identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A34D63FFBAD333FF27F9EB12BBB090.text	03A34D63FFBAD333FF27F9EB12BBB090.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tritegeus Berlese 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Tritegeus Berlese, 1913 (=  Sphodrocepheus Woolley &amp; Higgins, 1963 ) </p>
            <p> Type species:  Tritegeus bisulcatus Grandjean, 1953</p>
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	https://treatment.plazi.org/id/03A34D63FFBAD333FF27F9EB12BBB090	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBAD330FF27F93D125BB0E2.text	03A34D63FFBAD330FF27F93D125BB0E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tritegeus Berlese 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Synonymy of  Tritegeus and  Sphodrocepheus</p>
            <p> Woolley and Higgins (1963) proposed the genus  Sphodrocepheus without comparison with related genera. Balogh (1972) considered that this genus differs from  Tritegeus by having a pair of the longitudinal dorsolateral thickenings bordering the centrodorsal part of the notogaster, which bear the dorsolateral setae (whereas  Tritegeus lacks these thickenings). Sitnikova (1975 b) observed that  Sphodrocepheus is distinguished from  Tritegeus by the presence of the very large lamellae that cover the rostrum and are connected by a broad translamella (in contrast,  Tritegeus has lamellae that do not cover the rostrum and the translamella is not broad). Balogh and Balogh (1992) considered that  Sphodrocepheus differs from  Tritegeus by the presence of dorsolateral thickenings and rough sculpture on the notogaster, which are absent in  Tritegeus . </p>
            <p> However, a comparison of the descriptions and supplementary descriptions of the type species of  Sphodrocepheus and  Tritegeus —  S. tridactylus (see data of this paper) and  T. bisulcatus (see S. Bernini &amp; Bernini 1990; Pérez-Íñigo 1997; Weigmann 2006), respectively—reveals that the previously noted differences between these genera are inaccurate. Both  S. tridactylus and  T. bisulcatus share the following traits: 1) longitudinal dorsolateral thickenings bordering the centrodorsal part of the notogaster and bearing dorsolateral setae; 2) lamellae that do not cover the rostrum and are narrowly separated medially (translamella absent); 3) absence of rough sculpture on the notogaster. Woolley and Higgins (1963; Fig. 1 on p. 146) figured a wide translamella in  S. tridactylus , but their figure is inaccurate. In reality (see supplementary description below) the lamellae are separated, with their medial parts slightly protruding anteromedially, creating the illusion of a thin translamella. </p>
            <p> Hence, there is no difference between the genera  Sphodrocepheus and  Tritegeus . Therefore, I support the assumption of Bayartogtokh and Ermilov (2021) regarding the synonymy of  Sphodrocepheus with  Tritegeus , and propose final taxonomic proposals accordingly:  Tritegeus Berlese, 1913 (=  Sphodrocepheus Woolley &amp; Higgins, 1963 syn. nov. );  Tritegeus anthelionus (Woolley &amp; Higgins, 1968) comb. nov. ;  Tritegeus dentatus (Fujikawa, 1972) comb. nov. ;  Tritegeus mitratus (Aoki, 1967) comb. nov. ;  Tritegeus tridactylus (Woolley &amp; Higgins, 1963) comb. nov. ;  Tritegeus tuberculatus (Mahunka, 1988) comb. nov.</p>
            <p>Generic diagnosis</p>
            <p> Adult. With character states of  Cepheusidae (e.g. Sitnikova 1975 a; S. Bernini &amp; Bernini 1990; Weigmann 2006; Norton &amp; Behan-Pelletier 2009). Body size: Large (length more than 600). Integument: Surface microsculpturing microtuberculate; additionally, prodorsum partially foveate, notogaster frequently areolate, podosomal region partially tuberculate, anogenital region can be partially foveate or rugose. Prodorsum: Rostrum rounded or pointed. Lamella located dorsolaterally on prodorsum, comparatively long and broad; lamellae separated or connected by translamella mediodistally, sometimes entirely fused; cusp of lamella with outer tooth. Prolamella absent or vestigial. Sublamella and sublamellar porose area absent. Tutorium long, ridge-like, heavily dentate dorsally. Rostral, lamellar and interlamellar setae comparatively long, setiform/subflagellate/rod-like/thickened: le inserted on lamellar cusp; in inserted in interbothridial region; bothridial seta with or without head; exobothridial seta present. Bothridium cupshaped. Interbothridial region without tubercles. Dorsosejugal porose area and dorsophragma absent; pleurophragma present. Genal tooth large, elongate triangular. Notogaster: Anterior notogastral margin convex or straight medially. Humeral region with small, rounded projection. Pteromorph and octotaxic system absent. Ten pairs of notogastral setae (centrodorsal part of notogaster without setae) mainly rod-like/thickened, sometimes slightly dilated distally, inserted on notogastral surface or on tubercles; setae h 1 and p 1 inserted close to each other and usually shorter than dorsal setae.  Gnathosoma . Subcapitulum diarthric, with setiform subcapitular setae. Palp with setation: 0–2–1–3– 9(+ω); solenidion bacilliform, coupled with eupathidium mediodistally. Axillary saccule absent. Chelicera chelatedentate. Epimeral and lateral podosomal regions: Epimeral setal formula: 3–1–3–3; all epimeral setae setiform. Enantiophyses and tubercles absent. Pedotectum I represented by large scale divided into two parts (dorsal part triangular; ventral part simple); pedotectum II represented by small scale. Humeral porose area absent. Circumpedal carina and discidium present. Anogenital region: Setal formula: 6–1–2–3; all setae comparatively short, mainly setiform. Marginal porose area absent. Adanal lyrifissure oblique or transverse, lateral and slightly distanced from anterior margin of anal plate. Legs: All legs heterotridactylous, rarely monodactylous. Dorsoparaxial porose area on femora I–IV and on trochanters III, IV present; distoventral porose area on tibiae I–IV and proximoventral porose area in tarsi I–IV present or not observable. </p>
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	https://treatment.plazi.org/id/03A34D63FFBAD330FF27F93D125BB0E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB9D334FF27F8EE1236B41A.text	03A34D63FFB9D334FF27F8EE1236B41A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tritegeus tridactylus (Woolley & Higgins 1963) Ermilov 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tritegeus tridactylus (Woolley &amp; Higgins, 1963)</p>
            <p>(Figs 1, 2)</p>
            <p> Material.  Two specimens (two females): USA, Utah (G.F. Knowlton). The specimens were compared to the type material for a definite identification . </p>
            <p> All specimens (preserved in 70% solution of ethanol with a drop of glycerol) are deposited in the collection of the University of Tyumen,   Museum of Zoology , Tyumen, Russia  . </p>
            <p>Supplementary description of adult. Measurements. Body length: 765, 780; notogastral width: 570, 600.</p>
            <p>Integument (Figs 1A–C). Body color dark brown. Body and legs partially covered by thick gel-like layer of cerotegument, including dense microtubercles, sparsely located on notogaster and in marginal part of ventral plate. Surface with dense microtuberculate sculpturing; lamella, humeral process, pedotecta I, II, podosomal region, discidium, lateral part of prodorsum, lateral part of epimere I partially foveate; medioposterior part of notogaster (between setae lm, lp, h 2 h 3) with sparse, indistinct foveolae; lateral side of body between humeral process and acetabula II, III with dense large tubercles; additionally, podosomal region and dorsal part of pedotectum I partially rugose.</p>
            <p>Prodorsum (Figs 1A–C). Rostrum pointed, observable in dorsal aspect. Rostral region with bulge. Outer tooth of lamellar cusp well developed. Translamella absent but medial parts of lamellae slightly protruding anteromedially, creating an illusion of the presence of a thin translamella. Tutorium with numerous teeth and small scales dorsally. Rostral seta (90–94) setiform, with short, flexible tip, barbed; lamellar seta (105–109) with thickened mediobasal part and flagellate distal part, barbed; interlamellar seta (217–225) thickened, acuminate, distinctly barbed; bothridial seta (124–131) with longer stalk and short expanded head bearing numerous strong spines; exobothridial seta represented by alveolus (may be broken).</p>
            <p>Notogaster (Figs 1A–D). Centrodorsal region bordered by U-shaped thickening bearing dorsal notogastral setae. Setae h 1 (60–75) and p 1 (64–75) thickened, acuminate, heavily barbed; other setae (c, la, lm: 157–169; lp, h 2, h 3: 201–225; p 2, p 3: 134–146) thickened, acuminate, distinctly barbed. Opisthonotal gland opening and all lyrifissures observable.</p>
            <p> Gnathosoma (Figs 2A–C). Subcapitulum size: 165–172 × 116; subcapitular setae (a: 34; m: 15–19; h: 45–52) setiform, roughened; m thinnest; both adoral setae (15–19) setiform, smooth. Palp length: 101–109; postpalpal seta (11–13) spiniform, truncate, roughened. Chelicera length: 172–184; setae (cha: 43–45; chb: 28–30) setiform, barbed. </p>
            <p>Epimeral and podosomal regions (Figs 1B, C). All epimeral setae (3c, 4c: 67–75; 1b: 45–52; 3b, 4a, 4b: 34–37; 1a, 1c, 2a, 3a: 22) setiform, roughened. Discidium large, elongate triangular, rounded distally.</p>
            <p>Anogenital region (Figs 1B–D). Genital (g 1, g 2: 34–37; g 3: 26–30; others: 19–22) and aggenital (26–30) setae setiform, roughened; both anal setae (22–26) setiform, barbed; all adanal setae (37–41) slightly thickened, acuminate, comparatively heavily barbed. Adanal lyrifissure distinct, oblique.</p>
            <p> Legs (Figs 2D–G). All tarsi with three claws; median claw thicker than lateral claws; all claws slightly barbed on dorsal side. Trochanters III and IV with long tooth dorsoanteriorly. Formulas of leg setation and solenidia: I (1- 5-3-4-20) [1-2-2], II (1-4-3-4-16) [1-1-2], III (2-3-2-3-15) [1-1-0], IV (1-2-3-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1; comparison of morphology of some setae between  T. tridactylus ,  T. luissantossubiasi and  T. mitratus presented in Table 2. </p>
            <p>Note: Roman letters refer to normal setae; Greek letters to solenidia (except ɛ = famulus); single quotation mark (’) marks setae on the anterior and double quotation mark (”) setae on the posterior side of a given leg segment; parentheses refer to a pair of setae.</p>
            <p> Remarks. Based on the original description (Woolley &amp; Higgins, 1963) and the data presented here, the following diagnostic traits for  T. tridactylus are proposed: </p>
            <p>Adult. Body size: 720–780. Medioposterior part of notogaster (between setae la, lp, h 2 h 3) with sparse, indistinct foveolae. Rostrum pointed. Lamellar cusps separated medially; translamella absent. Rostral seta setiform, with short, flexible tip, barbed; lamellar seta with thickened mediobasal part and flagellate distal part, with slight and sparse barbs; interlamellar seta thickened, acuminate, distinctly barbed; in˃le˃ro; bothridial seta with shortly expanded head bearing numerous strong spines. Centrodorsal region of notogaster bordered by U-shaped thickening. Notogastral setae h 1 and p 1 comparatively short, slightly different in length, thickened, acuminate, heavily barbed; other setae long, thickened, acuminate, distinctly barbed; lp, h 2, h 3 ˃c, la, lm. All leg tarsi heterotridactylous; trochanters III and IV with long tooth dorsoanteriorly; seta pv’ on Ta I, II barbed (without ventral sparse spines); l” and v’ on Ti I, II thorn-like; l” on Ge I, II setiform.</p>
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	https://treatment.plazi.org/id/03A34D63FFB9D334FF27F8EE1236B41A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.text	03A34D63FFBDD33BFF27FDBA113BB53A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tritegeus luissantossubiasi Ermilov 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tritegeus luissantossubiasi sp. nov.</p>
            <p>(Figs 3, 4)</p>
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                 Type material.   Holotype (female) and three paratypes (three females): India, 28°36'56''N, 95°53'21''E, Arunachal Pradesh State,  
                <a title="Search Plazi for locations around (long 92.65222/lat 27.013334)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.65222&amp;materialsCitation.latitude=27.013334">Etalin</a>
                 , 700 m a.s.l., soil with litter in forest, 1–8.v.2012 (L. Dembický and O. Šauša). Two paratypes (two females):  India, 28°19'32''N 95°57'31''E, Arunachal Pradesh State, Hunli, 1300 m a.s.l., soil with litter in forest, 26.v.–1.vi.2012 (L. Dembický and O. Šauša). Two paratypes (two females): India, 27°00'48''N 92°39'08''E, Assam State, Bhalukpong, 150 m a.s.l., soil with litter in forest, 1–8.v.2012 (L. Dembický and O. Šauša).
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            <p> The holotype is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, Germany;   seven paratypes are deposited in the collection of the  University of Tyumen, Museum of Zoology, Tyumen, Russia  . All specimens are preserved in 70% solution of ethanol with a drop of glycerol. </p>
            <p>Diagnosis. Adult. Body size: 690–825. Medioposterior part of notogaster (between setae lp, h 2 h 3) with sparse, indistinct foveolae. Rostrum pointed. Lamellar cusps separated medially; translamella absent. Rostral seta setiform, with short, flexible tip, barbed; lamellar seta with thickened mediobasal part and flagellate distal part, with slight and sparse barbs; interlamellar seta thickened, acuminate, with slight and sparse barbs; in˃le˃ro; bothridial seta with shortly expanded head bearing numerous strong spines. Centrodorsal region of notogaster bordered by Ushaped thickening. Notogastral setae h 1 and p 1 comparatively short (p 1 ˃h 1), thickened, acuminate, heavily barbed; other setae long, thickened, acuminate, with slight and sparse barbs but with some strong barbs in distal part. All leg tarsi heterotridactylous; trochanters III and IV with long tooth dorsoanteriorly; seta pv’ on Ta I, II with ventral sparse spines; l” on Ti I, II and Ge I, II thickened; v’ on Ti I, II setiform, with ventral sparse spines.</p>
            <p>Description of adult. Measurements. Body length: 780 (holotype), 690–825 (paratypes); notogastral width: 600 (holotype), 555–600 (paratypes).</p>
            <p>Integument (Figs 3A–D). Body color dark brown. Body and legs partially covered by thick gel-like layer of cerotegument, including dense microtubercles. Surface with dense microtuberculate sculpturing, clusters of larger tubercles close and lateral to notogastral U-shaped thickening; lamella, humeral process, pedotecta I, II, podosomal region, discidium, lateral part of prodorsum, lateral part of epimere I partially foveate; medioposterior part of notogaster (between setae lp, h 2 h 3) with sparse, indistinct foveolae; lateral side of body between humeral process and acetabula II–IV with dense large tubercles; additionally, humeral process, podosomal region and dorsal part of pedotectum I partially rugose.</p>
            <p>Prodorsum (Figs 3A–C). Rostrum pointed, observable in dorsal aspect. Rostral region with bulge. Outer tooth of lamellar cusp well developed. Translamella absent but medial parts of lamellae slightly protruding anteromedially, creating an illusion of the presence of a thin translamella. Tutorium with numerous teeth and small scales dorsally. Rostral seta (90–105) setiform, with short, flexible tip, barbed; lamellar seta (154–176) with thickened mediobasal part and flagellate distal part, with slight and sparse barbs; interlamellar seta (172–206) thickened, acuminate, with slight and sparse barbs; bothridial seta (90–112) with longer stalk and short expanded head bearing numerous strong spines; exobothridial seta represented by alveolus (may be broken).</p>
            <p>Notogaster (Figs 3A–D). Centrodorsal region bordered by U-shaped thickening bearing dorsal notogastral setae. Setae h 1 (56–64) and p 1 (86–94) thickened, acuminate, heavily barbed; other setae (c, la, lm, lp, h 2, h 3: 146– 161; p 2, p 3: 101–124) thickened, acuminate, with slight and sparse barbs but with some strong barbs in distal part. Opisthonotal gland opening and all lyrifissures observable.</p>
            <p> Gnathosoma (Figs 4A–C). Subcapitulum size: 154–161 × 123–135; subcapitular setae (a: 34–41; m: 13–15; h: 30–41) setiform, roughened; m thinnest; both adoral setae (15–19) setiform, smooth. Palp length: 94–97; postpalpal seta (11) spiniform, truncate, roughened. Chelicera length: 161–155; setae (cha: 37–41; chb: 28–30) setiform, barbed. </p>
            <p>Epimeral and podosomal regions (Figs 3B, C). All epimeral setae (3c: 75; 4c: 56–71; 3b: 56–60; 1b, 4a, 4b: 41–52; 1c: 34–37; 1a, 2a, 3a: 15–19) setiform, roughened. Discidium large, elongate triangular, rounded distally.</p>
            <p>Anogenital region (Figs 3B–D). Genital (g 1, g 2: 37–45; g 3: 30–37; others: 22–30), aggenital (30–37), anal (19– 26), and adanal (41–56) setae setiform, roughened. Adanal lyrifissure distinct, oblique.</p>
            <p> Legs (Figs 4D–G). All tarsi with three claws; median claw thicker than lateral claws; all claws slightly barbed on dorsal side. Trochanters III and IV with long tooth dorsoanteriorly. Formulas of leg setation and solenidia: I (1- 5-3-4-20) [1-2-2], II (1-4-3-4-16) [1-1-2], III (2-3-2-3-15) [1-1-0], IV (1-2-3-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1; comparison of morphology of some setae between  T. luissantossubiasi ,  T. tridactylus and  T. mitratus presented in Table 2. </p>
            <p> Remarks.  Tritegeus luissantossubiasi is morphologically most similar to  T. tridactylus in main traits: rostrum triangular; bothridial seta with developed head; cusps of the lamellae separated medially; centrodorsal region of the notogaster bordered by U-shaped thickening, without rugose sculpturing; dorsal notogastral setae long (distinctly longer than distance lm–lp), thickened, acuminate; tridactylous legs. However, the new species differs from the latter by shorter dorsal notogastral setae lp, h 2, h 3 (not longer than distance lp– h 2 versus longer), roughened, with sparse barbs in distal part (versus distinctly and entirely barbed); the length of the posterior notogastral setae (p 1 distinctly longer than h 1 versus slightly different in length), and the morphology of the seta v’ on the leg tibiae I, II (setiform, with ventral sparse spines versus thorn-like). </p>
            <p>Etymology. This species is named after the late Prof. Dr. Luis. S. Subías, the famous taxonomist in oribatology.</p>
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	https://treatment.plazi.org/id/03A34D63FFBDD33BFF27FDBA113BB53A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB2D33EFF27FC5612A8B71E.text	03A34D63FFB2D33EFF27FC5612A8B71E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tritegeus mitratus (Aoki 1967) Ermilov 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tritegeus mitratus (Aoki, 1967)</p>
            <p>(Figs 5, 6)</p>
            <p> Material.   Three specimens (one male and two females): Japan,  Shikoku ,  Ishizuchi Mountain National Park , Tsuchigoya, Mt. Kamegamori, 1650 m a.s.l., abies and moss litter, with fungi and moss 15.VIII.1980 (S. Peck)  . </p>
            <p> All specimens (preserved in 70% solution of ethanol with a drop of glycerol) are deposited in the collection of the University of Tyumen,   Museum of Zoology , Tyumen, Russia  . </p>
            <p>Supplementary description of adult. Measurements. Body length: 780–825; notogastral width: 550–600.</p>
            <p>Integument (Figs 5A–C). Body color dark brown. Body and legs partially covered by thick gel-like layer of cerotegument, including dense microtubercles, larger near the U-shaped thickening on notogaster; lamella, epimere I, pedotecta I, II, discidium, lateral part of prodorsum partially foveate; lateral side of body between humeral process and acetabula II – IV with dense large tubercles; anogenital region with heavy ridges; marginal region of notogaster with some longitudinal ridges; additionally, humeral process, podosomal region and dorsal part of pedotectum I partially rugose.</p>
            <p>Prodorsum (Figs 5A–C). Rostrum pointed, not observable in dorsal aspect. Rostral region with short transverse ridge. Lamellar cusps fused mediodistally on significant continuation (hence, translamella absent); outer tooth of lamellar cusp well developed. Tutorium with numerous teeth and small scales dorsally. Rostral seta (86–94) setiform, with short, flexible tip, barbed; lamellar seta (131–154) with thickened mediobasal part and flagellate distal part, with sparse barbs; interlamellar seta (176–195) thickened, acuminate, roughened; bothridial seta (90–94) with longer stalk and short expanded head bearing numerous strong spines; exobothridial seta (56–60) setiform, roughened.</p>
            <p>Notogaster (Figs 5A–D). Centrodorsal region bordered by U-shaped thickening bearing dorsal notogastral setae. Setae h 1 (49–56) and p 1 (56–64) thickened, acuminate, heavily barbed; p 2, p 3 (34–41) slightly stiff, roughened; other setae (86–94) thickened, acuminate, roughened. Opisthonotal gland opening and all lyrifissures observable.</p>
            <p> Gnathosoma (Figs 6A–C). Subcapitulum size: 146–157 × 105–112; subcapitular setae (a: 30–34; m: 19–22; h: 30–37) setiform, roughened; m thinnest; both adoral setae (17–19) setiform, smooth. Palp length: 82–86; postpalpal seta (11) spiniform, roughened. Chelicera length: 146–161; setae (cha: 34–37; chb: 26–30) setiform, barbed. </p>
            <p>Epimeral and podosomal regions (Figs 5B, C). All epimeral setae (3c, 4c: 56–64; 1b: 37–45; 1c, 3b, 4a, 4b: 30; 1a, 2a, 3a: 11) setiform, roughened. Discidium large, elongate triangular, rounded distally.</p>
            <p>Anogenital region (Figs 5B–D). Genital (g 1, g 2: 26–30; g 3: 22–26; others: 15–19), aggenital (22–26), anal (22– 26), and adanal (ad 1: 30–34; others: 26–30) setae setiform, roughened. Adanal lyrifissure distinct, oblique.</p>
            <p> Legs (Figs 6D–G). All tarsi with three claws; median claw thicker than lateral claws; all claws slightly barbed on dorsal side. Trochanters III and IV with long tooth dorsoanteriorly. Formulas of leg setation and solenidia: I (1- 5-3-4-20) [1-2-2], II (1-4-3-4-16) [1-1-2], III (2-3-2-3-15) [1-1-0], IV (1-2-3-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1; comparison of morphology of some setae between  T. mitratus ,  T. tridactylus and  T. luissantossubiasi presented in Table 2. </p>
            <p> Remarks. Based on the original description (Aoki 1967) and the data presented here, the following diagnostic traits for  T. mitratus are proposed: </p>
            <p>Adult. Body size: 780–840. Notogaster foveolate. Rostrum pointed. Lamellar cusps fused mediodistally (rostrum not observable in dorsal aspect). Rostral seta setiform, with short, flexible tip, barbed; lamellar seta with thickened mediobasal part and flagellate distal part, with sparse barbs; interlamellar seta thickened, acuminate, roughened; in˃le˃ro; bothridial seta with shortly expanded head bearing numerous strong spines. Centrodorsal region of notogaster bordered by U-shaped thickening. Notogastral setae h 1 and p 1 comparatively short (p 1 slightly longer than h 1) thickened, acuminate, heavily barbed; p 2, p 3 shortest, slightly stiff, roughened; other setae medium-sized, thickened, acuminate, roughened. All leg tarsi heterotridactylous; trochanters III and IV with long tooth dorsoanteriorly; seta pv’ on Ta I, II with ventral sparse spines; l” on Ti I, II setiform; v’ on Ti I, II setiform, with ventral sparse spines; l” on Ge I, II setiform.</p>
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	https://treatment.plazi.org/id/03A34D63FFB2D33EFF27FC5612A8B71E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB7D33EFF27FEB615B8B38D.text	03A34D63FFB7D33EFF27FEB615B8B38D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tritegeus Berlese 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to known species of  Tritegeus (=  Sphodrocepheus ) </p>
            <p> 1. All legs monodactylous; cusp of lamella dentate laterally; body length: 700–714..........................................................................  Tritegeus dentatus (Fujikawa, 1972) . Distribution: Eastern Palaearctic region. </p>
            <p>- All legs heterotridactylous; cusp of lamella not dentate laterally................................................ 2</p>
            <p>2. Bothridial seta setiform or bacilliform, without distinct head (sometimes with unclear thickening)..................... 3</p>
            <p>- Bothridial seta with well-developed head.................................................................. 5</p>
            <p> 3. Rostrum pointed; notogastral seta la inserted posteriorly to c; body length: 725–860.......  Tritegeus bisulcatus Grandjean, 1953 (see also S. Bernini &amp; Bernini 1990; Pérez-Íñigo 1997; Weigmann 2006). Distribution: Western Palaearctic region. </p>
            <p>- Rostrum rounded; notogastral seta la inserted posterolaterally to c .............................................. 4</p>
            <p> 4. Bothridial seta long, longer than interlamellar seta; translamella present; body length: 1018.........................................................  Tritegeus major Golosova &amp; Karppinen, 1984 . Distribution: Eastern Palaearctic region. </p>
            <p> - Bothridial seta medium-sized, shorter than interlamellar seta; translamella absent; body length: 888–990.........................  Tritegeus brevisetus Sitnikova, 1975 (in 1975 a). Distribution: Eastern Mediterranean, Eastern Palaearctic region. </p>
            <p> 5. Cusps of lamellae entirely fused medially; body length: 780–840....................................................................  Tritegeus mitratus (Aoki, 1967) (also data presented here). Distribution: Eastern Palaearctic region. </p>
            <p>- Cusps of lamellae entirely or partially separated medially..................................................... 6</p>
            <p> 6. Interlamellar and dorsal notogastral (c, la, lm, lp, h 2, h 3) setae inserted on strong tubercles; body length: 681..................................................  Tritegeus tuberculatus (Mahunka, 1988) . Distribution: Vietnam, Nepal, Mexico. </p>
            <p>- Interlamellar and dorsal notogastral (c, la, lm, lp, h 2, h 3) setae not inserted on strong tubercles........................ 7</p>
            <p> 7. Centrodorsal part of notogaster simultaneously with areolae and ridges; notogastral seta h 1 swollen distally; body length: 750................................  Tritegeus sculptus S. Bernini &amp; Bernini, 1990 . Distribution: Western Mediterranean. </p>
            <p>- Centrodorsal part of notogaster not simultaneously with areolae and ridges; notogastral seta h 1 not swollen distally....... 8</p>
            <p>8. Dorsal notogastral setae la, lm, lp, h 2, h 3 medium-sized (as long as distance lm–lp)................................. 9</p>
            <p>- Dorsal notogastral setae la, lm, lp, h 2, h 3 long (distinctly longer than distance lm–lp)............................... 10</p>
            <p> 9. Bothridial seta medium-sized, distinctly shorter than interlamellar seta; posterior notogastral seta p 1 distinctly longer than h 1; notogaster with areolae; body length: 647–747......  Tritegeus janosbaloghi Ermilov &amp; Martens, 2022 . Distribution: Nepal. </p>
            <p> - Bothridial seta long, not shorter than interlamellar seta; posterior notogastral setae h 1 and p 1 slightly different in length; notogaster without areolae; body length: 870...................................................................................................  Tritegeus anthelionus (Woolley &amp; Higgins, 1968) . Distribution: Nearctic region. </p>
            <p> 10. Dorsal notogastral setae lp, h 2, h 3 longer (longer than distance lp– h 2), distinctly barbed; posterior notogastral setae h 1 and p 1 slightly different in length; seta v’ on leg tibiae I, II thorn-like; body length: 720–780..............................................  Tritegeus tridactylus (Woolley &amp; Higgins, 1963) (also data presented here). Distribution: Holarctic region. </p>
            <p> - Dorsal notogastral setae lp, h 2, h 3 shorter (not longer than distance lp– h 2), roughened, with sparse barbs in distal part; posterior notogastral seta p 1 distinctly longer than h 1; seta v’ on leg tibiae I, II setiform, with ventral sparse spines; body length: 690– 825....................................................  Tritegeus luissantossubiasi sp. nov. Distribution: India. </p>
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	https://treatment.plazi.org/id/03A34D63FFB7D33EFF27FEB615B8B38D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
