentheses, for characters with variations, refer to the holotype, figure 1 A – B): Ocularium with a pair of small tubercles separated from each other and near to the eyes; pair of tubercles of area I indistinct; area III with a pair of small tubercles; carapace with 13 – 33 (33) minute granules; area I with 3 – 9 (6), area II with 6 – 9 (6) and area III with 4 – 6 (4) small granules; dorsal scutum: posterior margin with high density of minute granules; lateral margin with medium density of minute granules; free tergites, anal operculum, stigmatic areas, and free sternites with low density of minute granules. Chelicerae: Segment I with 3 small dorsal basal granules and 1 small apical prolateral granule; segment II with medium density of minute granules. Pedipalps (figure 1 C – D): Trochanter with 2 central dorsal tubercles, 1 central ventral tubercle, and 2 small apical ventral granules; femur with 6 ventral elevations ' IiIiii', and 1 subapical prolateral spine; patella with 1 apical prolateral ventral tubercle; tibia with 1 small basal ventral tubercle, ventral armature with retrolateral ' IiIi' and prolateral ' IiIIi'; tarsus with ventral armature retrolateral ' IiIi' and prolateral ' IIi'. Leg I: Coxa with ventral row of very large granules; trochanter with low density of large ventral granules; femur and patella with small granules; tibia and metatarsus with minute granules. Leg II: Coxa with a ventral row of medium granules; trochanter with low density of ventral medium granules; femur and patella with small granules; tibia and metatarsus with minute granules. Leg III: Coxa with low density of minute granules; trochanter with low density of small granules; femur, patella and tibia with medium granules; metatarsus with minute granules. Leg IV (figure 1 B, E – G): Coxa with low density of minute granules; prolateral apophysis inserted almost dorsally, curved backwards and downwards at the apex; large apical retrolateral apophysis pointed and the same size as the trochanter; trochanter with a small, pointed and upward-curved prolateral sub-basal apophysis, retrolateral apical apophysis pointed, curved backwards and with a granule associated; femur short, with very strong inward curvature, apical retrolateral apophysis very large and curved backwards, prolateral apophysis very small, row 2 with flat granules, with a high density of disorganized granules in the apical third; patella with large prolateral granules; tibia with row 2 with a group of large granules close to each other, row 3 with small spines curved backwards along the entire length, increasing from base to apex; metatarsus with row 3 with small spines in the first half; tarsal process very large. Tarsal segmentation: 8 – 10 (9); 16 – 22 (20); 8 – 10 (9); 8 – 10 (9). Penis (figure 3 A – B): Ventral plate rectangular-shaped, longer than wide, with practically straight lateral margins, only a slight convexity in the apical region due to its elevation; apical margin slightly concave with elevations above the margin at the ends. All macrosetae pointed; group A with 3 macrosetae aligned almost longitudinally and slightly curved downwards; macrosetae B small, positioned more apically and ventrally than the A 1; group C with 3 macrosetae, C 1 being slightly separated from the others and slightly curved backwards; macrosetae D with half size of the macrosetae of group C and positioned below C 3; macrosetae E with half size of the macrosetae of group C and positioned more ventrally and between C 1 and C 2; glans with ventral process long, with a flabellum on apex, with many projections on apical margin, a little wider than the stem; stylus cylindrical with a dorsal apical beak. Measurements. Dorsal scutum. Length: 8.9 – 10.6 (10.6); width: 8.4 – 11.1 (11.1); pedipalps: 13 – 14.1 (13.9); legs: I: 24.1 – 28.8 (28.7); II: 44.6 – 56.6 (56.5); III: 34.9 – 43.2 (43.2); IV: 47 – 59 (58.8). Coloration (photos of live animals; figure 4): predominantly dark brown; chelicerae dark - yellowish; dorsal scutum dark brown, with yellowish grooves and carapace; apexes of trochanters of legs, tubercles of ocularium and granules of areas of dorsal scutum orange to yellow; articular membranes between coxae and trochanters pink; dry mark prominent, on the carapace like ‘ complete mask’, on the abdominal scutum following the grooves between areas I, II and III, between the tubercles of area III as two circles and present on apex of coxa, base of trochanter and femur IV. Female (figure 4): legs less armed and with weaker granulation compared to males; prolateral apophysis of coxa IV small, spine-like; trochanter IV with only a small apical retrolateral apophysis; posterior margin and free tergites with larger granules compared to males, and with a pair of small tubercles at the angles; tarsal processes smaller than in the male (approximately half the size of the tarsal claw). Serracutisoma Roewer 1930 now includes: S. proximum group: S. thalassinum (Simon 1879); S. proximum (Mello-Leitão 1922), type-species; S. molle (Mello-Leitão 1933); S. banhadoae (Soares & Soares 1947); S. fritzmuelleri DaSilva & Gnaspini 2010; S. pseudovarium DaSilva & Gnaspini 2010; and S. gnaspinii DaSilva 2014. S. inerme group: S. inerme (Mello-Leitão 1927); S. spelaeum (Mello-Leitão 1933); S. catarina (Machado, Pinto-da-Rocha & Giaretta 2002); S. guaricana DaSilva & Gnaspini 2010; and S. hara sp. nov..	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382844066104E1FE78C6C9FBDA.taxon	materials_examined	Type material: 1 male holotype (MZSP 18974) from Gruta do Botuverá, Botuverá, Santa Catarina. 9 / V / 1998, R. Pinto-da-Rocha.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382844066104E1FE78C6C9FBDA.taxon	diagnosis	Diagnosis: Similar to other species of S. inerme group. It differs by the combination of armature of femur III with spines close to each other and armature of row 2 of femur IV with spines only in the medial-apical portion.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382844066104E1FE78C6C9FBDA.taxon	etymology	Etymology: In honor of the opilionologist, author of many species of Gonyleptidae, and friend, Marcos Hara.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382844066104E1FE78C6C9FBDA.taxon	distribution	Distribution: Only known from the type locality.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
03A31D382844066104E1FE78C6C9FBDA.taxon	description	Description (figure 2 A – B): Ocularium with 1 pair of medium tubercles, separated and close to the eyes; carapace with 6 minute granules; areas I, II and III with, respectively, 10, 10 and 6 medium granules; area I with 1 pair of small tubercles; area III with 1 pair of medium spines; dorsal scutum: lateral margin with low density of minute granules; posterior margin and free tergites practically smooth; angles of the free tergites with a pair of medium granules. Chelicerae: First segment with 4 dorso-basal tubercles, 1 prolateral dorso-apical tubercle and 1 apical retrolateral tubercle; second segment with a medium density of granules. Pedipalps (figure 2 C): Trochanter with 2 medium granules and 1 apical ventral setiferous tubercle; femur with 8 ventral tubercles ‘ IiiiIiii’, and 1 prolateral spine; patella with 1 large prolateral granule; tibia with elevation pattern on the retrolateral ventral setae ‘ IiIi’, the penultimate larger, more thicker than the others, and fused with the last at the base and prolateral setae ‘ IiIi’, with 4 small ventral granules and 1 small retrolateral tubercle pointed; tarsus with prolateral ventral setae ‘ IIi’ and retrolateral setae ‘ IiIi’. Leg I: Coxa with 1 row of medium tubercles; trochanter with 1 large granule and 2 medium granules; femur, patella, and tibia with minute granules; metatarsus practically smooth. Leg II: Coxa with 1 row of small tubercles; trochanter with 1 medium granule and low density of small granules; femur, patella, and tibia with minute granules, row 4 of tibia with sparse small spines; metatarsus practically smooth. Leg III (figure 2 D): Coxa with 1 row of minute granules; trochanter with 1 medium granule and a low density of small granules; femur with row 3 with spines close to each other in the apical region; femur, patella, and tibia with minute granules; metatarsus practically smooth. Leg IV (figure 2 E): Coxa with low density of minute granules, apical prolateral apophysis long and almost transverse with falciform apex, apical retrolateral apophysis like a small tubercle; trochanter with low density of minute granules, basal prolateral apophysis as a strong tubercle, apical dorsal prolateral apophysis curved upwards, apical retrolateral apophysis small, pointed, and with half size of the dorsal-prolateral apophysis; femur slightly S-shaped almost straight, with row 2 with 2 strong spines, close to each other, in the median to last third region; femur, patella and tibia with medium spines in apical portion; metatarsus with minute granules; tarsal claws smooth; tarsal process very small. Tarsal segmentation: 9 – 10; 16 – 17; 11; 11 – 12. Penis (figure 3 C – D): Ventral plate hexagonal notably broader than long, lateral margin convex and apical margin slightly concave; all macrosetae spatulate; group A with 3 macrosetae arranged almost transversely; macrosetae B similar and aligned with the macrosetae of group A; group C with 3 pairs of macrosetae, C 1 separated of the others; macroseta D half the size of the macrosetae of group C, aligned basally to group C; two macrosetae E half the size of the macrosetae of group C, E 1 positioned between the two apical macrosetae of group C (C 1 and C 2) and E 2 more ventral and between the two more basal macrosetae C (C 2 and C 3); glans has a long ventral process with a wide flabellum at the apex, with many projections on ventral margin; stylus straight and cylindrical. Measurements. Dorsal scutum. Length: 8.6; width: 8.3; pedipalps: 13.8; legs: I: 47.2; II: 102.8; III: 71.9; IV: 95.3. Coloration (in alcohol): dorsal scutum, legs I – III, tibia, metatarsus IV, pedipalps and spines of area III have a light-yellow coloration; carapace, apophyses of the coxa and trochanter IV, leg IV, trochanters of the other legs, and the ventral side of the body dark brown; articular membranes light pink; dry mark only on trochanter IV and the base of femur IV. Female: Unknown.	en	Gomes-Silva, José Geovane, Dasilva, Marcio B. (2025): Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil. Zootaxa 5604 (1): 16-28, DOI: 10.11646/zootaxa.5604.1.2, URL: https://doi.org/10.11646/zootaxa.5604.1.2
