taxonID	type	description	language	source
03A34D63FFBAD333FF27F9EB12BBB090.taxon	materials_examined	Type species: Tritegeus bisulcatus Grandjean, 1953	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBAD330FF27F93D125BB0E2.taxon	description	Woolley and Higgins (1963) proposed the genus Sphodrocepheus without comparison with related genera. Balogh (1972) considered that this genus differs from Tritegeus by having a pair of the longitudinal dorsolateral thickenings bordering the centrodorsal part of the notogaster, which bear the dorsolateral setae (whereas Tritegeus lacks these thickenings). Sitnikova (1975 b) observed that Sphodrocepheus is distinguished from Tritegeus by the presence of the very large lamellae that cover the rostrum and are connected by a broad translamella (in contrast, Tritegeus has lamellae that do not cover the rostrum and the translamella is not broad). Balogh and Balogh (1992) considered that Sphodrocepheus differs from Tritegeus by the presence of dorsolateral thickenings and rough sculpture on the notogaster, which are absent in Tritegeus. However, a comparison of the descriptions and supplementary descriptions of the type species of Sphodrocepheus and Tritegeus — S. tridactylus (see data of this paper) and T. bisulcatus (see S. Bernini & Bernini 1990; Pérez-Íñigo 1997; Weigmann 2006), respectively — reveals that the previously noted differences between these genera are inaccurate. Both S. tridactylus and T. bisulcatus share the following traits: 1) longitudinal dorsolateral thickenings bordering the centrodorsal part of the notogaster and bearing dorsolateral setae; 2) lamellae that do not cover the rostrum and are narrowly separated medially (translamella absent); 3) absence of rough sculpture on the notogaster. Woolley and Higgins (1963; Fig. 1 on p. 146) figured a wide translamella in S. tridactylus, but their figure is inaccurate. In reality (see supplementary description below) the lamellae are separated, with their medial parts slightly protruding anteromedially, creating the illusion of a thin translamella. Hence, there is no difference between the genera Sphodrocepheus and Tritegeus. Therefore, I support the assumption of Bayartogtokh and Ermilov (2021) regarding the synonymy of Sphodrocepheus with Tritegeus, and propose final taxonomic proposals accordingly: Tritegeus Berlese, 1913 (= Sphodrocepheus Woolley & Higgins, 1963 syn. nov.); Tritegeus anthelionus (Woolley & Higgins, 1968) comb. nov.; Tritegeus dentatus (Fujikawa, 1972) comb. nov.; Tritegeus mitratus (Aoki, 1967) comb. nov.; Tritegeus tridactylus (Woolley & Higgins, 1963) comb. nov.; Tritegeus tuberculatus (Mahunka, 1988) comb. nov.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBAD330FF27F93D125BB0E2.taxon	diagnosis	Generic diagnosis Adult. With character states of Cepheusidae (e. g. Sitnikova 1975 a; S. Bernini & Bernini 1990; Weigmann 2006; Norton & Behan-Pelletier 2009). Body size: Large (length more than 600). Integument: Surface microsculpturing microtuberculate; additionally, prodorsum partially foveate, notogaster frequently areolate, podosomal region partially tuberculate, anogenital region can be partially foveate or rugose. Prodorsum: Rostrum rounded or pointed. Lamella located dorsolaterally on prodorsum, comparatively long and broad; lamellae separated or connected by translamella mediodistally, sometimes entirely fused; cusp of lamella with outer tooth. Prolamella absent or vestigial. Sublamella and sublamellar porose area absent. Tutorium long, ridge-like, heavily dentate dorsally. Rostral, lamellar and interlamellar setae comparatively long, setiform / subflagellate / rod-like / thickened: le inserted on lamellar cusp; in inserted in interbothridial region; bothridial seta with or without head; exobothridial seta present. Bothridium cupshaped. Interbothridial region without tubercles. Dorsosejugal porose area and dorsophragma absent; pleurophragma present. Genal tooth large, elongate triangular. Notogaster: Anterior notogastral margin convex or straight medially. Humeral region with small, rounded projection. Pteromorph and octotaxic system absent. Ten pairs of notogastral setae (centrodorsal part of notogaster without setae) mainly rod-like / thickened, sometimes slightly dilated distally, inserted on notogastral surface or on tubercles; setae h 1 and p 1 inserted close to each other and usually shorter than dorsal setae. Gnathosoma. Subcapitulum diarthric, with setiform subcapitular setae. Palp with setation: 0 – 2 – 1 – 3 – 9 (+ ω); solenidion bacilliform, coupled with eupathidium mediodistally. Axillary saccule absent. Chelicera chelatedentate. Epimeral and lateral podosomal regions: Epimeral setal formula: 3 – 1 – 3 – 3; all epimeral setae setiform. Enantiophyses and tubercles absent. Pedotectum I represented by large scale divided into two parts (dorsal part triangular; ventral part simple); pedotectum II represented by small scale. Humeral porose area absent. Circumpedal carina and discidium present. Anogenital region: Setal formula: 6 – 1 – 2 – 3; all setae comparatively short, mainly setiform. Marginal porose area absent. Adanal lyrifissure oblique or transverse, lateral and slightly distanced from anterior margin of anal plate. Legs: All legs heterotridactylous, rarely monodactylous. Dorsoparaxial porose area on femora I – IV and on trochanters III, IV present; distoventral porose area on tibiae I – IV and proximoventral porose area in tarsi I – IV present or not observable.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB9D334FF27F8EE1236B41A.taxon	description	(Figs 1, 2)	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB9D334FF27F8EE1236B41A.taxon	materials_examined	Material. Two specimens (two females): USA, Utah (G. F. Knowlton). The specimens were compared to the type material for a definite identification. All specimens (preserved in 70 % solution of ethanol with a drop of glycerol) are deposited in the collection of the University of Tyumen, Museum of Zoology, Tyumen, Russia. Supplementary description of adult. Measurements. Body length: 765, 780; notogastral width: 570, 600. Integument (Figs 1 A – C). Body color dark brown. Body and legs partially covered by thick gel-like layer of cerotegument, including dense microtubercles, sparsely located on notogaster and in marginal part of ventral plate. Surface with dense microtuberculate sculpturing; lamella, humeral process, pedotecta I, II, podosomal region, discidium, lateral part of prodorsum, lateral part of epimere I partially foveate; medioposterior part of notogaster (between setae lm, lp, h 2 h 3) with sparse, indistinct foveolae; lateral side of body between humeral process and acetabula II, III with dense large tubercles; additionally, podosomal region and dorsal part of pedotectum I partially rugose. Prodorsum (Figs 1 A – C). Rostrum pointed, observable in dorsal aspect. Rostral region with bulge. Outer tooth of lamellar cusp well developed. Translamella absent but medial parts of lamellae slightly protruding anteromedially, creating an illusion of the presence of a thin translamella. Tutorium with numerous teeth and small scales dorsally. Rostral seta (90 – 94) setiform, with short, flexible tip, barbed; lamellar seta (105 – 109) with thickened mediobasal part and flagellate distal part, barbed; interlamellar seta (217 – 225) thickened, acuminate, distinctly barbed; bothridial seta (124 – 131) with longer stalk and short expanded head bearing numerous strong spines; exobothridial seta represented by alveolus (may be broken). Notogaster (Figs 1 A – D). Centrodorsal region bordered by U-shaped thickening bearing dorsal notogastral setae. Setae h 1 (60 – 75) and p 1 (64 – 75) thickened, acuminate, heavily barbed; other setae (c, la, lm: 157 – 169; lp, h 2, h 3: 201 – 225; p 2, p 3: 134 – 146) thickened, acuminate, distinctly barbed. Opisthonotal gland opening and all lyrifissures observable. Gnathosoma (Figs 2 A – C). Subcapitulum size: 165 – 172 × 116; subcapitular setae (a: 34; m: 15 – 19; h: 45 – 52) setiform, roughened; m thinnest; both adoral setae (15 – 19) setiform, smooth. Palp length: 101 – 109; postpalpal seta (11 – 13) spiniform, truncate, roughened. Chelicera length: 172 – 184; setae (cha: 43 – 45; chb: 28 – 30) setiform, barbed. Epimeral and podosomal regions (Figs 1 B, C). All epimeral setae (3 c, 4 c: 67 – 75; 1 b: 45 – 52; 3 b, 4 a, 4 b: 34 – 37; 1 a, 1 c, 2 a, 3 a: 22) setiform, roughened. Discidium large, elongate triangular, rounded distally. Anogenital region (Figs 1 B – D). Genital (g 1, g 2: 34 – 37; g 3: 26 – 30; others: 19 – 22) and aggenital (26 – 30) setae setiform, roughened; both anal setae (22 – 26) setiform, barbed; all adanal setae (37 – 41) slightly thickened, acuminate, comparatively heavily barbed. Adanal lyrifissure distinct, oblique. Legs (Figs 2 D – G). All tarsi with three claws; median claw thicker than lateral claws; all claws slightly barbed on dorsal side. Trochanters III and IV with long tooth dorsoanteriorly. Formulas of leg setation and solenidia: I (1 - 5 - 3 - 4 - 20) [1 - 2 - 2], II (1 - 4 - 3 - 4 - 16) [1 - 1 - 2], III (2 - 3 - 2 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 3 - 3 - 12) [0 - 1 - 0]; homology of setae and solenidia indicated in Table 1; comparison of morphology of some setae between T. tridactylus, T. luissantossubiasi and T. mitratus presented in Table 2. Note: Roman letters refer to normal setae; Greek letters to solenidia (except ɛ = famulus); single quotation mark (’) marks setae on the anterior and double quotation mark (”) setae on the posterior side of a given leg segment; parentheses refer to a pair of setae.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB9D334FF27F8EE1236B41A.taxon	discussion	Remarks. Based on the original description (Woolley & Higgins, 1963) and the data presented here, the following diagnostic traits for T. tridactylus are proposed: Adult. Body size: 720 – 780. Medioposterior part of notogaster (between setae la, lp, h 2 h 3) with sparse, indistinct foveolae. Rostrum pointed. Lamellar cusps separated medially; translamella absent. Rostral seta setiform, with short, flexible tip, barbed; lamellar seta with thickened mediobasal part and flagellate distal part, with slight and sparse barbs; interlamellar seta thickened, acuminate, distinctly barbed; in ˃ le ˃ ro; bothridial seta with shortly expanded head bearing numerous strong spines. Centrodorsal region of notogaster bordered by U-shaped thickening. Notogastral setae h 1 and p 1 comparatively short, slightly different in length, thickened, acuminate, heavily barbed; other setae long, thickened, acuminate, distinctly barbed; lp, h 2, h 3 ˃ c, la, lm. All leg tarsi heterotridactylous; trochanters III and IV with long tooth dorsoanteriorly; seta pv’ on Ta I, II barbed (without ventral sparse spines); l ” and v’ on Ti I, II thorn-like; l ” on Ge I, II setiform.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.taxon	description	(Figs 3, 4)	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.taxon	materials_examined	Type material. Holotype (female) and three paratypes (three females): India, 28 ° 36 ' 56 '' N, 95 ° 53 ' 21 '' E, Arunachal Pradesh State, Etalin, 700 m a. s. l., soil with litter in forest, 1 – 8. v. 2012 (L. Dembický and O. Šauša). Two paratypes (two females): India, 28 ° 19 ' 32 '' N 95 ° 57 ' 31 '' E, Arunachal Pradesh State, Hunli, 1300 m a. s. l., soil with litter in forest, 26. v. – 1. vi. 2012 (L. Dembický and O. Šauša). Two paratypes (two females): India, 27 ° 00 ' 48 '' N 92 ° 39 ' 08 '' E, Assam State, Bhalukpong, 150 m a. s. l., soil with litter in forest, 1 – 8. v. 2012 (L. Dembický and O. Šauša). The holotype is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, Germany; seven paratypes are deposited in the collection of the University of Tyumen, Museum of Zoology, Tyumen, Russia. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.taxon	diagnosis	Diagnosis. Adult. Body size: 690 – 825. Medioposterior part of notogaster (between setae lp, h 2 h 3) with sparse, indistinct foveolae. Rostrum pointed. Lamellar cusps separated medially; translamella absent. Rostral seta setiform, with short, flexible tip, barbed; lamellar seta with thickened mediobasal part and flagellate distal part, with slight and sparse barbs; interlamellar seta thickened, acuminate, with slight and sparse barbs; in ˃ le ˃ ro; bothridial seta with shortly expanded head bearing numerous strong spines. Centrodorsal region of notogaster bordered by Ushaped thickening. Notogastral setae h 1 and p 1 comparatively short (p 1 ˃ h 1), thickened, acuminate, heavily barbed; other setae long, thickened, acuminate, with slight and sparse barbs but with some strong barbs in distal part. All leg tarsi heterotridactylous; trochanters III and IV with long tooth dorsoanteriorly; seta pv’ on Ta I, II with ventral sparse spines; l ” on Ti I, II and Ge I, II thickened; v’ on Ti I, II setiform, with ventral sparse spines.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.taxon	description	Description of adult. Measurements. Body length: 780 (holotype), 690 – 825 (paratypes); notogastral width: 600 (holotype), 555 – 600 (paratypes). Integument (Figs 3 A – D). Body color dark brown. Body and legs partially covered by thick gel-like layer of cerotegument, including dense microtubercles. Surface with dense microtuberculate sculpturing, clusters of larger tubercles close and lateral to notogastral U-shaped thickening; lamella, humeral process, pedotecta I, II, podosomal region, discidium, lateral part of prodorsum, lateral part of epimere I partially foveate; medioposterior part of notogaster (between setae lp, h 2 h 3) with sparse, indistinct foveolae; lateral side of body between humeral process and acetabula II – IV with dense large tubercles; additionally, humeral process, podosomal region and dorsal part of pedotectum I partially rugose. Prodorsum (Figs 3 A – C). Rostrum pointed, observable in dorsal aspect. Rostral region with bulge. Outer tooth of lamellar cusp well developed. Translamella absent but medial parts of lamellae slightly protruding anteromedially, creating an illusion of the presence of a thin translamella. Tutorium with numerous teeth and small scales dorsally. Rostral seta (90 – 105) setiform, with short, flexible tip, barbed; lamellar seta (154 – 176) with thickened mediobasal part and flagellate distal part, with slight and sparse barbs; interlamellar seta (172 – 206) thickened, acuminate, with slight and sparse barbs; bothridial seta (90 – 112) with longer stalk and short expanded head bearing numerous strong spines; exobothridial seta represented by alveolus (may be broken). Notogaster (Figs 3 A – D). Centrodorsal region bordered by U-shaped thickening bearing dorsal notogastral setae. Setae h 1 (56 – 64) and p 1 (86 – 94) thickened, acuminate, heavily barbed; other setae (c, la, lm, lp, h 2, h 3: 146 – 161; p 2, p 3: 101 – 124) thickened, acuminate, with slight and sparse barbs but with some strong barbs in distal part. Opisthonotal gland opening and all lyrifissures observable. Gnathosoma (Figs 4 A – C). Subcapitulum size: 154 – 161 × 123 – 135; subcapitular setae (a: 34 – 41; m: 13 – 15; h: 30 – 41) setiform, roughened; m thinnest; both adoral setae (15 – 19) setiform, smooth. Palp length: 94 – 97; postpalpal seta (11) spiniform, truncate, roughened. Chelicera length: 161 – 155; setae (cha: 37 – 41; chb: 28 – 30) setiform, barbed. Epimeral and podosomal regions (Figs 3 B, C). All epimeral setae (3 c: 75; 4 c: 56 – 71; 3 b: 56 – 60; 1 b, 4 a, 4 b: 41 – 52; 1 c: 34 – 37; 1 a, 2 a, 3 a: 15 – 19) setiform, roughened. Discidium large, elongate triangular, rounded distally. Anogenital region (Figs 3 B – D). Genital (g 1, g 2: 37 – 45; g 3: 30 – 37; others: 22 – 30), aggenital (30 – 37), anal (19 – 26), and adanal (41 – 56) setae setiform, roughened. Adanal lyrifissure distinct, oblique. Legs (Figs 4 D – G). All tarsi with three claws; median claw thicker than lateral claws; all claws slightly barbed on dorsal side. Trochanters III and IV with long tooth dorsoanteriorly. Formulas of leg setation and solenidia: I (1 - 5 - 3 - 4 - 20) [1 - 2 - 2], II (1 - 4 - 3 - 4 - 16) [1 - 1 - 2], III (2 - 3 - 2 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 3 - 3 - 12) [0 - 1 - 0]; homology of setae and solenidia indicated in Table 1; comparison of morphology of some setae between T. luissantossubiasi, T. tridactylus and T. mitratus presented in Table 2.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.taxon	discussion	Remarks. Tritegeus luissantossubiasi is morphologically most similar to T. tridactylus in main traits: rostrum triangular; bothridial seta with developed head; cusps of the lamellae separated medially; centrodorsal region of the notogaster bordered by U-shaped thickening, without rugose sculpturing; dorsal notogastral setae long (distinctly longer than distance lm – lp), thickened, acuminate; tridactylous legs. However, the new species differs from the latter by shorter dorsal notogastral setae lp, h 2, h 3 (not longer than distance lp – h 2 versus longer), roughened, with sparse barbs in distal part (versus distinctly and entirely barbed); the length of the posterior notogastral setae (p 1 distinctly longer than h 1 versus slightly different in length), and the morphology of the seta v’ on the leg tibiae I, II (setiform, with ventral sparse spines versus thorn-like).	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFBDD33BFF27FDBA113BB53A.taxon	etymology	Etymology. This species is named after the late Prof. Dr. Luis. S. Subías, the famous taxonomist in oribatology.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB2D33EFF27FC5612A8B71E.taxon	description	(Figs 5, 6)	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB2D33EFF27FC5612A8B71E.taxon	materials_examined	Material. Three specimens (one male and two females): Japan, Shikoku, Ishizuchi Mountain National Park, Tsuchigoya, Mt. Kamegamori, 1650 m a. s. l., abies and moss litter, with fungi and moss 15. VIII. 1980 (S. Peck). All specimens (preserved in 70 % solution of ethanol with a drop of glycerol) are deposited in the collection of the University of Tyumen, Museum of Zoology, Tyumen, Russia. Supplementary description of adult. Measurements. Body length: 780 – 825; notogastral width: 550 – 600. Integument (Figs 5 A – C). Body color dark brown. Body and legs partially covered by thick gel-like layer of cerotegument, including dense microtubercles, larger near the U-shaped thickening on notogaster; lamella, epimere I, pedotecta I, II, discidium, lateral part of prodorsum partially foveate; lateral side of body between humeral process and acetabula II – IV with dense large tubercles; anogenital region with heavy ridges; marginal region of notogaster with some longitudinal ridges; additionally, humeral process, podosomal region and dorsal part of pedotectum I partially rugose. Prodorsum (Figs 5 A – C). Rostrum pointed, not observable in dorsal aspect. Rostral region with short transverse ridge. Lamellar cusps fused mediodistally on significant continuation (hence, translamella absent); outer tooth of lamellar cusp well developed. Tutorium with numerous teeth and small scales dorsally. Rostral seta (86 – 94) setiform, with short, flexible tip, barbed; lamellar seta (131 – 154) with thickened mediobasal part and flagellate distal part, with sparse barbs; interlamellar seta (176 – 195) thickened, acuminate, roughened; bothridial seta (90 – 94) with longer stalk and short expanded head bearing numerous strong spines; exobothridial seta (56 – 60) setiform, roughened. Notogaster (Figs 5 A – D). Centrodorsal region bordered by U-shaped thickening bearing dorsal notogastral setae. Setae h 1 (49 – 56) and p 1 (56 – 64) thickened, acuminate, heavily barbed; p 2, p 3 (34 – 41) slightly stiff, roughened; other setae (86 – 94) thickened, acuminate, roughened. Opisthonotal gland opening and all lyrifissures observable. Gnathosoma (Figs 6 A – C). Subcapitulum size: 146 – 157 × 105 – 112; subcapitular setae (a: 30 – 34; m: 19 – 22; h: 30 – 37) setiform, roughened; m thinnest; both adoral setae (17 – 19) setiform, smooth. Palp length: 82 – 86; postpalpal seta (11) spiniform, roughened. Chelicera length: 146 – 161; setae (cha: 34 – 37; chb: 26 – 30) setiform, barbed. Epimeral and podosomal regions (Figs 5 B, C). All epimeral setae (3 c, 4 c: 56 – 64; 1 b: 37 – 45; 1 c, 3 b, 4 a, 4 b: 30; 1 a, 2 a, 3 a: 11) setiform, roughened. Discidium large, elongate triangular, rounded distally. Anogenital region (Figs 5 B – D). Genital (g 1, g 2: 26 – 30; g 3: 22 – 26; others: 15 – 19), aggenital (22 – 26), anal (22 – 26), and adanal (ad 1: 30 – 34; others: 26 – 30) setae setiform, roughened. Adanal lyrifissure distinct, oblique. Legs (Figs 6 D – G). All tarsi with three claws; median claw thicker than lateral claws; all claws slightly barbed on dorsal side. Trochanters III and IV with long tooth dorsoanteriorly. Formulas of leg setation and solenidia: I (1 - 5 - 3 - 4 - 20) [1 - 2 - 2], II (1 - 4 - 3 - 4 - 16) [1 - 1 - 2], III (2 - 3 - 2 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 3 - 3 - 12) [0 - 1 - 0]; homology of setae and solenidia indicated in Table 1; comparison of morphology of some setae between T. mitratus, T. tridactylus and T. luissantossubiasi presented in Table 2.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
03A34D63FFB2D33EFF27FC5612A8B71E.taxon	discussion	Remarks. Based on the original description (Aoki 1967) and the data presented here, the following diagnostic traits for T. mitratus are proposed: Adult. Body size: 780 – 840. Notogaster foveolate. Rostrum pointed. Lamellar cusps fused mediodistally (rostrum not observable in dorsal aspect). Rostral seta setiform, with short, flexible tip, barbed; lamellar seta with thickened mediobasal part and flagellate distal part, with sparse barbs; interlamellar seta thickened, acuminate, roughened; in ˃ le ˃ ro; bothridial seta with shortly expanded head bearing numerous strong spines. Centrodorsal region of notogaster bordered by U-shaped thickening. Notogastral setae h 1 and p 1 comparatively short (p 1 slightly longer than h 1) thickened, acuminate, heavily barbed; p 2, p 3 shortest, slightly stiff, roughened; other setae medium-sized, thickened, acuminate, roughened. All leg tarsi heterotridactylous; trochanters III and IV with long tooth dorsoanteriorly; seta pv’ on Ta I, II with ventral sparse spines; l ” on Ti I, II setiform; v’ on Ti I, II setiform, with ventral sparse spines; l ” on Ge I, II setiform.	en	Ermilov, Sergey G. (2024): Taxonomic contribution to knowledge of the oribatid mite genus Tritegeus (Acari, Oribatida, Cepheusidae). Zootaxa 5556 (1): 37-50, DOI: 10.11646/zootaxa.5556.1.4, URL: https://doi.org/10.11646/zootaxa.5556.1.4
