taxonID	type	description	language	source
03AE87B79C13FF8F7DC1FC6528ADF825.taxon	type_taxon	Type species. Concavapsocus parallelus Wang et al., 2019. Included species. Concavapsocus parallelus Wang et al., 2019; Concavapsocus valvaculeiformus sp. nov.	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C13FF8F7DC1FC6528ADF825.taxon	diagnosis	Revised diagnosis. Ocelli absent; antennae 23 - segmented; mx 2 and mx 4 without sensilla; forewing subrectangular (length: width = 3: 1), apical margin distinctly concave; vannal region well-developed; veins reduced, nearly parallel, with sparse setae along veins and posterior margin; R 2 + 3 fused with R 1 at middle of wing, distally bifurcating into R 2 and R 3; CuP and A 1 meeting at the middle of wing. Hind wing membranous, posterior margin with setae. Legs with thick setae, one coxal rasp present; tibiae with two apical spurs; tarsi three-segmented.	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C13FF8F7DC1FC6528ADF825.taxon	discussion	Remarks. Concavapsocus Wang et al., 2019 is distinguished from the other Psyllipsocidae genera by its unique forewing morphology and venation. This genus exhibits a consistent forewing structure regardless of wing type: macropterous or brachypterous forms both display an inwardly concave apical margin, a well-developed vannal region, and a subrectangular outline length-to-width ratio of 3: 1. In contrast, others Psyllipsocidae genera exhibit brachypterous species with rounded apices, while macropterous forms bear elongate, strip-like wings (e. g., Psyllipsocus Selys-Longchamps, 1872). In Concavapsocus, veins are reduced and nearly parallel; pterostigma, cell, and nodulus are absent, and M, CuA and A veins are unbranched, these features differ significantly from other genera: Psyllipsocus with two radial cells are present: second radial cell is either pentagonal or hexagonal in shape, and CuA branched; Khatangia with a triangular pterostigma and closed pentagonal radial cell bordered by straight R 1; in Libanopsyllipsocus, M vein bifurcated, areola postica elongate and free, CuP and A 1 converge to form a nodulus; In Globopsocus, Rs and Cu are unbranched, M vein branched. In addition, antennae, maxillary palp and hindwing can also distinguished Concavapsocus: antennae 23 - segmented, with cuticular sculpture (vs. part of Psyllipsocus antennae shorter than 20 - segmented); mx 2 and mx 4 lack sensillum (vs. Psyllipsocus mx 4 without large sensillum, or with smaller ones near distal end); hindwing R and M unbranched, CuA bifurcating into CuA 1 and CuA 2 (vs. Psyllipsocus and Libanopsyllipsocus hindwing with a basal cell and branched Rs and M veins); tibiae with two apical spurs, tarsi three-segmented, preapical claws without preapical tooth (vs. Khatangia, Libanopsyllipsocus and some Psyllipsocus preapical claws with preapical tooth) (Vishnyakova 1975; Mockford 1993; Perrichot et al. 2003; Azar & Engel 2008; Azar & Nel 2011; Zhang et al. 2016; Hakim et al. 2018).	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C10FF8A7DC1FF5D2A76FA6F.taxon	description	(Figs 1 – 2)	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C10FF8A7DC1FF5D2A76FA6F.taxon	materials_examined	Materials. All specimens are adult. Holotype: CNU-PSO-MA 2015005, female, well-preserved. Paratype: CNU-PSO-MA 2015006, female, well-preserved. CNU-PSO-MA 2015007, male, body covered by many impurities, head and legs broken; CNU-PSO-MA 2015008, male, terminalia blurred and covered by wings; CNU-PSO-MA 2015009, female, wings broken. Locality and horizon. Hukawng Valley, Kachin State, Northern Myanmar. mid-Cretaceous, lowermost Cenomanian.	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C10FF8A7DC1FF5D2A76FA6F.taxon	etymology	Etymology. The species name is a combination of the Latin words “ valv- ” (valve) and “ aculeiformus ” (needle like), referencing the elongated needle-like structure of the ventral valves.	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C10FF8A7DC1FF5D2A76FA6F.taxon	diagnosis	Diagnosis. Distinct ecdysial line, lacinia developed with four apical teeth. Macropterous or brachypterous. Legs covered dense setae, tibiae with two apical spurs. Paraproct lacking anal spine; V 1 slender, base unfused and tip sharp, needle-like apex, approximately three times the length of V 2; V 2 rod-like, twice as thick as V 1; V 3 short, broad and rounded, flanking ventral valve laterally; subgenital plate sparsely setose.	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C10FF8A7DC1FF5D2A76FA6F.taxon	discussion	Remarks. We assigned the new species to Concavapsocus Wang et al., 2019 based on the following characteristics: (1) ocelli absent; (2) antennae with 23 - segmented; (3) mx 2 and mx 4 without conical sensillum; (4) lacinia developed; (5) forewings membranous, with several setae on veins and posterior margin; (6) basal portion of Sc short, ending free in membrane; (7) basal part of R 1 fused with R 2 + 3, A 1 fused with CuP near middle of forewing; (8) hind wing present, membranous; (9) coxae with several setae; one coxal rasp present; (10) tibiae with two apical spurs; tarsi with three-segmented. Therefore, the new specimens can be clearly classified into Concavapsocus. The differences between Concavapsocus valvaculeiformus sp. nov. and C. parallelus include: (1) C. valvaculeiformus with a prominent vertexal ecdysial cleavage line (vs. C. parallelus ecdysial cleavage line absent); (2) lacinia developed, bearing four teeth of nearly uniform size (vs. C. parallelus lacinia only with two apical teeth); (3) macropterous and brachypterous (vs. C. parallelus only has brachypterous); (4) paraproct without an anal spine (vs. C. parallelus paraproct with an anal spine); (5) V 1 slender, the base unfused and the tip sharp, fine needle-like, approximately third times the length of V 2, V 2 rod-like, twice times thicker than V 1, V 3 short, broad and rounded, located on both sides of ventral valve, subgenital plate with few setae (vs. C. parallelus V 1 reduced to rod-like, not sclerotized, V 2 slender, needle-like, V 3 broad without posterior setae, subgenital plate with dense setae, covering at most part of basal V 3).	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
03AE87B79C10FF8A7DC1FF5D2A76FA6F.taxon	description	Description. Head: Oval, width 0.2 mm, length 0.36 mm; dostinct vertexal ecdysial line, covered by long dense setae. Compound eyes well-developed, located on both ends of head, diameter 0.11 mm, IO / d = 0.30, ocelli absent. Postclypeus broad, diameter longer than half of compound eyes. Antennae 23 - segmented, with secondary annulation, ca. 1.12 mm long, scape and pedicel ca. 0.08 mm long, broader than flagellomeres; scape, pedicel, f 1 and f 2 with setae, f 1: 0.10 mm, f 2: 0.09 mm (Fig. 2 A). Chewing mouthparts, maxillary palps four-segmented, ca. 0.29 mm long, mx 2 and mx 4 longest, mx 1: 0.02 mm, mx 2: 0.10 mm, mx 3: 0.05 mm, mx 4: 0.12 mm, mx 4 rod-like, mx 2 and mx 3 with setae. Lacinia developed, with four apical teeth (left lacinia of CNU-PSO-MA 2015006 with only three teeth visible); labial palps two-segmented, terminal segment rounded, thicker than maxillary palp and lacinia, ca. 0.08 mm long (Fig. 2 B). Thorax: Neck relatively wider, with a transversely band-like pronotum exhibiting convex on the sides and concave in the middle. Wings: Macropterous, forewing 1.37 mm in length, 0.47 mm in width, with a length-to-width ratio of three to one; forewing vannal fold developed, subrectangular and apex concave inwardly, veins parallel, bearing setae. forewing Sc short, not reaching R or wing margin, ends in membrane at 0.18 mm, pterostigma absent; R with four branches, bifurcating into Rs and R 4 + 5 at 0.35 mm distal of wing base, R 4 + 5 reaching wing margin at 1.35 mm, Rs bifurcating into R 1 and R 2 + 3 at 0.58 mm distal of wing base, R 2 + 3 bifurcating into R 2 and R 3 at 1.01 mm distal of wing base, reaching wing margin at 1.18 mm and 1.32 mm respectively; R fused with M basally, then bifurcate at 0.18 mm; base of CuA interrupted, M and CuA unbranched, reaching wing margin at 1.33 mm and 1.30 mm respectively; A unbranched, reaching wing margin at 0.88 mm; CuP and A 1 meeting at middle of wing at 0.38 mm at distal of wing base; posterior margin bearing setae. Hind wing present, 1.30 mm in length, 0.42 mm in width, membranous; Sc absent, R and M unbranched, reaching wing margin at 1.29 mm and 1.23 mm respectively, Cu bifurcating into CuA and CuP at 0.52 mm distal of wing base, A absent; radial cell absent (Fig. 2 C). Leg: Coxae setose, hind legs with coxal rasp (Fig. 2 D), tibiae bearing dense setae with two apical spurs; tarsi three-segmented (t 1> t 2> t 3), t 1 and t 2 with spurs. Preapical claws simple, symmetrical without preapical tooth and pulvilli (Fig. 2 E). Female genitalia: Paraproct without anal spine. V 1 slender, base unfused apex needle-like, approximately third times the length of V 2; V 2 rod-like, and twice times thicker than V 1; V 3 short, broad and rounded, flanking valve laterally, subgenital plate with sparse setae (Fig. 2 F, G).	en	Yang, Xiao-Yue, Li, Sheng, Ren, Dong, Yao, Yun-Zhi (2025): New species of Concavapsocus (Psocodea: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar. Zootaxa 5637 (3): 594-600, DOI: 10.11646/zootaxa.5637.3.7, URL: https://doi.org/10.11646/zootaxa.5637.3.7
