taxonID	type	description	language	source
03A887C7FFDA39213FB2FF2C1BA85849.taxon	type_taxon	Type species. Atracodes indutus Melichar, 1902 (by subsequent designation of Melichar 1923: 118).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFDA39213FB2FF2C1BA85849.taxon	description	Amended description. Head and eyes much narrower than pronotum at caudal margin. Vertex in dorsal view longer than broad bearing median carina, lateral carinae weakly foliate, subparallel proximally, curved medially near head apex. Head apex in dorsal view roundly produced, in lateral view projected for length greater than maximum width of eye (subequal in length to prothorax at midline). Frons elongated, much longer at midline than wide at widest point (near lower margin of eyes), narrowed dorsally, apex with callosity. Frontoclypeal suture nearly linear (weakly sinuate). Eye round in lateral view (or nearly so), lateral ocelli below eye (just in front of eye midline and above level of antennae). Antennae short, scape cylindrical in cross-section, about as tall as wide, pedicle distally enlarged, taller than wide; flagellum bristle-like with bulbous base. Pronotum roughly trapezoidal from dorsal view (with anterior apex truncate), about as long at midline as vertex, broadest posteriorly, strongly narrowing cephalad to less than width of head with eyes; anterior margin strongly arched (may be medially truncate or notched), posterior margin broadly concave, disc tricarinate (may be obscured by wax), carinae obsolete anteriorly; lateral margins of dorsal disc carinate, disc bearing pair of variably distinct depressions (one on either side of midline); paradiscal region broad, extending ventrad below level of antennae. Mesonotum broad and about as long at midline as vertex and pronotum combined, tricarinate (may be obscured by wax). Forewings broad, held broadly tent-like (a subfamily feature), costal margin undulating and expanded to form very broad postcostal cell, humeral region bearing lobe projecting anteriad of tegulae; a small bulla bearing a cluster of pustules in proximal portion of Sc + RA; nodal line just distad of midlength (forewing often slightly inflected or curved downward near level of nodus); distal portion of forewing more reticulate than proximal; forewing apex with two irregular subapical lines (defining two irregular rows of cells, proximal band poorly defined in some species); in lateral view, forewings often inflected ventrally near nodal line. The number of lateral spines on the hind tibia differ between species, and range from 2 – 4 spines. The apical spinulation differs between species and ranges from 6 – 8 spines on the first and second tarsal segments, with the third segment always having 2 spines. Pygofer of male terminalia lacking lateral or medioventral projections. Gonostyli in lateral view elongate and bearing prominent capitulum (simple and elongate hooked). Aedeagal complex (Fig. 6; aedeagus + phallotheca) bilaterally symmetrical, stout, apically bifurcate and ornamented with varied processes; approximately subequal in length to gonostyli in lateral view, shaft straight to weakly upcurved, bearing varied apical and lateral processes; aedeagus simple, usually distinguishable medioventrally from phallotheca. Anal tube (lateral view) narrow and elongated, usually exceeding gonostyli, distally down curved.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFDA39213FB2FF2C1BA85849.taxon	biology_ecology	Plant associations. Atracodes indutus: Coccoloba uvifera (L.) L. (seagrape, Polygonaceae; label information); Atracodes albus comb. nov.: Inga vera Willd. (Fabaceae), Lantana sp. (Verbenaceae; label information).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFDA39213FB2FF2C1BA85849.taxon	discussion	Remarks. The American Flatoidinae comprises seven genera (excluding species misplaced in the Old World genus Flatoides) and about 70 species. Three genera are reported only from the continental mainland (Flataloides Metcalf, Metcracis Medler, and Phalaenomorpha Amyot & Serville), one is widespread in both the mainland and the Caribbean (Flatoidinus Melichar), and three are reported only in the Caribbean region (Atracodes, Flatarina Metcalf & Bruner, and Flatarissa Metcalf & Bruner). Of the Caribbean genera, two (Flatarissa and Flatarina) are Cuban endemics. Keys that include American Flatoidinae can be found in Melichar (1901), Metcalf (1938), Metcalf & Bruner (1948), Caldwell & Martorell (1951), and Bartlett et al. (2014), although none of these include all American genera. Among the genera of Flatoidinae in the Caribbean region, Atracodes can be most readily diagnosed by the projected head (vertex much longer than broad, or in lateral view, head projected in front of eye for a distance equal or greater than the greatest width of the eye), and the presence of an undulating costal margin of the forewing. The Cuban endemic Flatarissa has an undulating costal margin, but the vertex is broader than wide. The Cuban endemic Flatarina does not have either feature. The genus Flatoidinus is widespread and diverse. Flatoidinus bears a pointed head, but the head is generally broader than long (or as broad as long), and the costal margin of the forewings is not undulate. We have found the number of lateral spines on the hind tibiae to be an unreliable feature. Within Atracodes, the number of lateral spines on the hind tibiae varies from 2 – 4. Atracodes was originally described by Melichar (1902) as having three lateral spines and subsequently as having two by Melichar (1923). We have found that Atracodes indutus Melichar, 1902 often has four lateral spines, with one very small. Similarly, the waxy pruinosity that covers the wings and body of many flatids may bear helpful diagnostic patterns, but at the same time obscure coloration and surface morphological features (such as carinae and wing venation). Coloration may vary geographically and possibly by the age of the individual resulting in considerable variation within a species. From a diagnostic standpoint, the pruinosity can be problematic, and removing it (perhaps by brushing with ammonia) may be desirable to review obscured features. Within Atracodes, there appear to be two species groups based on the male terminalia, the tortrix- group, and the indutus- group. The tortrix- group contains A. tortrix, comb. nov., A. fasciculosus, comb. nov., A. fasciatus comb. nov. & stat. nov., A. insularis comb. nov., A. vittatus comb. nov. & stat. nov. and A. lichenoides comb. nov. This group is characterized by: anal tube (dorsal view) broad and of uniform width, aedeagal complex (lateral view) straight or downcurved, phallotheca (dorsal or ventral view) deeply bifurcated (to about half length), bearing 2 – 4 processes (Fig. 11), with aedeagus extending from the dorsal side of the phallotheca, between the bifurcations; tapered to a convex apex. The indutus- group contains A. indutus, A. longifrons, A. sanctaeluciae, and A. albus comb. nov. This group is characterized by: anal tube (dorsal view) narrowed in proximal half (broader distally); aedeagal complex curved upward distally, phallotheca (dorsal or ventral view) less deeply bifurcated (to about three quarters length), bearing three pairs of processes (Fig. 6), with aedeagus extending ventrodorsally, apex upcurved.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD9392C3FB2F9291FE05D62.taxon	description	(Figs 2 – 6)	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD9392C3FB2F9291FE05D62.taxon	materials_examined	Type locality. St. Thomas, US Virgin Islands. Amended description. Color. General color from mottled brown, to greige, to ivory, with an irregular waxy pruinescence (which obscures surface color patterns; Fig. 4). Head in lateral view (Fig. 2 B, 3 B) with a dark line extending anteriorly from eye to dark apical callosity. A similar line pronotum from posterior margin of eye to tegulae (may be irregular or obscured by wax). Mesonotum anteriorly embrowned, bearing three darkened lines tracing carinae to apical margin and bearing pair of dark spots near anterior margin, a second pair near lateral margin behind midline, and a third pair on either side of midline near scutellum. Holotype (Fig. 2) forewing bearing dark markings on tegulae and angled lines on forewing, obscured or wanting on other specimens (Fig. 4); holotype forewing with irregular dark spots in clavus; a curved dark line from A 1 to CuP; a dark spot is located at 1 ⁄ 4 the length of the CuP from the basal cell, with a lighter patch behind it, where the CuA splits; a lateral runs down from the CuP to MP, and runs along MP; dark patch located in the middle of the cells between Pc + CP and ScP; dark line running from MP to MP 1. Structure. Body length (with wing) male: 7.0 – 13.0 mm; female: 9.0 – 15.0 mm. Head (in dorsal view, Figs 2 A, 3 A) narrow, distinctly projecting in front of eyes for distance greater than eye width; apex bluntly pointed bearing callosity. Vertex roughly quadrate, length and width subequal; median carina present, appearing apically forked into strongly serpentine carinae directed laterad to meet lateral carinae of vertex, lateral margins subparallel, slightly foliate, linear for most of length, subapically appearing medially inflected into strongly serpentine carinae reaching median carina; in lateral view, lateral carinae of vertex meeting lateral carinae of frons at sharp angle, ~ 30 degrees); posterior margin appearing concave, hidden by anterior margin of pronotum. Frons nearly quadrate, from ventrofrontal view (e. g., Figs 2 C, 3 D), broadest near level of antenna, narrowed distally and proximally; apex bluntly rounded at callosity, lateral carinae foliate, median carina absent; frontoclypeal suture truncate. Clypeus triangular, bearing median carina. Rostrum reaching hind trochanters. Hind tibiae (Fig. 3 C) with 4 lateral teeth (proximal tooth small); apical spinulation 6 (2 + 4) - 7 - 2. Pronotum approximately triangular from dorsal view, anterior margin convexly rounded and carinate (carina extending diagonally laterad to form lateral margin of pronotal disc), posterior margin broadly and narrowly concave; length subequal to vertex at midline; faintly tricarinate, a pair pits on either side of midline in anterior half of disc; in lateral view pronotum near same level as vertex, Fig. 3 B); paradiscal region broad, approximately quadrate, bearing transverse carinae between eye and tegulae. Mesonotum broad, from dorsal view roughly lozenge-shaped (from lateral view raised above level of pronotum), obscurely tricarinate. Forewings broad, leading margin expanded into board postcostal cell bearing many crossveins (mostly simple, except forked near wing apex), lateral margin undulate, humeral angle expanded into anteriorly projecting lobe; distal portion of wing inflected downward at nodal line; nodal line and subapical marginal vein arched and irregular, forming 2 rows of elongate apical cells; a proximal bulla (bearing cluster of pustules) on proximal portion of Sc + R; clavus exceeding midlength of wing, A 1 close to marginal A 2; CuA forked well before wing midlength; MP twice forked before wing midlength; RP-MP, Sc + RA, and PC + CP, run to marginal vein without branching. Pygofer from lateral view (Fig. 2 D) broad and irregularly shaped, broadest subventrally, caudal margin convexly curved; anterior margin sinuately convex; medioventral process absent (ventral view), margin distinctly concave (similar to Fig. 10 D). Gonostyli in lateral view broad, exerted portion roughly quadrate bearing large elongate, falcate capitulum (on subapical dorsal margin); apex concave (with indentation near ventral margin); in ventral view gonostyli diverging (defining median triangle, similar to Fig. 10 D); narrowed distally, inner margins linear, lateral margin weakly sinuate. Phallotheca bilaterally symmetrical; apex (in dorsal or ventral view) appearing wide and broadly bilobed (from dorsal, Fig. 6 B, or ventral view, Fig. 6 C); upturned apically (from lateral view) (Fig. 6 A) bearing subapical dorsal spine just before upward inflection; phallotheca bearing four pairs of retrorse processes; three pairs extending from phallotheca distal apex, one stout pair reading phallotheca midlength and two shorter pair (one pair slender, one pair flattened and bladelike); last pair originating from subapical ventral margin, slender and elongated, distally upcurved exceeding phallotheca midlength, apex usually resting above dorsal margin of phallotheca. Aedeagus appearing medioventral (visible from ventral view, Fig. 6 C), apex attenuated to a point and upcurved between lateral lobes of phallotheca. Anal tube (lateral view, Fig. 2 D) hood-like (distal half downturned), relatively slender, ventral margin concave, dorsal margin convex; ental surface bearing small ventral projection in distal third (Fig. 2 D); paraproct short cylindrical, epiproct rounded.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD9392C3FB2F9291FE05D62.taxon	discussion	Remarks: Atracodes indutus is most similar to Atracodes albus comb. nov. but can be distinguished by size (A. indutus is about two thirds the size of A. albus comb. nov.), A. indutus has dark spots in each of the marginal cells, at the costal margin in the forewing, whereas A. albus comb. nov. has dark spots in only some of the marginal cells. The anal tube in A. albus comb. nov. (Caldwell & Martorell 1951, plate 48 a) from lateral view has a large ventral lobe on the lateral margin just before midlength (absent in A. indutus). The gonostyli in A. indutus extend further past the carinae, than in A. albus comb. nov. Atracodes albus and A. indutus can be most easily differentiated from the other eight species in the Atracodes, by the dark lateral lines distad and proximad to the eyes. Atracodes indutus was briefly described by Melichar (1902) with a dorsal habitus illustration (viz. plate 8, fig. 7). Specimens from Guana Island, and Virgin Gorda Island, do not closely fit the patterning (viz. The dark markings on the forewings), or coloration of the original description. Specimens range from brown to greige, to ivory, with irregular and often obscure patterning. Plant associations. Guana Island specimens were collected at U. V. lights near Coccoloba uvifera (L.) L. (seagrape, Polygonaceae).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD9392C3FB2F9291FE05D62.taxon	distribution	Distribution. US Virgin Islands (St. Thomas), British Virgin Islands (Frenchman’s Cay, Guana, Virgin Gorda).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD9392C3FB2F9291FE05D62.taxon	materials_examined	Type material examined. Holotype, male: St. Thomas / Coll. Signoret // indutus / det. Melichar // Atracodes / indutus // Coll. Nat. - Mus. Wien // Holotype / Atracodes / indutus / det. ♂ Melichar — NHMW. Other material examined. BRITISH VIRGIN ISL. / Guana Island, 0 – 80 m. / 13 – 26 July 1986 / S. E. Miller & M. G. Pogue // North Bay, Coccoloba / forest, U. V. light / trap, sea level / 15 – 25 July 1986 // USNM // Atracodes / indutus Melichar, 1902 / Det. NNLevia 2023 // UDCC _ TCN 00101345, 00101346, 00101362 – 00101369, 00101371, 00101373 (USNM, 12 Males). BRITISH VIRGIN ISL. / Guana Island, 0 – 80 m. / 13 – 26 July 1986 / S. E. Miller & M. G. Pouge // USNM // Atracodes / indutus Melichar, 1902 / Det. NNLevia 2023 // UDCC _ TCN 00101336 – 00101344, 00101355 – 00101361, 00101265, 00101266, 00101372 (USNM, 12 Male, 7 Female). BRITISH VIRGIN ISLANDS / Virgin Gorda Island / V. Gorda Peak, ca. 400 m / 17 – 19 July 1986 / S. E. Miller & M. G. Pogue // USNM // Atracodes / indutus Melichar, 1902 / Det. NNLevia 2023 // UDCC _ TCN 00101374 (USNM, male). BRITISH VIRGIN IS. / Guana Island / July 1986 / Scott E. Miller // Pseudoflatoides / albus Caldwell / Det: CRBartlett ‘ 99 // UDCC _ TCN 00103107 (NCSU, male). BRIT. VIRGIN ISLS. / Guana Island / 1 – 14 July 1984 / S. E. & P. M. Miller // Atracodes / indutus Melichar, 1902 / Det. NNLevia 2024 // UDCC _ TCN 00103111 – 00103115, 00102961 (USNM, 4 male, 1 female, 1 nymph).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD439293FB2FCDD1F625DA7.taxon	materials_examined	Type locality. San Vicente, Pinar del Rio Province, Cuba. Amended description. Color. General color chalky to yellowish white (Figs 8 A, 9 A), forewing veins darker, variably patterned. Head with dark patch behind eye (line anterior to eye absent; Figs 8 B, 9 B). Mesonotum variably embrowned, especially posteriorly (may be broken into three pairs of large black spots). Forewings bearing variably developed dark transverse serpentine band near midlength (at nodal line, not reaching clavus); apical marginal cells bearing small dark spots. Structure. Body length (including wings) male: 15.0 – 19.5 mm; female 19.2 – 21.1 mm. Head (in dorsal view, Figs 8 A, 9 A) narrow projecting in front of eyes for distance greater than eye width, bluntly pointed bearing apical callosity. Vertex quadrate, about 1.5 x as long as wide; medial carinae present, distally forked; lateral carinae keeled, serpentine, subapically inflected medially, then strongly serpentine to join median carina; posterior margin concave (partly hidden by anterior margin of the pronotum); in lateral view, lateral carinae of vertex meeting lateral carinae of frons at sharp angle, ~ 50 degrees). Frons from ventral view (Fig. 8 D), roughly elongate quadrate (length about 2 x greatest width); widest proximad of midlength (narrowed distally and proximally), median carinae obsolete, lateral carinae foliate. Frontoclypeal suture approximately linear. Clypeus elongate and triangular (median carina present, obscure). Rostrum reaching or exceeding hind trochanter. Hind tibiae (Fig. 8 C) bearing two lateral teeth, apical spinulation 8 (5 + 3) - 8 - 2. Pronotum (Figs 8 A, 9 A) roughly triangular in dorsal view, anterior margin keeled, convexly rounded, anterior keels projecting caudad diagonally to form lateral margins of disc; median carina obscure, disc bearing pair of pits on either side of midline just anterior to midlength; paradiscal region broad (extending lateral just below level of antennae), bearing transverse carina between eye and tegula. Mesonotum broad (about equal in length to pronotum and vertex combined at midlength), roughly lozenge-shaped with the lateral corners rounded, obscurely tricarinate; in lateral view mesonotum raised above the pronotum. Forewings with broad postcostal cell bearing many marginal veins often distally forked; claval apex well distad of forewing midlength, A 1 close to marginal A 2 vein, CuA vein forked near claval midlength at about level of first fork of MP. Pygofer (Figs 10 A, B) in lateral view irregular, widest subventrally, caudal margin convexly rounded, anterior margin concave; medioventral process absent, ventral margin concave (Fig. 10 D). Phallotheca stout, bilaterally symmetrical (Figs 10 A, 11 A), distally narrowed, bifurcate with medial aedeagus, weakly downcurved; bearing three pairs of subapical retrorse processes (A 1 – A 6), A 1 and A 2 arise dorsoapically (dorsal apical processes), long and slender, reaching phallathecal base; A 3 and A 4 (dorsal subapical processes) arise just proximad of A 1 and A 2. stout and blade-like reaching near phallathecal midlength; A 5 and A 6 arise on subapical ventral margin, shorter and stouter than A 1 and A 2 nearly reaching phallathecal base (Fig. 11 A). Aedeagus extends dorsomedially from phallotheca to about half the length of the bifurcation; broad, tapering to upcurved, convex apex. Gonostyli comparatively slender (Figs 10 A, B), exerted portion elongate quadrate (dorsal and ventral margins nearly straight) bearing large apical hooked capitulum on dorsal margin; apical margin weakly arched. Anal tube in lateral view (Figs 10 A, B) slender and elongated, dorsal margin sinuately curved, distally downcurved, apex sharply inflected downward; ventral margin sinuately convex; paraproct elongated caudally.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD439293FB2FCDD1F625DA7.taxon	discussion	Remarks. Metcalf and Bruner (1948: 109) described this species as a subspecies of Pseudoflatoides fasciculosus, differing from the nominate subspecies because it was ‘ heavily marked’, but ‘ the structure and details of genitalia agree with the typical variety’. The typical subspecies was illustrated (Metcalf and Bruner 1948, plate VIII, Figs 7 – 8, plate XVI, fig. 5), but P. f. fasciatus was not. The terminalia of the paratype from Camagűey differs markedly from that illustrated by Metcalf and Bruner (1948, plate XVI, fig. 6) in the pygofer is broader in A. fasciatus, the anal tube shape differs (most notably not inflected downward at apex, versus A. fasciatus, Figs 10 A, B), and the aedeagus processes (in the typical form, fasciculosus processes A 3 and A 4 are slender, not stout, and processes A 5 and A 6 are short and hooked instead of reaching the aedeagal base). These differences are sufficiently great to revise the status of the subspecies to a full species as Atracodes fasciatus, comb. nov. & stat. nov. Atracodes fasciatus comb. nov. & stat. nov. appears similar to A. tortrix comb. nov. (Fig. 7 D, Metcalf & Bruner 1948, plate XVI, fig. 4) in that in the latter species the aedeagus appears shorter, aedeagal processes A 3 and A 4 appear more elongated, and the apex of the anal tube is not inflected downward. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD439293FB2FCDD1F625DA7.taxon	distribution	Distribution. Cuba (Camagűey, Artemisa, Pinar del Río).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD439293FB2FCDD1F625DA7.taxon	materials_examined	Type material examined. HOLOTYPE: San Vincente / 4 - 6 - 9 - [19] 22 / cd. s. c. B. / g j. a // ♂ // Holotype // 8768 // Pseudoflatoides / fasciculosus / fasciatus / Metc. & Brun. / det. Z. P. M (NSCU, male). ALLOTYPE: San Vincente / En Go? m? [illegible] / 4 - 9 - [19] 22 // jc 34 J. A // 8768 // Allotype // Pseudoflatoides / fasciculosus / fasciatus / Metc. & Brun. / det. Z. P. M // UDCC _ TCN 00102976 [2 D barcode label] (NCSU, female). PARATYPES: Camagűey, Cuba, / J. Acuba, Col. / Aug. 20 1924 // Paratype // Pseudoflatoides / fasciculosus / fasciatus / Metc. & Brun. / det. Z. P. M // UDCC _ TCN 00102977 (NCSU, female). Cubitas // Camagűey, Cuba / July 30, 1923 / J. Acuna, Col. // Paratype // Pseudoflatoides / fasciculosus / fasciatus / Metc. & Brun. / det. Z. P. M // UDCC _ TCN 00103104 (NCSU, female). Sierra Rangel / P. del Río, Cuba / Aug. 29, 1927 / Alt. 1500 feet // Col. J. Acuña, y S. C. Bruner. // Paratype // Pseudoflatoides / fasciculosus / fasciatus / Metc. & Brun. / det. Z. P. M. // UDCC _ TCN 00103105 (NCSU, female).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD139363FB2FBA31D505A17.taxon	materials_examined	Type Locality. Camagűey, Camagűey Province, Cuba. Amended description. Color. Ochraceous buff (Figs 12 A, 13 A), tinged with olive green, heavily marked with brownish fuscous. Eyes with dark semicircular lines. Dark lateral line from posterior margin of eye to wing base (Fig. 12 B). Lateral margins of mesonotum largely bordered with dark lines that may be broken up into dots (Figs 12 A, 13 A). Forewings heavily covered in whitish pruinescence, patterning variable, usually bearing proximal fuscous markings on remigium; dark line tracing claval margin proximally then arched diagonally across remigium to reach leading wing margin subapically; transverse dark line following nodus, sometimes with broad, strongly serpentine, irregular fuscous marking from apex of clavus to wing apex. Structure. Body length (head apex to forewing tip) male: 15.2 mm; female: 16.8 – 18.0 mm. Head (dorsal view, Figs 12 A, 13 A) narrow, distinctly projecting in front of the eyes for distance greater than eye width, posterior margin hidden by anterior margin of pronotum. Vertex about twice as long as width between eyes; median carina present, forked at apex to form serpentine carinae contiguous with lateral carinae; lateral carinae nearly parallel posteriorly, distal portion medially serpentine, joining with median carina near head apex. Head apex blunt with medial callosity. Frons (Figs 12 D, 13 C) elongated, approximately narrowly quadrate, apex rounded; 3 x as long as broad, lateral margins arched, widest near level of antennae; median carina absent, lateral carinae foliate. Frontoclypeal suture linear (upcurved laterally), clypeus elongate triangular. Eyes circular (Figs 12 B, 13 B), lateral ocelli obscure below eye near midlength (anterior to and slightly above level of antennae). Rostrum reaching hind trochanters. Pronotum (Figs 12 A, 13 A) approximately triangular in dorsal view, with the apical margin rounded, anterior end convexly rounded, carinate (from lateral view lateral carinae of vertex meeting lateral carinae of frons at strong angle). Mesonotum broad, roughly lozenge-shaped with the lateral corners rounded, and raised above the pronotum, tricarinate. Hind tibiae with two lateral teeth (Fig. 12 C); apical spinulation 7 (3 + 2) - 7 - 2. Forewings broad, humeral angle forming a lobe that projects toward the cephalad tegulae, lateral margin undulate, postcostal cell broad, marginal crossveins mostly forked, clavus extending well past forewing midlength, A 1 close to marginal A 2 vein, MP forked proximad to fork of CuA. Pygofer in lateral view (Figs 14 A, B) broad, irregularly shaped, with extended area dorsally on apical side. Phallotheca bilaterally symmetrical (Fig. 15), in lateral view (Fig. 15 A) narrowing distally past midlength; in ventral and dorsal views (Figs 15 B, C) apex bifurcate with medial aedeagus and three pairs of processes (Figs 14 A, 15 A); A 1 (process on opposite side hidden) very slender, extending from phallotheca apex, curved in upward medial catch, then broadly curved to anteriomedial direction (nearly crossing medially); A 2 and A 3; thicker, originating dorsally just subapically, crossing medially; A 5 and A 6 arising well subapically on ventral margin, arched to nearly cross midventrally (appearing forceps-like); aedeagus extending medially from the phallotheca; broad, moderately tapers, apex convex, extend about half the length of the phallathecal bifurcation. Gonostyli in lateral view (Fig. 14 A, B) broad, bearing a large smoothly hooked capitulum at the distal apex of the dorsal margin, with a small projection near apex, nearly straight along ventral margin, ventral margin folded medially, from ventral view (Fig. 14 D), medial margin nearly linear. Anal tube in lateral view (Fig. 14 A, B) elongate, distally downcurved; in dorsal view (Fig. 14 C) elongate-elliptical, apex bluntly rounded.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD139363FB2FBA31D505A17.taxon	discussion	Remarks. Atracodes vittatus comb. nov. & stat. nov. can be distinguished from the most similar species A. fasciculosus comb. nov., in the following features: The pygofer is broader in A. vittatus comb. nov. & stat. nov. (Fig. 14 A vs. Metcalf & Bruner 1948. Plate XVI, fig. 5), the gonostyli of A. vittatus comb. nov. & stat. nov. bears an apical projection on the ventral margin (smoothly rounded in A. fasciculosus comb. nov.), the ventral processes of the aedeagus of A. vittatus comb. nov. & stat. nov. are long (exceeding aedeagal midlength), but short and hooked in A. fasciculosus comb. nov. Metcalf and Bruner (1948) described A. vittatus comb. nov. & stat. nov. as a subspecies of A. fasciculosus comb. nov. due to similarity in coloration and patterning and they asserted that the general structure of the male terminalia were similar. We found that the male terminalia were substantially too different to be considered the same species. The male holotype in the NCSU collection is missing its head and prothorax. The mapped distribution is based on available specimens. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD139363FB2FBA31D505A17.taxon	distribution	Distribution. Cuba (Camagűey, Guantánamo, “ Santa Clara ” Provinces).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFD139363FB2FBA31D505A17.taxon	materials_examined	Type material examined. HOLOTYPE: Camagüey, Cuba. / J. Acuña, Col. / Aug. 20 1924 // ♂ // Holotype // Pseudoflatoides / fasciculosus / vittatus / Metc. & Brun. / det. Z. P. M. (NSCU, male). PARATYPES: San Blas, / Sta. Clara. / G. C. Rowe / 10 / 15 / 19 [31] // Paratype // Pseudoflatoides / fasciculosus / vittatus / Metc. & Brun. / det. Z. P. M. // UCC _ TCN 00103106 [2 D barcode label] (NCSU, male). Camagűey, Cuba, / J. Acuna, Col. / Aug. 15 1924 // Paratype // Pseudoflatoides / fasciculosus / vittatus / Metc. & Brun. / det. Z. P. M. // UDCC _ TCN 00102981 (NCSU, female). ALLOTYPE: Guantanamo / Cuba // Allotype // Pseudoflatoides / fasciculosus / vittatus / Metc. & Brun. / det. Z. P. M. // UDCC _ TCN 00102980 (NSCU, female).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCE39353FB2FB081EEA5FA7.taxon	description	(Fig. 16)	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCE39353FB2FB081EEA5FA7.taxon	materials_examined	Type locality. Rio Piedras, Puerto Rico. Synoptic diagnosis. Atracodes albus comb. nov. can be recognized from congeners by: covered in a waxy white to creamy white waxy pruinescence; head without dark lateral line near eye; hind tibia with 3 lateral teeth, apical spinulation 6 (4 + 2) - 7 - 2; phallothecal complex curving upward distally with three pairs of processes, one pair extending from apex, one stout pair extending from lateral margin near apex for about ½ the length of phallothecal complex, 1 pair extending near upcurve for length of phallothecal complex; anal tube in lateral view with ventral lobe on near mid length; apical margin of gonostyli nearly straight. Plant associations. Inga vera Willd. (Fabaceae), Lantana sp. (Verbenaceae).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCE39353FB2FB081EEA5FA7.taxon	distribution	Distribution. Puerto Rico.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCE39353FB2FB081EEA5FA7.taxon	materials_examined	Material examined. HOLOTYPE: P. R. Acc. No. / Rio Piedras, P. R. / 11 - 16 - [19] 47 // Caldwell / Martorell / Collectors // Pseudoflat / HOLOTYPE / alba // JSCaldwell / Collection / 1959 // Pseudoflatoides / albus male / Holotype / Caldwell (USNM, male); ALLOTYPE: P. R. Acc. No. 12 - 43. / Rio Piedras, P. R. / May 1, 1943 // G. N. Wolcott / Coll. // Pseudoflat / ALLOTYPE / alba // Pseudoflatoides / albus / det 49 / JSCaldwell // USNMENT 01145568. Other material examined. PUERTO RICO: / El Verde Forest / 7 Jul 1969 / R Bonilla, Coll / Light Trap // Pseudoflatoides / albus Caldwell / Det. J. A. Ramos // NCSU _ ENT 00286735 (NCSU, male). PUERTO RICO: / El Verde Forest / 7 Jul 1969 / R Bonilla, Coll / Light Trap // Pseudoflatoides / albus Caldwell / Det. J. A. Ramos // NCSU _ ENT 00286736 (NCSU, female). PUERTO RICO: / El Verde Forest / 7 Jul 1969 / R Bonilla, Coll / Light Trap // Pseudoflatoides / albus Caldwell / Det. J. A. Ramos // NCSU _ ENT 00286740 (NCSU, male). Castaner, P. R. / May 23, 1956 / ON Inga vera / Coll. Mario Perez // Pseudoflatoides / albus Caldwell / Det. J. A. Ramos // UDCC _ TCN 00102974, 00103108 (NCSU, 1 male, 1 female). PR: Cayey; mountain / pasture; 31 Jul 1996 / 18 07 N x 66 10 W / T. J. Henry and / R. G. Wheeler, Jr. // on / Lantana / sp. // Pseudoflatoides / albus Caldwell & Martorell, 1951 / Det. NNLevia 2024 // USNMENT / 01145523 (USNM, female). PUERTO RICO / Luquillo Forest / I- 4 - 1963, Paul & / Phylllis Spangler // El Yunque Biol. / Sta. MolinderoRoad / elevation 2.100 ’ / at black light // Pseudoflatoides / albus Caldwell & Martorell, 1951 / Det. NNLevia 2024 // UDCC _ TCN 00102962 – 00102965 (USNM, 2 male, 2 female). PUERTO RICO / Caribbean Nat. For / El Verde Field Sta. / Dec. 4, 1968 // J. Macdonald C. / Coll. 1996 // Pseudoflatoides / albus Caldwell & Martorell, 1951 // Det. NNLevia 2024 // UDCC _ TCN 00103116 (NCSU, male).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCD39343FB2F8981F835C17.taxon	materials_examined	Type locality. Cuba. Synoptic diagnosis. Atracodes fasciculosus comb. nov. can be distinguished from congeners by: white in color, vertex three times as long as broad; many cross veins in costal cells, forewing with nearly “ S ” shaped dark line present in the middle of the wing extending from ScP to A 2; hind tibia with 2 lateral teeth, apical spinulation 7 (5 + 2) - 7 - 2; pygofer narrow, apical margin nearly straight; phallotheca broad, weakly downcurved, abruptly narrowed near apex with subapical ventral lobe, with three pairs of processes, 2 slender dorsal pairs (apical longer than subapical) and 1 short stout ventral pair, gonostyli broad in lateral view, ventral margin nearly linear. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCD39343FB2F8981F835C17.taxon	distribution	Distribution. Cuba.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCD39343FB2F8981F835C17.taxon	materials_examined	Material examined. E. E. A / Ento. N 2. / 8768 // San Vincent / Pide’ Rio / 4 - 9 - 22 / SCByJ. A. // Pseudoflatoides / fasciculosus / var. / SCB // UDCC _ TCN 00103128 (NSCU, female). Camagűey, Cuba / J. Acuna, Col. / Aug. 15 1924 // Pseudoflatoides / fasciculosus / Melichar / det. Z. P. M // DLS 26 - D- 74 C / NCSU // UDCC _ TCN 00102975 (NCSU, female). Atracodes fasciculosus griscescens. 8768 // San Vincent / 4 - 6 - 9 - 22 / col. S. C. B. J / J. a. // Blank // ♂ // Pseudoflatoides / fasciculosus / griseus / Melichar / det. Z. P. M. // Pseudoflatoides / griseus / Mel. // UDCC _ TCN 00102979 (NCSU, male).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39343FB2FD091FE75BB7.taxon	materials_examined	Type locality. Cuba. Synoptic diagnosis. Atracodes insularis comb. nov. may be distinguished from congeners by: dirty yellow in color, forewings creamy white with yellow veins, mesonotum with two dark lateral lines near apical margin (Melichar 1902, plate 9. Fig. 10); three nearly lateral dark lines in coastal cells, dark line extending from costal margin near posterior of wing to ⅓ the length broken continuing to CuP, ScP darkened, two dark curved lines present, 1 near the base of the wing, 1 ¼ the length of the wing, dark curved line present about half the length of wing between ScP and CuP; vertex nearly as long as broad, frons nearly three times as long as broad; hind tibia with 2 lateral teeth, apical spinulation 2 - 6 - 8 (Medler 1994).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39343FB2FD091FE75BB7.taxon	discussion	Remarks. This species is excluded from the key because the holotype is female (Melichar 1902, Medler 1994) without definitively associated males. Medler (1994) restored this taxon to species status (after being treated as a subspecies) because comparative features of the ‘ tortrix complex of species’ (Medler 1994: 222) were not properly elucidated by Metcalf and Bruner (1948). A comparative study is needed, but is beyond the scope of the current project. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39343FB2FD091FE75BB7.taxon	distribution	Distribution. Cuba.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39343FB2FD091FE75BB7.taxon	materials_examined	Material examined. Cuba / Wright // PRUhler / Collection // Pseudoflatoides / insularis /? Mel. // USNMENT 01145546 (USNM, female).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39333FB2F9A91FB85E8F.taxon	description	(Fig. 7 B)	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39333FB2F9A91FB85E8F.taxon	materials_examined	Type locality. Pico Turquino, Santiago de Cuba (as Oriente) Province, Cuba. Synoptic diagnosis. Atracodes lichenoides comb. nov. can be distinguished from congeners by: body and tegmina relatively darkly marked, general color ochraceous and tawny; crossveins ir, r-m, m-cu, icu and between CuA and A 1 darkened; hind tibia with 2 lateral teeth, apical spinulation 7 (5 + 2) - 10 - 2; phallothecal complex nearly straight (Metcalf & Bruner 1948, plate XVI, fig. 3), appearing distally bilobed bearing two pairs of processes present, a slender subapical pair and a stout pair extending from lateral ventral margin; gonostyli broad in lateral view, ventral margin nearly linear. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39333FB2F9A91FB85E8F.taxon	distribution	Distribution. Cuba.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCC39333FB2F9A91FB85E8F.taxon	materials_examined	Material examined. June 10 – 29 / [19] 36 / J. Acuña, col. // Pico Turquino / Cuba, 1650 ft. / [illegible] // Pseudoflatoides / lichenoides / Metc. & Brun. det. Z. P. M. // UDCC _ TCN 00102983 (NCSU, male; this specimen may be the holotype).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCB39323FB2F92C1DCB5FFF.taxon	description	(Fig. 7 B)	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCB39323FB2F92C1DCB5FFF.taxon	materials_examined	Type locality. Dominica. Synoptic diagnosis. Atracodes longifrons can be distinguished from congeners by: yellowish-brown to stramineous color bearing a grayish green pruinescence; a dark lateral line present before eye, tegmina irregularly patterned; frons elongate lateral margins sinuate; hind tibia with 3 lateral teeth, apical spinulation 7 (5 + 2) - 8 - 2; phallothecal complex (Fennah 1965, Figs 93 – 95) slightly distally curved upward, bearing three pairs of processes, 2 dorsal pair slender and elongated, angled caudad basally and sharply curved retrorsely, last pair originating lateroventrad, directed retrorsely, curving dorsad; aedeagus distinct ventrally; anal tube from lateral view with concave ventral margin, without ventral median lobe on ental surface, subapical lateral margin angled and small ventral projection near apex.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCB39323FB2F92C1DCB5FFF.taxon	discussion	Remark. The type specimen of this species (reported to be at BMNH, Fennah 1965) could not be located. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCB39323FB2F92C1DCB5FFF.taxon	distribution	Distribution. Dominica.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCB39323FB2F92C1DCB5FFF.taxon	materials_examined	Material examined. DOMINICA / 2.2 mi E. of / Pont Casse / V- 2 - 1964 / O. S. Flint, jr. // Atracodes / longifrons Fennah, 1965 / Det. NNLevia 2023 // UDCC _ TCN 00102531 (UDCC, male).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39323FB2FE611DC55DD3.taxon	materials_examined	Type locality. Castries, St. Lucia Synoptic diagnosis. Atracodes sanctaeluciae can be distinguished from congeners by: patterning similar to A. indutus, covered in a grayish-white pruinescence; vertex 1.5 times longer than width between eyes (Fennah 1965, fig. 99); hind tibia with 3 lateral teeth, apical spinulation 6 - 5 - 2; phallothecal complex stout (Fennah 1965, fig. 100), curved upward bearing three pairs of elongated slender processes, 2 subapical pairs and 1 pair from the ventrolateral subapical margin projecting retrorsely, strongly curving dorsad. Anal tube in lateral view narrow, deflexed in distal third (Fennah 1965). Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39323FB2FE611DC55DD3.taxon	distribution	Distribution. St. Lucia.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39323FB2FE611DC55DD3.taxon	materials_examined	Material examined. HOLOTYPE: Type // ST. LUCIA / Castries / 4. XI. 1935 / H. E. Box // Atracodes / sanctaeluciae / det. RGFennah Fenn. // NHMUK 015981665 (BMNH, male).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39313FB2FC4D1D875FDA.taxon	description	(Fig. 7 D)	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39313FB2FC4D1D875FDA.taxon	materials_examined	Type locality. Cuba. Synoptic diagnosis. Atracodes tortrix comb. nov. can be distinguished from congeners by: color pale olive to buff bearing a dark median spot on the pronotum and two near posterior margin of mesonotum; vertex ⅕ longer than width between eyes; tegminal patterning irregular; hind tibia with 2 lateral teeth, apical spinulation 7 - 9 - 2 (Medler 1993 b); phallothecal complex (Medler 1993, fig. 7) in lateral view stout and broad, dorsal and ventral surfaces strongly irregular, bearing 3 pairs of processes, 2 pair dorsal (apical and subapical) very elongated, both pair reaching pygofer, 1 subapical ventral pair blade like; gonostyli in lateral view broad with ventral margin distinctly convex, apex rounded; anal tube in lateral view narrow and elongated, ventral margin broadly concave, without ventral processes.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39313FB2FC4D1D875FDA.taxon	discussion	Remarks. Medler (1993 b) designated and illustrated (ibid., fig. 7) a lectotype for this species in the MNMH and indicated that his previous citation of a holotype in the Naples Museum (Medler 1988) was in error and these specimens should be paralectotypes. Medler (1988) indicated that Metcalf and Bruner (1948) had misinterpreted this species (e. g., indicating the species was 9.0 – 10.5 mm whereas type material was 14 – 16 mm, Metcalf & Bruner 1948, Medler 1988, 1993 b). Review of species allied with A. tortrix comb. nov. was beyond the intended scope of this study. Plant associations. None.	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39313FB2FC4D1D875FDA.taxon	distribution	Distribution. Cuba, reported from Hispaniola (Uhler 1884).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
03A887C7FFCA39313FB2FC4D1D875FDA.taxon	materials_examined	Material examined. Lectotype: MUSEUM PARIS / CUBA / COLL. GUÉRIN-MENÉV. / ex COLL. A. SALLÉ 1897 // 4 // Museum Paris / MNMH (EH) / 609 // Phalaenomorpha / tortrix. Guer. / Voy Sagra (type) / Cuba // Type // LECTOTYPE / Flata / tortrix Guérin-M. / Desig. J. T. Medler, 1997 (MNMH, male). Other material examined. Camaguey, Cuba, / J. Acuna, Col. / Aug. 20 1924 // Pseudoflatoides / tortrix / Guer. Men. / det. Z. P. M. // UDCC _ TCN 00102982 (NCSU, female). Camaguey, Cuba, / J. Acuna Col. / Aug. 15 1924 // Pseudoflatoides / tortrix / Guer. Men. / det. Z. P. M. // UDCC _ TCN 00103109, 00103110 (NCSU, 2 females). Pico Turquino, / Cuba, [illegible, crossed out] / 375 feet. // June 10 – 29 / [19] 36 / J. Acuna, col. // Pseudoflatoides / tortrix / Guer. Men. / det. Z. P. M. // UDCC _ TCN 00103129 (NCSU, female).	en	Levia, Nathaniel N., Bartlett, Charles R. (2025): A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region. Zootaxa 5590 (2): 231-255, DOI: 10.11646/zootaxa.5590.2.4, URL: https://doi.org/10.11646/zootaxa.5590.2.4
