taxonID	type	description	language	source
03B187E6FFE51B54FF380FC4FB3350FF.taxon	materials_examined	Type species: Thoradonta dentata Hancock, 1909, by original designation.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE51B54FF380FC4FB3350FF.taxon	discussion	Remarks. This genus consists of 19 species distributed in India, China, Thailand, Vietnam, Malaysia, Indonesia, and Papua New Guinea; of them one species is known from the Philippines (Cigliano et al., 2025).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE51B54FF380D23FDAC563C.taxon	description	Figs 1, 2	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE51B54FF380D23FDAC563C.taxon	materials_examined	Material examined. Philippines: S Palawan, env. of Brooke’s Point (E cost), 6 – 8.03.2004, 2 ♀, leg. A. V. Gorochov (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE51B54FF380D23FDAC563C.taxon	discussion	Remarks. This species was described from the north part of Palawan Island (Günther, 1938). Here it is firstly recorded from the south part of this island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380EADFF395356.taxon	materials_examined	Type species: Paratettix personatus (Bolívar, 1887), by subsequent designation (Hancock, 1907). Composition. This large genus included 71 species from the continental Asia and Oriental region (Cigliano et al., 2025).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380F88FC6150FF.taxon	description	Figs 3, 4	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380F88FC6150FF.taxon	materials_examined	Material examined. Philippines: Mindanao Island, Mount Kitanglad Range Natural Park, 8.18 ° N, 124.94 ° E, h = 1400 m, 5.02.2024, 2 ♂, 2 ♀, leg. M. M. Omelko (FSCB).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380F88FC6150FF.taxon	distribution	Distribution. This species is widely distributed in India, Malaysia, Indonesia, Philippines, and Papua New Guinea (Bolívar, 1887; Blackith & Blackith, 1987; Bhaskar et al., 2019; Tumbrinck, 2019; Cigliano et al., 2025). Here E. personatus is recorded from Mindanao Island for the first time.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF38088DFE7855B2.taxon	materials_examined	Material. Philippines: Mindoro Island, env. of Puerto Galera (N cost), 11 – 13.03.2004, 1 female nymph of last instar, leg. A. V. Gorochov (FSCB).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF38088DFE7855B2.taxon	distribution	Distribution. This species was known from the Luzon Island. In this paper P. dentifer is recorded from Mindoro Island for the first time.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380BA6FD1E5451.taxon	materials_examined	Type species: Eugavialidium formosanum Tinkam, 1936, by original designation.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380BA6FD1E5451.taxon	discussion	Remarks. The genus consists of five species known from the south part of continental China (Fujian), Taiwan Island, and the Philippines (Cigliano et al., 2025).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380A46FC16577D.taxon	materials_examined	Material. Philippines: N Palawan, env. of Taytay (E cost), 25 – 26.02.2004, 1 ♀, 2 ♂, leg. A. V. Gorochov; S Palawan, env. of Brooke’s Point (E cost), 6 – 8.03.2004, 5 ♀, 6 ♂, leg. A. V. Gorochov (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380A46FC16577D.taxon	distribution	Distribution. This species was known from Binaluan in the north part of the Palawan Island (Günther, 1938). Here S. boettcheri is firstly recorded from the southern part of this island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380D5BFDE3561C.taxon	materials_examined	Type species: Tetrix producta Serville, 1838, by monotypy.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE61B57FF380D5BFDE3561C.taxon	discussion	Remarks. The genus consists of 23 species widely distributed in Oriental region from India and China to New Guinea Island (Cigliano et al., 2025).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE71B56FF380EE5FCC253C7.taxon	materials_examined	Type species: Diotarus verrucifer Stål, 1877	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE71B56FF380EE5FCC253C7.taxon	discussion	Remarks. This genus consists of four species from Philippines: Diotarus verrucifer Stål, 1877, endemic to Mindanao, D. galeatus Bolívar, 1887, endemic to Luzon (Angat and Norzagaray), D. ikonnikovi Bey-Bienko, 1935, also endemic to Luzon (Los Baños), and D. pupus Bolívar, 1887, known from Luzon (Camarines Sur) (Stål, 1877; Bolívar, 1887; Bey-Bienko, 1935; Mohagan et al., 2020).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE71B56FF380C78FDF65103.taxon	materials_examined	Material examined. Philippines: Mindanao Island, Cotabato Province, Apo Mt., 7.03 ° N, 125.22 ° E, h = 1350 m, 26 – 30.01.2024, 1 ♀, leg. M. M. Omelko (FSCB).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE71B56FF380C78FDF65103.taxon	distribution	Distribution. This species was known from the following localities in Mindanao Island: Zamboanga, Iligan City, Marilog Forest Reserve, Davao City (Mohagan et al., 2020). Here Diotarus verrucifer is recorded from a new locality situated not far from Davao.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE71B56FF380DF7FA45554A.taxon	materials_examined	Type species: Arulenus punctatus Stål, 1877, by monotypy.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE01B50FF380EADFBEB543E.taxon	description	Figs 5 – 14	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE01B50FF380EADFBEB543E.taxon	materials_examined	Material examined. Holotype — ♀, Philippines: Mindanao Island, Mount Kitanglad Range Natural Park, 8.18 ° N, 124.94 ° E, h = 1400 m, 27.01.2024, leg. M. M. Omelko (ZIN). Paratype: 1 ♂, the same locality and collector as for holotype but 05.02.2024 (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE01B50FF380EADFBEB543E.taxon	diagnosis	Diagnosis. The differences of a new species from other congeners are given in a key above.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE01B50FF380EADFBEB543E.taxon	description	Description. Female (holotype). Body medium-size for genus. Antennae with 12 black antennomeres: 1 st massive scapus, it width 2.2 times more than width of frontal ridge near the base of antennae; 2 nd short pedicel; 3 – 6 th antennomeres robust and shorter than the central segments; 7 – 8 th antennomeres elongated; 9 th and 10 th antennomeres flattened, 2.3 – 2.4 times as long as wide; apical segments fused and small. Length of antenna 2 times longer than length of fore femur. Eyes globose, in lateral view the upper margin of the eye not elevated above the pronotum. In dorsal view, vertex not projecting before the eyes, its width between eyes 1.4 times wider than width of an eye. Lateral carinae of the vertex not elevated above the eyes; medial carina of the vertex weak in the apical fourth. Frontal costa short, reaching the mid ocellus only. Lateral ocelli and bifurcation of frontal costa situated in the upper fourth of the compound eyes height. Dorsal margin of the antennal groove placed above the lower margin of compound eyes. Pronotum covering the whole abdomen, dorsally finely granulated. Anterior margin of pronotum truncated; median carina of pronotum low, almost straight in profile; prozonal carinae distinct, convergent; hind margin of lateral lobe of the pronotum without tegminal (upper) sinus; lower sinus deep; lower part of lateral lobe directed outwards, with obliquely truncate apex; infrascapular area narrow, lower margin of area straight. Tegmina and hind wings absent. Upper and lower carinae of fore and mid femora finely serrated; fore femur 5.6 times, mid femur 5.1 times as long as wide. Hind femur stout, 2.6 times as long as wide; upper and lower carinae finely dentate. Upper side of hind tibia with 3 outer and 4 inner teeth. First tarsal segment of hind legs as long as 3 rd segment (without claws), ventrally with 3 pulvilli almost equal in length. Subgenital plate as long as wide, with triangle posterior margin. Cerci conical with tapered apex. Ovipositor elongated and strongly armed by teeth and strong setae. Upper valvulae wider in lateral view than the lower ones. Upper valve of ovipositor 3.7 times longer than its maximum width; lower valve 5.2 times longer than its maximum width. Body blackish brown. Head brown; labrum black; maxillary palps light brown with blackish marks; eyes whitish. Antennae black. Dorsal side of the pronotum yellowish-brown with narrow black stripe along the median carina and wide black band between the bases of hind femora. Lateral lobes of pronotum blackish. Fore and mid legs black. Hind femora blackish brown with light marks. Hind tibiae black with light brown dorsal side. Hind tarsi black. Basal abdominal tergites black, apical tergites light brown. Sternal plate and abdominal sternites blackish brown; subgenital plate black. Ovipositor black. Male (paratype). Similar to female but smaller. Antennae 12 - segmented, 9 th and 10 th antennomeres flattened, 2.7 and 3.0 times as long as wide, respectively. Length of antenna 2.5 times longer than length of fore femur. Fastigium of vertex 1.7 times wider than width of one eye from above. Frontal ridge narrow. Pronotum as in female. Fore femur 2.7 times, mid femur 2.6 times, hind femur 3.45 times as long as wide. For and mid femora 5.0 times, hind femur 2.7 times as long as wide. Epiproct narrow triangular, with pointed apex. Subgenital plate in lateral view triangle. Body colored as in female but antennae blackish brown, dorsal side of the pronotum in anterior part light brown and in apical part blackish brown; lateral lobes of the pronotum light brown. Subgenital plate, 9 th abdominal sternite and cerci blackish. Measurements (in mm). Length of body (from frontal ridge to apex of subgenital plate) male 7.5, female 9.3; pronotum male 7.0, female 9.4; antenna male 4.9, female 5.6; fore femur male 2.1, female 2.8; mid femur male 2.0, female 2.6; hind femur male 4.6, female 5.5; ovipositor 1.3.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE01B50FF380EADFBEB543E.taxon	distribution	Distribution. Philippines: Mindanao Island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE01B50FF380EADFBEB543E.taxon	etymology	Etymology. The species name refers to the type locality, Kitanglad Mountain.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE11B53FF380860FA4550F0.taxon	materials_examined	Type species: Cleostratus monocerus Stål, 1877, by subsequent designation (Kirby, 1910).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE21B5DFF380D23FF2952E2.taxon	description	Figs 15 – 23	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE21B5DFF380D23FF2952E2.taxon	materials_examined	Material examined. Holotype — ♂, Philippines: Mindanao Island, Mount Kitanglad Range Natural Park, 8.18 ° N, 124.94 ° E, h = 1400 m, 15.02.2024, leg. M. M. Omelko (ZIN). Paratype: 1 ♀, the same locality and collector as for holotype but 13.02.2024 (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE21B5DFF380D23FF2952E2.taxon	diagnosis	Diagnosis. The differences of a new species from other congeners are given in a key above.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE21B5DFF380D23FF2952E2.taxon	description	Description. Male (holotype). Body of moderate size for genus, slender. Antennae 14 - segmented; length of antenna 1.3 times as long as fore femur; mid segments of antennae 7.5 – 7.8 times as long as wide. Antennal grooves situated at the level of the upper margin of eyes. Fastigium of vertex forming the relatively short triangular rostrum directed forwards; rostrum 1.1 times shorter than the horizontal diameter of one eye seen from lateral view; fastigium between eyes almost as wide as one eye seen from dorsal view; median carina of fastigium absent; supraocular lobes small; in dorsal view rostrum narrowly triangular with truncate apex. Frontal ridge in lateral view almost straight between eyes; apex of ridge directed gently upward; lateral carinae of the frontal ridge in frontal view almost parallel; width of first antennal segment 1.3 times as wide as frontal ridge between antennae. Pronotum in dorsal view with straight anterior margin; posterior process of pronotum far surpass the apex of hind femora. Median carina of pronotum in lateral view low, straight; lateral carinae in prozona distinct, straight; prozona subsquare; humero-apical carinae distinct; external lateral carinae almost straight; extralateral carinae reaching apex of pronotum; interhumeral carinae distinct. Posterior margin of the lateral lobe of the pronotum with tegminal (upper) sinus as broad as ventral sinus; lower side of lateral lobes in dorsal view gently directed outwards, broadly rounded. Visible part of tegmen 3.0 times as long as wide and almost as wide as maximal width of mid femora. Hindwing long, surpass the apex of the posterior process of pronotum for 0.2 mm. Fore and mid femora with straight upper and lower margins, 4.7 – 4.9 times as long as wide. Hind femur 3.3 times as long as wide, with gently dentate upper and lower carinae. Dorsal side of hind tibia with 6 outer and 5 inner spines and numerous denticles between them. First tarsal segment of hind legs as long as 3 rd segment (without claws), ventrally with 3 pulvilli almost equal in length. Subgenital plate in lateral view elongated; in dorsal view plate 2 times as long as wide, apex of plate with narrow excision. Cerci conical, 1.7 times as long as wide near the base. Female (paratype). Similar to male but apical segments of antennae and both hind legs missing. Rostrum 1.1 times shorter than the horizontal diameter of one eye seen from lateral view; fastigium between eyes 1.15 times as wide as one eye seen from dorsal view; width of first antennal segment 1.4 times as wide as frontal ridge between antennae. Visible part of tegmen 2.8 times as long as wide. Hindwings surpass the apex of the posterior process of pronotum on 1.3 mm. Fore femur 4.4 times, mid femur 4.6 times as long as wide. Epiproct narrowly triangular with pointed apex. Subgenital plate 1.1 times as long as wide, posterior margin triangle. Cerci conical, 1.6 times as long as wide. Valves of ovipositor narrow; upper valve dorsally and lower valve ventrally with row of teeth and armed by long setae; upper valve 3.7 times, lower valve 5.2 times as long as its maximum width. General coloration of the male and female body blackish-brown. Eyes light brown; face in frontal view brown with blackish marks, black rostrum and lower part of the labrum; antennae black with narrow light rings at apices of antennomeres. Dorsal side of pronotum blackish; lateral lobes of pronotum brown with blackish marks. Visible part of tegmen black with light crossveins. Hindwings black with light spots. Fore and mid femora brown; hind femur with black marks. Hind tibia black. Abdomen blackish brown; female subgenital plate and ovipositor blackish brown. Measurements (in mm). Length of body (from frontal ridge to apex of subgenital plate) male 10.8, female 11.9; pronotum male 13.2, female 13.1; antenna male 2.8; tegmen male 1.2, female 1.7; fore femur male 2.1, female 2.2; mid femur male 2.3, female 2.3; hind femur male 5.9; ovipositor 1.4.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE21B5DFF380D23FF2952E2.taxon	distribution	Distribution. Philippines: Mindanao Island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE21B5DFF380D23FF2952E2.taxon	etymology	Etymology. Named for arachnologist Mikhail Mikhailovich Omelko, who collected the type specimens of this species.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5DFF380F1CFBCD5024.taxon	materials_examined	Type species: Centrosotettix centrosus (Bolívar, 1887), by original designation. Composition. Till now this genus included a type species only known from Malaysia (Borneo Island: Sarawak Province) (Günther, 1938; Cigliano et al., 2025). A second species is found in the Philippines and described here.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5DFF380F1CFBCD5024.taxon	discussion	Remarks. The genus Centrosotettix is new for the fauna of the Philippines.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5CFF380C5FFEA1543A.taxon	description	Figs 24 – 30	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5CFF380C5FFEA1543A.taxon	materials_examined	Material examined. Holotype — ♀, Philippines: Mindanao Island, Mount Kitanglad Range Natural Park, 8.18 ° N, 124.94 ° E, h = 1400 m, 17.02.2024, leg. M. M. Omelko (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5CFF380C5FFEA1543A.taxon	diagnosis	Diagnosis. New species similar to Centrosotettix centrosus but differs from later by eyes in lateral view slightly protruding above pronotum, visible part of tegmen with light crossveins, and hind tibia black (in C. centrosus eyes in lateral view strongly protruding above pronotum, visible part of tegmen black, and hind tibia light brown).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5CFF380C5FFEA1543A.taxon	description	Description. Female (holotype). Body slender, medium-size for genus. Eyes in lateral view slightly protruding above pronotum. Antennae filiform, 16 - segmented; antennal grooves situated at the level of lower margin of eyes. Length of antenna 1.9 times longer than length of fore femur; mid segments of antennae 7.5 – 8.5 times as long as wide. Lateral ocelli placed slightly above the middle part of the eyes. Fastigium of vertex almost reaching the front of eyes from dorsal aspect, 1.5 times narrower than width of one eye from above; anterior margin of fastigium almost straight; median carina short; lateral carinae slightly elevated. Width of 1 st antennal segment 2.5 times more than width of frontal ridge near base of antennae. Pronotum in dorsal view with straight anterior margin; posterior process of pronotum far surpass the apex of hind femora. Median carina of pronotum in profile weakly elevated between shoulders and straight in apical part of pronotum. Prozonal carinae parallel, prozona 1.5 times as wide as long. Hind margin of lateral lobe of pronotum bisinuate, tegminal (upper) sinus distinct; lower sinus deep; lower part of lateral lobe directed outwards, with obliquely truncate apex; infrascapular area narrow, lower margin of area straight. Tegmina elongated, narrow, width of visible part of tegmen almost equal to the width of mid femur; hind wings reaching the apex of the posterior process of pronotum. Fore and mid femora 4.0 – 4.4 times as long as wide, with straight and finely dentate upper and lower margins. Hind femur slender, 3.3 times as long as wide; upper and lower carinae finely dentate. Upper side of hind tibia with 6 outer and 4 – 5 inner teeth. First tarsal segment of hind legs as long as 3 rd segment (without claws). Subgenital plate transverse, with excised posterior margin and triangle middle projection. Cerci conical, 1.5 times as long as wide near base, with acute apex. Valves of ovipositor long and narrow; upper valve dorsally and lower valve ventrally with row of teeth and long setae; upper valve 4.6 times, lower valve 7.8 times as long as its maximum width. Body grayish brown with black marks. Face in frontal view light brown with black spots; clipeal triangle and lower part of the labrum black; maxillary palps whitish. Antennae blackish; antennomeres of flagellum with light rings at apex; apical antennomeres whitish. Dorsal side of the pronotum grayish brown with small black spots along median carina; lateral lobes light brown with black oblique stripe. Visible part of tegmen with light crossveins. Fore and mid femora grayish brown. Fore and mid tibiae brown with two black rings. First segment of fore and mid tarsi blackish, 2 nd light brown with black apex; claws brown. Lower outer area of hind femora black. Hind tibiae black with brown ring in the basal part. First segment of hind tarsi whitish, 2 nd segment black, 3 rd segment with light with black apex; claws brown. Abdominal tergites black with small light brown spots. Sternal plate black. Abdominal sternites yellowish with black longitudinal stripe. Subgenital plate yellowish. Ovipositor yellowish with brown apex. Male unknown. Measurements (in mm). Length of female body (from frontal ridge to apex of subgenital plate) 9.5; pronotum 13.0; antenna 3.8; tegmen 1.5; fore femur 2.01; mid femur 2.2; hind femur 5.6; ovipositor 1.6.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5CFF380C5FFEA1543A.taxon	distribution	Distribution. Philippines: Mindanao Island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEC1B5CFF380C5FFEA1543A.taxon	etymology	Etymology. The species name is given because the holotype was found on the high altitude in the mountain region of the island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFED1B5FFF380865FA4453BE.taxon	type_taxon	(type species: Bolotettix validispinus Hancock, 1907, by original designation).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFED1B5FFF380865FA4453BE.taxon	discussion	Taxonomic remarks. The type species of the genera Bolotettix (B. validispinus from Malaysia: Sarawak: Kuching) and Syzygotettix (S. boettcheri from Philippines: Palawan: Binaluan) characterized by almost the same diagnostic features and undoubtedly belongs to one genus. Therefore, a new synonymy is proposed: Bolotettix Hancock, 1907 = Syzygotettix Günther, 1938, syn. nov. Here five species are transferred from Syzygotettix to Bolotettix and new combinations are established: Bolotettix baeri (Bolívar, 1887), comb. nov., Bolotettix boettcheri Günther, 1938, comb. nov., Bolotettix semperi Günther, 1938, comb. nov., Bolotettix uncinatus (Stål, 1877), comb. nov., and Bolotettix vicinus Günther, 1938, comb. nov.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEE1B5FFF380FE0FD0955CD.taxon	description	(Figs 31 – 41)	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEE1B5FFF380FE0FD0955CD.taxon	materials_examined	Material examined. Holotype — ♀, Philippines: Mindanao Island, Mount Kitanglad Range Natural Park, 8.18 ° N, 124.94 ° E, h = 1400 m, 28.01.2024, leg. M. M. Omelko (ZIN). Paratype: 1 ♂, the same data as for holotype (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEE1B5FFF380FE0FD0955CD.taxon	diagnosis	Diagnosis. New species is most similar to Bolotettix perminutus described from Luzon Island by Bolívar (1887) by oblique truncated lateral lobes of the pronotum but easily recognizable from it by small size and stout hind femora (in latter species, length of pronotum in male 8.4 – 10, in female 8.5 – 12.8 mm, and hind femur 3 – 3.2 times as long as wide).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEE1B5FFF380FE0FD0955CD.taxon	description	Description. Female (holotype). Body of small size for genus. Antennae filiform, 15 - segmented; antennal grooves situated slightly above level of the lower margin of eyes. Length of antenna 2 times longer than length of fore femur; mid segments of antennae 5.2 – 5.4 times as long as wide. Lateral ocelli placed slightly above the level of lower margin of eyes. Fastigium of vertex with concavities between lateral carinae, reaching the front of eyes from dorsal aspect, 1.1 times narrower than width of one eye from above; anterior margin of fastigium broadly rounded; median carina short; lateral carinae slightly elevated. Width of 1 st antennal segment equal to width of frontal ridge near base of antennae. Median carina of pronotum in profile low, slightly wavy. Prozonal carinae parallel; prozona 1.5 times as wide as long. Hind margin of lateral lobe of pronotum bisinuate, tegminal (upper) sinus distinct; lower sinus deep; lower part of lateral lobe obliquely truncate; infrascapular area narrow, lower margin of area straight. Tegmina elongated, width of visible part of tegmen equal to width of mid femur. Hindwings almost reaching the apex of the posterior process of pronotum. Fore and mid femora with straight and finely dentate upper and lower margins. For femur 3.5 times, mid femur 3.6 times as long as wide. Hind femur stout, 2.7 times as long as wide. Upper side of hind tibia with 6 outer and 5 – 6 inner teeth. First tarsal segment of hind legs 1.4 times as long as 3 rd segment (without claws); tarsal pulvilli elongated, almost equal in size. Epiproct narrow triangular, with pointed apex. Subgenital plate elongated; apex of plate triangle. Cerci conical. Valves of ovipositor short; upper valve dorsally and lower valve ventrally with row of teeth and short setae; upper valve 3 times, lower valve 4.3 times as long as its maximum width. Body blackish brown. Face in frontal view light brown; clipeal triangle blackish; lower part of labrum black; maxillary palps whitish. Basal part of antennae light brown, apical antennomeres blackish. Dorsal side of pronotum blackish; lateral lobes light brown with black oblique stripe. Visible part of tegmen black. Fore and mid tibiae brown with two black rings. First segment of fore and mid tarsi blackish, 2 nd light brown with black apex; claws brown. Outer area of hind femora black with light spot at base. Hind tibiae black. Abdominal tergites laterally black, dorsally light brown. Abdominal sternites and subgenital plate light brown. Ovipositor brown. Male (paratype). Similar to female but smaller. Antennae 15 - segmented. Width of the eye from above 1.1 times wider than width of fastigium of vertex between eyes. Width of 1 st antennal segment 1.1 times more than width of frontal ridge near base of antennae. Pronotum, tegmina, and hind wings as in female. Fore femur 2.7 times, mid femur 2.6 times, hind femur 2.7 times as long as wide. Epiproct narrow triangular, with pointed apex. Subgenital plate in lateral view triangle with acutely rounded apex; in ventral view apex of plate weakly excised. Cerci conical, with pointed apex. Body colored as in female but dorsal side of pronotum and outer area of hind femora black; abdominal tergites black except brown apical ones; cerci and subgenital plate light brown. Measurements (in mm). Length of body (from frontal ridge to apex of subgenital plate) male 5.9, female 6.1; pronotum male 6.5, female 7.0; antenna male 2.3, female 2.8; tegmen male 0.9, female 1.0; fore femur male 1.3, female 1.4; mid femur male 1.4, female 1.6; hind femur male 3.8, female 4.4; ovipositor 0.9.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEE1B5FFF380FE0FD0955CD.taxon	distribution	Distribution. Philippines: Mindanao Island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEE1B5FFF380FE0FD0955CD.taxon	etymology	Etymology. From Latin ‘ nanus’ means dwarf.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEF1B58FF38092BFD0C53C2.taxon	description	(Figs 42 – 49)	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEF1B58FF38092BFD0C53C2.taxon	materials_examined	Material examined. Holotype — ♂, Philippines: Mindanao Island, Mount Kitanglad Range Natural Park, 8.19 ° N, 124.92 ° E, h = 1350 m, 3.02.2024, leg. M. M. Omelko (ZIN). Paratype: 1 ♂, the same data as for holotype (ZIN).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEF1B58FF38092BFD0C53C2.taxon	diagnosis	Diagnosis. The new species is most similar to Bolotettix semperi by small size and stout hind femora, but differs from latter in the shape of spine of lateral lobes of pronotum (in B. semperi spine directed outwards). By directed obliquely backwards spine B. parvus sp. nov. is similar to B. luzonicus, but easily recognizable from it by stout hind femora (in latter species hind femur 2.9 – 3 times as long as wide).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEF1B58FF38092BFD0C53C2.taxon	description	Description. Male (holotype and paratype). Body of small size for genus. Antennae filiform, 15 - segmented; antennal grooves situated at the level of the lower margin of eyes. Length of antenna 1.8 times longer than length of fore femur; mid segments of antennae 4.5 – 4.7 times as long as wide. Lateral ocelli placed distinctly above the level of lower margin of eyes. Fastigium of vertex with concavities between lateral carinae, reaching the front of eyes from dorsal aspect, 1.1 times narrower than width of one eye from above; anterior margin of fastigium rounded; median carina reaching middle of eyes; lateral carinae slightly elevated. Width of 1 st antennal segment 1.1 times wider than width of frontal ridge near base of antennae. Median carina of pronotum in profile low, wavy. Prozonal carinae converge backwards, prozona 1.2 times as wide as long. Hind margin of lateral lobe of pronotum bisinuate, tegminal (upper) sinus distinct; lower sinus deep; lower part of lateral lobe with spine directed obliquely backwards. Tegmina elongated; width of visible part of tegmen 1.5 – 1.6 times narrower than width of mid femur. Hindwings short, not reaching the apex of the posterior process of pronotum. Fore and mid femora with finely dentate upper and lower margins. For femur 2.8 – 3 times, mid femur 3 times as long as wide. Hind femur stout, 2.1 times as long as wide. Upper side of hind tibia with 4 – 5 outer and 4 inner teeth. First tarsal segment of hind legs 1.2 times as long as 3 rd segment (without claws); tarsal pulvilli elongated, almost equal in size. Epiproct narrow triangular, with pointed apex. Subgenital plate in lateral view elongated; in ventral view apex of plate excised. Cerci conical, 1.6 times as long as wide. Body of holotype blackish brown. Head blackish; antennae brown. Dorsal side of the pronotum blackish. Fore and mid femora brown with indistinct black marks. Fore and mid tibiae light brown with two black rings. Upper outer area of hind femora brown, lower area black. Hind tibiae black. Abdominal tergites black, apical segments dorsally light brown. Abdominal sternites light brown. Body of paratype colored as in holotype but lateral lobes and dorsal side of pronotum before shoulders light brown. Female unknown. Measurements (in mm). Length of male body (from frontal ridge to apex of subgenital plate) 5.5 – 6.4; pronotum 5.4 – 5.6; antenna 2.3; fore femur 1.1 – 1.3; mid femur 1.4 – 1.5; hind femur 3.9.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEF1B58FF38092BFD0C53C2.taxon	distribution	Distribution. Philippines: Mindanao Island.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFEF1B58FF38092BFD0C53C2.taxon	etymology	Etymology. From Latin ‘ parvus’ means small.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE91B58FF380C7CFED2563B.taxon	materials_examined	Type species: Systolederus haani Bolívar, 1887, by subsequent designation (Rehn, 1904).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE91B58FF380C7CFED2563B.taxon	discussion	Remarks. During long time, Teredorus Hancock, 1907 and Systolederus Bolívar, 1887 were considered as separate genera (Otte, 1997). Recently, almost all species of the genus Teredorus from the Old World were transferred to the genus Systolederus (Devriese & Husemann, 2023). Undoubtedly, three species of Teredorus described from China and Cambodia (Zheng, 2015; Zha & Ding, 2017; Storozhenko, 2019) must be also placed in Systolederus and new combinations are proposed here: Systolederus longidorsalis (Zheng, 2015), comb. nov., Systolederus jinshaensis (Zha et Ding, 2017), comb. nov., and Systolederus vicinus (Storozhenko, 2019), comb. nov. Nowadays four species are known from the Philippines: S. femoralis (Walker, 1871), S. angusticeps (Stål, 1877), S. haani Bolívar, 1887, and S. boettcheri Günther, 1939 (Walker, 1871; Stål, 1877; Bolívar, 1887; Günther, 1939; Kevan, 1966; Cigliano et al., 2025).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE91B58FF380A67FBD2571C.taxon	materials_examined	Material. Philippines: Mindoro Island, env. of Puero Galera (N cost), 11 – 13.03.2004, 4 ♂, 6 ♀, leg. A. V. Gorochov (ZIN and FSCB).	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
03B187E6FFE91B58FF380A67FBD2571C.taxon	distribution	Distribution. This Philippine species was known from the Luzon, Panaon and Mindanao islands (Günther, 1939; Kevan, 1966). Here it is recorded from Mindoro Island for the first time.	en	Storozhenko, Sergey Yu. (2025): New and little known pygmy grasshoppers (Orthoptera: Tetrigidae) from the Philippines. Zootaxa 5661 (3): 351-366, DOI: 10.11646/zootaxa.5661.3.3, URL: https://doi.org/10.11646/zootaxa.5661.3.3
