taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03A04148B132FE010BC5FF75F82EF0E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882762/files/figure.png	https://doi.org/10.5281/zenodo.17882762	FIGURE 1. Trichodrilus benati sp. nov. A, anterior part of body. B, Clitellum, surface view. C, nephridium in 12/13. D, atrium. E, atrial duct and penis, detail. F, spermatheca. G, H, atrium. (A–E, photos from the holotype, Mairulegorreta Cave; F, from paratype; G, H, from Arandilla River source).	FIGURE 1. Trichodrilus benati sp. nov. A, anterior part of body. B, Clitellum, surface view. C, nephridium in 12/13. D, atrium. E, atrial duct and penis, detail. F, spermatheca. G, H, atrium. (A–E, photos from the holotype, Mairulegorreta Cave; F, from paratype; G, H, from Arandilla River source).	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B132FE010BC5FF75F82EF0E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882764/files/figure.png	https://doi.org/10.5281/zenodo.17882764	FIGURE 2. Trichodrilus benati sp. nov. Drawing of the male duct, from the holotype.	FIGURE 2. Trichodrilus benati sp. nov. Drawing of the male duct, from the holotype.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B132FE010BC5FF75F82EF0E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882766/files/figure.png	https://doi.org/10.5281/zenodo.17882766	FIGURE 3. Trichodrilus medius Hrabě, 1960, syntypes. A–E, atrium. F, spermatheca. A, transverse section (Hr1517-30 #9). B–F, sagittal histological sections (Hr 1517-9 #1).	FIGURE 3. Trichodrilus medius Hrabě, 1960, syntypes. A–E, atrium. F, spermatheca. A, transverse section (Hr1517-30 #9). B–F, sagittal histological sections (Hr 1517-9 #1).	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B136FE0D0BC5FCA0F969F1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882768/files/figure.png	https://doi.org/10.5281/zenodo.17882768	FIGURE 4. Trichodrilus tenuis Hrabě, 1960, from Spanish populations. A–D, atrium. A, Coventosa Cave. B, Ermita San Isidro, Huelva. C, Cullalvera Cave. D, Ojo Guareña Cave. E, Spermatheca from Velamazan Well. A, whole-mounted individual, B–E from dissected worms.All scales represent 50 µm. Some photos are formed by the juxtaposition of two photos, one of them with marked edges, since the atrial duct is usually in another plane than the atrial ampulla.	FIGURE 4. Trichodrilus tenuis Hrabě, 1960, from Spanish populations. A–D, atrium. A, Coventosa Cave. B, Ermita San Isidro, Huelva. C, Cullalvera Cave. D, Ojo Guareña Cave. E, Spermatheca from Velamazan Well. A, whole-mounted individual, B–E from dissected worms.All scales represent 50 µm. Some photos are formed by the juxtaposition of two photos, one of them with marked edges, since the atrial duct is usually in another plane than the atrial ampulla.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B136FE0D0BC5FCA0F969F1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882770/files/figure.png	https://doi.org/10.5281/zenodo.17882770	FIGURE 5. Trichodrilus tenuis Hrabě,1960, syntypes. A, B, D, atrium in Hr 1517-7 (#1). C, atrium and penis Hr1772 (#19). E: Hr 1517-9 (#4) atrium and penis. F, spermatheca Hr 1517-7 (#1). All sagittal histological sections, except for E, from a wholemount. All scales 50 µm.	FIGURE 5. Trichodrilus tenuis Hrabě,1960, syntypes. A, B, D, atrium in Hr 1517-7 (#1). C, atrium and penis Hr1772 (#19). E: Hr 1517-9 (#4) atrium and penis. F, spermatheca Hr 1517-7 (#1). All sagittal histological sections, except for E, from a wholemount. All scales 50 µm.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B136FE0D0BC5FCA0F969F1B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882772/files/figure.png	https://doi.org/10.5281/zenodo.17882772	FIGURE 6. Distribution map of Trichodrilus tenuis (in red) and T. moravicus (in green) (from Timm & Abarenkov), and Iberian T. tenuis from the present study (in black). Karst areas of Spain (Oliveira et al. 2009) in the upper left square, marked in black.	FIGURE 6. Distribution map of Trichodrilus tenuis (in red) and T. moravicus (in green) (from Timm & Abarenkov), and Iberian T. tenuis from the present study (in black). Karst areas of Spain (Oliveira et al. 2009) in the upper left square, marked in black.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B13AFE0B0BC5FC14FE1FF68A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882774/files/figure.png	https://doi.org/10.5281/zenodo.17882774	FIGURE 7. Trichodrilus stygodytes sp. nov. A, schematic drawing of the reproductive system in segments X and XI. B, anterior body region. C, atrium and vasa deferentia junctions to the basal side of the atrial ampulla. D, atrium. E, atrial duct and penis. F, spermathecal duct with vestibule. (A–C from paratypes, E from holotype, D from an individual in the first author’s collection).	FIGURE 7. Trichodrilus stygodytes sp. nov. A, schematic drawing of the reproductive system in segments X and XI. B, anterior body region. C, atrium and vasa deferentia junctions to the basal side of the atrial ampulla. D, atrium. E, atrial duct and penis. F, spermathecal duct with vestibule. (A–C from paratypes, E from holotype, D from an individual in the first author’s collection).	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B13AFE0B0BC5FC14FE1FF68A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882778/files/figure.png	https://doi.org/10.5281/zenodo.17882778	FIGURE 8. T. moravicus Hrabě, 1937 and T. stygodytes sp. nov. A–C, T. moravicus, Říčka River type collection: A, atrium in transverse histological section, paralectotype (Hr1015-2 #17). B, atrium and posterior vas deferens junction in longitudinal section, lectotype (Hr1007-2(IV)#11). C, spermatheca in transverse sections (Hr1015-2 #17). D–H, T. stygodytes sp. nov., Býčí Skála collection: D–G, atrium. H, spermatheca.All scales 50 µm.	FIGURE 8. T. moravicus Hrabě, 1937 and T. stygodytes sp. nov. A–C, T. moravicus, Říčka River type collection: A, atrium in transverse histological section, paralectotype (Hr1015-2 #17). B, atrium and posterior vas deferens junction in longitudinal section, lectotype (Hr1007-2(IV)#11). C, spermatheca in transverse sections (Hr1015-2 #17). D–H, T. stygodytes sp. nov., Býčí Skála collection: D–G, atrium. H, spermatheca.All scales 50 µm.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B13CFE160BC5FB01F9C9F5B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882778/files/figure.png	https://doi.org/10.5281/zenodo.17882778	FIGURE 8. T. moravicus Hrabě, 1937 and T. stygodytes sp. nov. A–C, T. moravicus, Říčka River type collection: A, atrium in transverse histological section, paralectotype (Hr1015-2 #17). B, atrium and posterior vas deferens junction in longitudinal section, lectotype (Hr1007-2(IV)#11). C, spermatheca in transverse sections (Hr1015-2 #17). D–H, T. stygodytes sp. nov., Býčí Skála collection: D–G, atrium. H, spermatheca.All scales 50 µm.	FIGURE 8. T. moravicus Hrabě, 1937 and T. stygodytes sp. nov. A–C, T. moravicus, Říčka River type collection: A, atrium in transverse histological section, paralectotype (Hr1015-2 #17). B, atrium and posterior vas deferens junction in longitudinal section, lectotype (Hr1007-2(IV)#11). C, spermatheca in transverse sections (Hr1015-2 #17). D–H, T. stygodytes sp. nov., Býčí Skála collection: D–G, atrium. H, spermatheca.All scales 50 µm.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
03A04148B13CFE160BC5FB01F9C9F5B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17882772/files/figure.png	https://doi.org/10.5281/zenodo.17882772	FIGURE 6. Distribution map of Trichodrilus tenuis (in red) and T. moravicus (in green) (from Timm & Abarenkov), and Iberian T. tenuis from the present study (in black). Karst areas of Spain (Oliveira et al. 2009) in the upper left square, marked in black.	FIGURE 6. Distribution map of Trichodrilus tenuis (in red) and T. moravicus (in green) (from Timm & Abarenkov), and Iberian T. tenuis from the present study (in black). Karst areas of Spain (Oliveira et al. 2009) in the upper left square, marked in black.	2025-10-16	Rodríguez, Pilar;Fend, Steven		Zenodo	biologists	Rodríguez, Pilar;Fend, Steven			
