identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B18E5CFF9DFFC45AAD4AF6FA4BFC55.text	03B18E5CFF9DFFC45AAD4AF6FA4BFC55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neurigoninae	<div><p>Key to tribes of Neurigoninae and genera of Neotropical Neurigonini (modified from Naglis 2003)</p><p>1 Thorax strongly elongated (Naglis 2001a, figs 3A, C); abdomen about as long as thorax and dorsoventrally flattened; vertex excavated dorsally between vt s and ocellar tubercle; male It5 with ventral comb of short blunt spines (Naglis 2001a, fig. 3D); arista subapical (Naglis 2001a, fig. 1B); surstylus with one single lobe (Naglis 2001a, figs 1E, 2C, 3G)........ Coeloglutini</p><p>– Thorax not elongated (Figs 1A, 3C); abdomen usually about 1.3–1.5 times longer than thorax and not dorsoventrally flattened; vertex not excavated (Figs 1B, 3B); male It5 without ventral comb of short spines; arista usually dorsal (Fig. 1C); surstylus divided into dorsal and ventral lobes (Fig. 2A–D)........................................................... 2</p><p>2 Thorax usually metallic green (except Naglisia, with thorax yellow); dc s consisting of 2–3 pairs of setae (except Argentinia Parent, with 6–7 pairs); wing vein M S-shaped, joining costa distinctly anteriad of apex close to vein R 4+5, with distance between R 4+5 and M 1 at costa usually less than half-length of crossvein dm-m; legs usually without major setae on tibia II and/or tibia III; female oviscapt with tergites 9+10 blunt and rounded, with sternite 10 (“cercus” in Naglis’ terminology) free from tergite 9+10 (Quevedo et al. 2025, fig. 2A–E)................................................... Dactylomyiini</p><p>– Thorax metallic green or yellow; dc s consisting of 5–7 pairs of setae; wing vein M 1 usually straight or bent, joining costa at apex or posteriad of apex, with distance between R 4+5 and M 1 at costa usually more than half-length of crossvein dm-m (except Mberu, Figs 4F, 5G, 6F); legs usually with major setae on tibia II and/or tibia III; female oviscapt with tergites 9+10 elongate and tapered, with sternite 10 fused with tergite 9+10 (Fig. 22B, C) … Neurigonini .................................. 3</p><p>3 Male abdominal segment 5 with long sickle-shaped ventral projection covering entire ventral border of hypopygium (Naglis 2002a, fig. 3A); hypopygium large and at base remarkably projecting above dorsum of abdomen (Naglis 2002a, fig. 3A); male coxa II with ventral lobe bearing long sinuate or flattened setae (Naglis 2002a, fig. 3A, B); male It4+5 broadened and pennate bilaterally (Naglis 2002a, figs 3A, 5A); phallus divided into two arms with multi-furcate tip.......... Bickelomyia Naglis</p><p>– Male abdominal segment 5 with or without ventral projection, if present then not sickle-shaped and only partially covering ventral border of hypopygium; hypopygium at base not projecting above dorsum of abdomen; male coxa II unmodified; male It4+5 rarely pennate; phallus usually present as single arm with pointed tip....................................... 4</p><p>4 Postpedicel triangular, with acute apex (Figs 1C, 5C, 6C, 16C, 20C); costal section between R 4+5 and M 1 shorter than half-length of crossvein dm-m, M 1 ending anterior to wing apex (Figs 1F, 4F, 5G); It1–5 with ventral surface entirely bare (Fig. 5F); female tergite 9+10 with 4–6 spines at apex (Fig. 22B, C, D; Capellari &amp; Amorim 2011, figs 8, 9)..... Mberu Capellari &amp; Amorim</p><p>– Postpedicel subtriangular, with rounded apex; costal section between R 4+5 and M 1 usually longer than half-length of crossvein dm-m, M 1 ending at or posteriad to wing apex (Naglis 2003, fig. 1A); It1–5 covered by setae on ventral surface; female tergite 9+10 without spines at apex............................................................................ 5</p><p>5 Thorax entirely metallic green; abdomen metallic bronze-green with segments 1 and 2 often yellow; tibia II usually with only 1–2 ad setae; IIt1 devoid of major setae; proboscis without pair of long ventral hairs; male It4 usually dorsoventrally flattened, with ventral pile; male cercus often elongated, with modified setation.............................. Viridigona Naglis</p><p>– Thorax mainly ochreous-yellow, metallic green at most on mesonotal depression and scutellum, or sometimes dark brown, with or without dense pruinosity; abdomen ochreous-yellow, usually with dark tergal bands; tibia II usually with ad, pd and ventral setae; IIt1 usually with strong posterior seta basally and additional strong setae; proboscis with pair of long sinuous ventral hairs; male It4 not modified; if modified, not dorsoventrally flattened with ventral pile; male cercus short and rounded, with short setae........................................................................... Neurigona Rondani</p></div>	https://treatment.plazi.org/id/03B18E5CFF9DFFC45AAD4AF6FA4BFC55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF9CFFC45AAD4804FAE3F9D0.text	03B18E5CFF9CFFC45AAD4804FAE3F9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu Capellari & Amorim 2011	<div><p>Mberu Capellari &amp; Amorim</p><p>Mberu Capellari &amp; Amorim, 2011: 39 . Type species: Mberu pepocatu Capellari &amp; Amorim, 2011: 40 (original designation).</p><p>Diagnosis. Male. Antennal postpedicel triangular, with acute apex, arista-like stylus dorsally inserted (Figs 1C, 5C, 20C). Posterior slope of mesonotum distinctly flattened; acr s biseriate, six pairs of dc s (three posteriormost conspicuously larger). Wing membrane hyaline (Figs 4F, 6F) or sometimes with conspicuous modifications of the venation (some species of pepocatu group) (Figs 3F, 5G, 7D), veins R 4+5 and M 1 beyond dm-m parallel (Fig. 6F) or converging at wing apex (Figs 3F, 4F, 5G); distance between R 4+5 and M 1 at costal section between R 4+5 and M 1 shorter than half-length of crossvein dm-m (Figs 3F, 4F, 5G, 6F, 7D); M 1 ending anterior of wing apex (Figs 1F, 3F, 4F, 5G, 6F, 7D). Legs sometimes with conspicuous modified rows of setae (MSSC) (Figs 3E, 4C, D, 5D, 6D, 16D, 18C, E); tarsus I with ventral surface entirely bare (MSSC) (Fig. 5F). Female. Similar to male, except for MSSCs and as noted: Wing membrane hyaline or sometimes with conspicuous modifications of the venation (some species of pepocatu group, but less pronounced than in males) (Figs 3G, 5H, 6G). Oviscapt telescoping into proximal segments, acanthophorites (tergites 9+10) flattened, prolonged and fused with cercus, dorsally divided in two disconnected hemitergites, and with apical spines (Capellari &amp; Amorim 2011, figs 8, 9; Fig. 22B–D).</p></div>	https://treatment.plazi.org/id/03B18E5CFF9CFFC45AAD4804FAE3F9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF9CFFC35AAD4A89FA4BF9B9.text	03B18E5CFF9CFFC35AAD4A89FA4BF9B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu undefined-1 Capellari & Amorim 2011	<div><p>Key to males of Mberu species</p><p>1 Base of femur I with anterior bush-like comb of setae (Figs 6D, 7B); It1 longer than remaining tarsomeres combined (Fig. 5D); femur II with ventral surface bare, except for anteroventral row of setae, decreasing in length towards apex, and 1 av long and sometimes sinuous near apex (Figs 4D, 5E) … pepocatu group................................................ 2</p><p>- Base of femur I without conspicuous differentiated setae; It1 shorter than remaining tarsomeres combined; ventral surface of femur II completely bare............................................................................... 7</p><p>2 Wing venation without modifications and spurious veins, dm-m straight, M 4 complete, ending at wing margin (Figs 1F, 4F, 6F)................................................................................................ 3</p><p>- Wing venation with modifications well defined, reticulated, with short spurious veins arising from R 4+5, M 1, dm-m or M 4; dm-m distorted, S-shaped; M 4 incomplete, not reaching wing margin (if reaching margin, just branch, not main vein) (Figs 3F, 5G, 7D)................................................................................................ 5</p><p>3 Eyes touching on face (Fig. 6B); wing vein M 1 with distinct flexion S-shaped beyond crossvein dm-m, R 4+5 and M 1 parallel apically to costa (Fig. 6F); femur II and tibia II without conspicuous row of setae (Fig. 6E) [Brazil: Goiás and Mato Grosso do Sul]............................................................................... Mberu lopesi sp. nov.</p><p>- Eyes not touching on face (Figs 1B, 4B); wing vein M 1 slightly concave at anterior margin, without distinct flexion, R 4+5 and M 1 beyond crossvein dm-m converging apically to costa (Figs 1F, 4F); femur II with row of ad setae, tibia II with row of anterior setae (Figs 1E, 3E, 7C)................................................................................ 4</p><p>4 Lower surface of occiput with ventral pocl s sparse and straight, as long as vt s, and some sparse and straight white ocp s, increasing in size ventrally, the ventralmost 1.3 times as long as ventral pocl s (Fig. 1B, C); setae on anterior surface of coxa I shorter than width of coxa (Fig. 1D); wing membrane light brown, slightly darker along anterior margin and at apex (Fig. 1F) [Brazil: Goiás].................................................................... Mberu altamiro sp. nov.</p><p>- Lower surface of occiput with ventral pocl s dense and sinuous, two times as long as oc s, and numerous dense and sinuous white ocp s, increasing in size ventrally, the ventralmost as long as lower pocl s (Fig. 4B, C); setae on anterior surface of coxa I longer than width of coxa (Fig. 4C); wing membrane completely hyaline (Fig. 4F) [Brazil: Distrito Federal and Goiás]...................................................................................... Mberu ericae sp. nov .</p><p>5 Wing venation slightly modified, with short spurious veins arising from M 1, dm-m and M 4; R 4 +5 without spurious veins; M 4 ending in small brown spot at apex (Fig. 3F); It 1 not swollen at basal 1/3 [Brazil: Mato Grosso do Sul]................................................................................................ Mberu bodoquena sp. nov .</p><p>- Wing venation conspicuously modified, reticulated, with short spurious veins arising perpendicularly from R 4+5, M 1, dm-m and M 4 (Figs 5G, 7D); M 4 not ending in spot; It1 ventrally swollen at basal 1/3 (Fig. 5D)............................... 6</p><p>6 Wing membrane light brown, slightly darker along anterior margin, apex of R 4 +5 and M 1 slightly anastomosed, but not forming brownish spots at their apices (Fig. 7D) [Brazil: Minas Gerais]............... Mberu pepocatu Capellari &amp; Amorim, 2011</p><p>- Wing membrane hyaline, apex of R 4 +5 and M 1 greatly anastomosed, with anastomoses forming brownish spots touching each other or nearly so (Fig. 5G) [Brazil: Distrito Federal, Goiás and Mato Grosso do Sul].............. Mberu jawara sp. nov .</p><p>7 Tibia III dorsally swollen on apical half and comb of posterior pale setae inclined to apex (Figs 12E, 18E); DSur (Figs 13D, 19D) irregularly shaped and robust, with curved ventral margins and truncated, rounded distal lobe; phallus with short dorsal projection near apex (Figs 13G, 19F)..................................................................... 8</p><p>- Tibia III not conspicuously swollen and without comb of setae; DSur narrow and elongated, with straight ventral margins (Figs 9D, 11D, 15D, 17D, 21D); phallus without dorsal projection near apex (Figs 9G, 11G, 17F, 21E)...................... 9</p><p>8 Coxa I with conspicuous posterior hook-shaped projection (Fig. 12C); It1 with distinct row of spine-like setae (Fig. 12D) [Brazil: Mato Grosso do Sul and Rio Grande do Sul; Paraguay: Coronel Oviedo; Argentina: Corrientes]................................................................................................... Mberu lamasi sp. nov.</p><p>- Coxa I without conspicuous projection (Fig. 18C); It1 without modified setae (Fig. 18D) [Brazil: Santa Catarina and Rio Grande do Sul]..................................................................... Mberu soaresi sp. nov .</p><p>9 Eyes touching on face (Figs 14B, 20B); MEp branched at apex (Figs 15G, 21H).................................. 10</p><p>- Eyes not touching on face (Figs 8B, 10C, 12B, 16B, 18B); MEp not branched at apex (Figs 9F, 11E, 13E)............. 11</p><p>10 Body length about 2 mm; postpedicel about 2.4 times as long as basal width (Fig. 20B, C); male hypopygium large, elongate, reaching forward to abdominal segment 2 (Fig. 21A, B) [Brazil: Goiás]...................... Mberu takwajasu sp. nov.</p><p>- Body length less than 1.6 mm; postpedicel about 1.6 times as long as basal width (Fig. 14B, C); male hypopygium stout, not elongate, reaching forward to abdominal segment 4 (Fig. 15A) [Brazil: Mato Grosso do Sul]... Mberu murtiniensis sp. nov.</p><p>11 DSur about 3.7 times as long as high, curved near apex (Fig. 17D), conspicuously projecting beyond VSur (Fig. 17A, B) [Brazil: São Paulo]................................................................. Mberu periotoi sp. nov.</p><p>- DSur about 2.7–3.0 times as long as high, straight, not conspicuously projecting beyond VSur (Figs 9A, B, D; 11A, B, D).. .................................................................................................. 12</p><p>12 Legs completely yellow (Fig. 8A); MEp stout, about 2.5 times narrower than VSur, with rough dorsal face (Fig. 9C, F) [Brazil: Rio Grande do Sul]................................................................ Mberu amorimi sp. nov.</p><p>- Legs mostly dark brown (Fig. 10A); MEp narrow, about 4.5 times narrower than VSur, with smooth dorsal face (Fig. 11C, E) [Brazil: Goiás and São Paulo].......................................................... Mberu britoi sp. nov .</p></div>	https://treatment.plazi.org/id/03B18E5CFF9CFFC35AAD4A89FA4BF9B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF9BFFC35AAD4A9AFAE2F8CE.text	03B18E5CFF9BFFC35AAD4A9AFAE2F8CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu pepocatu Capellari & Amorim 2011	<div><p>The pepocatu group</p><p>Diagnosis (Based on males). Base of femur I with 2 long pv setae, with anterior bush-like comb of setae (Figs 6D, 7B); It1 longer than remaining tarsomeres combined (Fig. 5D); femur II with ventral surface bare, except for av row of setae, decreasing in length towards apex, ending in one long and sometimes sinuous av seta near apex (Figs 4D, 5E); the hypopygial morphology is quite similar for all species in the group (as in Fig. 2, see Discussion).</p></div>	https://treatment.plazi.org/id/03B18E5CFF9BFFC35AAD4A9AFAE2F8CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF9AFFC05AAD4CA6FC51F80D.text	03B18E5CFF9AFFC05AAD4CA6FC51F80D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu altamiro Silva & Capellari & Oliveira 2025	<div><p>Mberu altamiro sp. nov.</p><p>(Figs 1, 2)</p><p>Description. Male. Body length: 2.23 mm. Wing length: 2.24 mm long, 0.57 mm wide (n = 1). Body mostly dark brown to black, with metallic reflections. Head (Fig. 1B, C, D). Frons width 0.3 times head width, at narrowest part; eyes almost touching on face, narrowest distance in middle of face less than diameter of anterior ocellus; face slightly wider at ventral and dorsal ends, with silver pruinosity ventral to antenna, darker in middle, lower part of face and clypeus with dense silver pruinosity; palpus dark brown, with 2–3 apical setulae; proboscis dark brown, with short whitish hairs; one proclinate convergent vt s; one reclinate divergent oc s, as long as vt s; one reclinate convergent poc s, 0.4 times as long as vt s; occiput distinctly concave above, shiny dark brown; lower surface of occiput covered by complete row of white pocl s, increasing in size ventrally, ventralmost sparse and straight, as long as vt s; lower occiput covered by some white, straight ocp s, increasing in size ventrally, 1.3 times as long as lower pocl s (Fig. 1B, C, D). Scape short, yellow, dorsal surface bare; pedicel yellow with dorsal surface dark brown, with crown of dark apical setae; postpedicel dark brown, triangular, about 1.5 times longer than basal width, pubescent; stylus about 5 times longer than postpedicel, dorsally inserted, bare, displaced laterally, bi-articulated at base. Thorax. Dark brown to black, with some dark green reflections, 0.7 times as long as abdomen, posterior slope of mesonotum distinctly flattened, postpronotal lobe with lower pale spot. Short scattered setae from anterior side of mesonotum to slightly beyond transverse suture; two rows of 13–15 small acr s, ending at mesonotal slope; 6 dc s, increasing in size posteriorly, posterior two conspicuously longer; 1 psut ial s, 1 presut spal s, 2 large psut spal s; 2 pprn s, one strong and one tiny white seta; 2 npl s, 2 pal s, one large and one setula anteriad; 1 pair of strong l sctl s and 1 pair of smaller laterad setae, about half as long as l sctl s; 2 white prepst s. Wing (Fig. 1F). Membrane light brown, with darker brown spot below wing apex; veins brown. C ending before wing apex, only slightly extended beyond M 1; Sc ending at middle of R 1; R 1 ending at about basal third of wing; R 2+3 and R 4+5 strongly displaced towards anterior margin as compared to R 1; anterior cells strongly compressed, R 2+3 ending in C at apical fourth of wing; R 4+5 and M 1 convergent near wing margin; M 4 complete, reaching wing margin; CuA+CuP strong, not reaching wing margin. Lower calypter, halter and cilia white. RMx ratio = 1.7, CuAx ratio = 0.3. Legs. Mostly brown, yellow at apex of femur II, base of tibia II, apex of femur III, middle of tibia III, basal half of IIIt1; pair of claws present on all legs. Pulvilli small. Leg I (Fig. 1D). Coxa I covered with white, fine setae (shorter than coxa I width), decreasing in length towards apex, apex with 3–4 strong, pale apical setae anteriorly. Femur I with 4–5 small, pale pv setae at base, with anterior bush-like comb of brown setae (e.g. Fig. 7B). Tibia I covered by fine setae, longer dorsally. It1 with ventral surface entirely bare (e.g. Fig. 5F), dorsal surface covered by dark setae; It1 longer than remaining tarsomeres combined (Fig. 1D). Leg II. Coxa II with a few pale setulae anteriorly, bare laterally. Femur II (Fig. 1E) with ad row of small, inconspicuous brown setae along apical third, increasing in length towards apex; ventral surface bare, except for anteroventral row of 7–8 sparse, pale setae along basal 3/4, decreasing in length towards apex, and one sinuous, brown av seta (e.g. Fig. 4D). Tibia II with row of dark, erect dorsal setae (Fig. 1E). Leg III. Coxa III with one conspicuous, long, pale seta near base and one setula above. Femur III with dorsal row of dark setae, decreasing in length towards apex. Tibia III with one dorsal brown seta at distal 1/4, followed by one row of dark, dorsal setulae apically; comb of pale, apical setae, projecting towards to tarsal excavation. IIIt1 ventrally excavated at base. Relative podomere ratios: I: 14.7/ 11.5/ 6.2/ 1.3/ 1/ 1/ 1.2; II: 23.4/ 23.8/ 11.6/ 1/ 1/ 1.4/ 2; III: 13.1/ 13.9/ 5.3/ 3.6/ 1.6/ 1/ 1. Abdomen. Mostly dark brown, with some dark blue to green reflections on tergites 4–5. Tergites 1–5 with short, black setae. Segment 6 partially hidden under tergite 5, tergite 6 bare; segment 7 forming short peduncle, tergite and sternite 7 bare; sternite 8 small, triangular. Sternites 1–3 visible, sclerotized, light brown, sternite 4 membranous. Hypopygium (Fig. 2). Epandrium elongate, 1.3 times as long as high, dark (Fig. 2B). MEp elongate, about 7.5 times as long as wide (Fig. 2G). DSur and VSur covering internal appendages (Fig. 2B); VSur (Fig. 2C) enlarged, twice higher than DSur (Fig. 2D), with two inner apicoventral setae, probably LEp fused with VSur (see Quevedo et al. 2025); DSur (Fig. 2D) irregular and robust, with curved ventral margins and truncated, rounded distal lobe. Hypandrium basally fused to epandrium. Phallus with short, dorsal projection near apex (Fig. 2F). Cercus (Fig. 2H) divided in two parts, ventral part curved and stout, with 3–4 spines ventrally, dorsal part rounded and covered by fine setae, with digitiform projection apically.</p><p>Female. Unknown.</p><p>Type material. HOLOTYPE ♂, “ BRAZIL, GO [Goiás], Goianápolis | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.11381&amp;materialsCitation.latitude=-16.53488" title="Search Plazi for locations around (long -49.11381/lat -16.53488)">Parque Estadual [State Park] Altamiro de Moura</a> | Pacheco-PEAMP | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.11381&amp;materialsCitation.latitude=-16.53488" title="Search Plazi for locations around (long -49.11381/lat -16.53488)">Malaise mata [Seasonal Semi-deciduous Forest]</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.11381&amp;materialsCitation.latitude=-16.53488" title="Search Plazi for locations around (long -49.11381/lat -16.53488)">margem do córrego [stream bank]</a> | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.11381&amp;materialsCitation.latitude=-16.53488" title="Search Plazi for locations around (long -49.11381/lat -16.53488)">Carapina – Trilha da Onça</a> | -16.534879, -49.113812 [16°32′05.6″S 49°06′49.7″W] - 790m | 29.vi.2022 - 02.viii.2022 | Cols. Oliveira, S.S. &amp; Lopes, W.R. ”, “HOLOTYPE | Mberu altamiro | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia dissected and stored in microvial with glycerin, pinned beneath the specimen; left wing mounted in permanent slide, glued to paper triangle and pinned beneath specimen.</p><p>4+5</p><p>Etymology. Toponymic, referring to the type-locality and treated as a noun in apposition.</p><p>Remarks. Mberu altamiro sp. nov. can be distinguished from the other species of the pepocatu group (those with vein M 1 converging with R 4+5 at costa) by the absence of elongated ventral pocl s and ocp s (Fig. 1B, C), which in the other species are as long as the vt s (Figs 3B, C) and by the lack of long anterior basal setae on coxa I (Fig. 1D). Additionally, M. altamiro sp. nov. has a light brown wing membrane with a dark brown spot below the wing apex, and it lacks spurious veins on the wings (Fig. 1F). Furthermore, the rows of setae on femur II and tibia II are less developed than in the other species that exhibit these structures (Fig. 1E).</p><p>Distribution (Fig. 24). Brazil: state of Goiás (Cerrado biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFF9AFFC05AAD4CA6FC51F80D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF97FFCF5AAD4CA6FC7CF8C7.text	03B18E5CFF97FFCF5AAD4CA6FC7CF8C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu bodoquena Silva & Capellari & Oliveira 2025	<div><p>Mberu bodoquena sp. nov.</p><p>(Fig. 3)</p><p>Male. Body length: 2.41–2.75 mm. Wing length: 2.16–2.86 mm longer, 0.54–0.80 mm wide (n = 4). Similar to M. altamiro sp. nov. in both general habitus and overall chaetotaxy, except as noted. Head (Figs 3B, C). Complete row of white pocl s, increasing in size ventrally, ventralmost dense and sinuous, as long as anterior setae at base of coxa I (Fig. 3C); lower occiput covered by numerous white, sinuous ocp s, increasing in size ventrally, as long as lower pocl s (Fig. 3B, C). Pedicel yellow, with dorsal surface brown; postpedicel about 1.45 times longer than basal width; stylus about 4.9 times longer than postpedicel. Thorax (Fig. 3C). Brown, with some dark green reflections. Wing (Fig. 3F). Membrane hyaline, veins brown; venation reticulated, with five short spurious veins arising from M 1 and one from crossvein dm-m; dm-m distorted, S-shaped; M 4 not reaching wing margin, ending at small brown spot near wing margin; RMx ratio = 2.8, CuAx ratio = 2.5. Legs (Fig. 3B–E). Legs mostly brown, yellow at apex of tibia II, IIt1, femur III at apex, and tibia III near base and on apical 3/4. Leg I. Anterior surface of coxa I largely covered with white, fine setae, decreasing in length towards apex, basalmost ones longer, sinuous, and conspicuously longer than width of coxa I (Fig. 3C) (MSSC); base of femur I with 2–4 long, white pv setae, sometimes bunched together and 2–3 tiny hairs posteriad (MSSC); apex of femur I with 2–3 strong dorsal setae (MSSC); tibia I with row of 8–10 ad brown setae at basal half, decreasing in size towards apex (MSSC). Leg II. Femur II with ad row of strong, conspicuous brown setae along apical 2/3, decreasing in length towards apex (Fig. 3E) (MSSC). Tibia II with anterior row of long, conspicuous brown setae, strongly curved medially (Fig. 3E) (MSSC). Leg III. Coxa III with one conspicuous, long, pale seta near base, one pale setula posteriad and one pale setula anteriad. Relative podomere ratios: I: 1.5/ 10.4/ 6.6/ 1.5/ 1.3/ 1/ 1.1; II: 13.8/ 9.8/ 10.9/ 3.7/ 2.4/ 1.4/ 1; III: 11.8/ 13.4/ 5.6/ 5.8/ 3.3/ 1.4/ 1.1.</p><p>Hypopygium. As in M. altamiro sp. nov. (Fig. 2) (see Discussion). Female. Body length: 2.8–3.3 mm. Wing: 2.5–2.9 mm long, 0.9–1.1 mm wide (n = 3). Habitus similar to male, except for MSSCs and as noted. Head. Face slightly broader (about as broad as ocellar tubercle), slightly wider towards clypeus, clypeal suture more evident. Wing (Fig. 3G). R 4+5, M 1 and crossvein dm-m without spurious veins, sometimes just with some dark points near veins, variable among specimens; R 4+5 only slightly concave on distal half, not as close to margin as in males; M 1 complete, reaching wing margin; crossvein dm-m less distorted; RMx ratio = 4.4, CuAx ratio = 0.6. Legs. Mostly yellow. Coxa II and III dark brown; femora yellow to pale brown; coxa I covered by short, scattered pale setae anteriorly; tibia II with one ad at 1/4 and one near middle, and four apical setae. Relative podomere ratios: I: 9.9/ 8.9/ 4.5/ 2.6/ 1.5/ 1/ 1.2; II: 9.5/ 9.2, 7.2/ 3.5/ 1.9/ 1.3/ 1; III: 9.6/ 11.8/ 3.9/ 4.4/ 2.3/ 1.4/ 1. Abdomen. Tergite 1–5 with short vestiture of black setae. Sternites light brown, covered by a few black setae. Oviscapt telescoping into presegments, acanthophorites (tergites 9+10) flattened, prolonged and fused to cerci, with apical spines (e.g. Fig. 22).</p><p>Type material. HOLOTYPE ♂, “SISBIOTA - | CNPQ/FAPESP [Vertical line] | BRASIL [BRAZIL], MS [Mato Grosso do Sul], Bodoquena, | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.880386&amp;materialsCitation.latitude=-20.69886" title="Search Plazi for locations around (long -56.880386/lat -20.69886)">Fazenda [Farm] Califórnia</a> (topo [top]) | 20º41′55.9″S 56º52′49.4″W | Malaise 6, 8-22.viii.2011, | Lamas, Nihei et al. leg.”, “HOLOTYPE | Mberu bodoquena | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Mato Grosso do Sul. Same data as holotype (1 ♂, 3 ♀, one dissected, MZUSP). Same data except, 22.viii.2011 (1 ♂, MZUSP). Same data except, 22.viii–6.ix.2011 (1 ♂, MZUSP) .</p><p>Etymology. Toponymic, referring to the type-locality and treated as a noun in apposition.</p><p>Remarks. Mberu bodoquena sp. nov. is similar to M. ericae sp. nov., M. pepocatu and M. jawara sp. nov., with a number of male secondary sexual characters, such as modified setation on the legs and conspicuously modified wing venation, but M. bodoquena sp. nov. can be differentiated by wing venation in males slightly modified, with short spurious veins arising from M 1 and dm-m; dm-m distorted, S-shaped; M 4 not reaching wing margin, ending in a little brown spot near wing margin (Fig. 3F), while in M. ericae sp. nov. the wing membrane is unmodified (Fig. 4F) and in M. pepocatu and M. jawara sp. nov. the wing membrane bears more pronounced modifications (Figs 5G, 7D).</p><p>Distribution (Fig. 24). Brazil: state of Mato Grosso do Sul (Cerrado biome).</p><p>Comments. Mberu bodoquena sp. nov. is one of the morphospecies collected in Mato Grosso do Sul (Brazil) during SISBIOTA-Diptera Brazilian Network (see Lamas et al. 2023).</p></div>	https://treatment.plazi.org/id/03B18E5CFF97FFCF5AAD4CA6FC7CF8C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF95FFCD5AAD4CA6FBBAF9C3.text	03B18E5CFF95FFCD5AAD4CA6FBBAF9C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu ericae Silva & Capellari & Oliveira 2025	<div><p>Mberu ericae sp. nov.</p><p>(Fig. 4)</p><p>Description. Male. Body length: 2.38–3.09 mm. Wing length: 2.20–2.61 mm long, 0.58–0.68 mm wide (n = 7). Similar to M. altamiro sp. nov. in both general habitus and chaetotaxy, except as noted. Body mostly dark brown with metallic reflections. Head (Fig. 4B, C) Complete row of white pocl s, increasing in size ventrally, ventralmost dense and sinuous, as long as anterior setae at base of coxa I; lower occiput covered by numerous white, sinuous ocp s, increasing in size ventrally, as long as lower pocl s. Pedicel completely yellow; postpedicel about 1.3 times longer than basal width; stylus about 5.3 times longer than postpedicel. Thorax (Fig. 4C). Brown, with some dark green reflections. Wing (Fig. 4F). Membrane completely hyaline. RMx ratio = 3.3, CuAx ratio = 0.2. Legs (Fig. 4A, C–E). Mostly brown, yellow on apical 1/4 of tibia II, apical 1/4 of IIt1, femur III at apex, tibia III near base and at apex, and basal 1/4 of IIIt1. Leg I. Anterior surface of coxa I largely covered with white setae, decreasing in length towards apex, basalmost setae sinuous, conspicuously longer than width of coxa I (Fig. 4C); base of femur I with 2–4 long pv white setae, sometimes bunched together, and 2–3 tiny hairs posteriad, apex of femur I with 2–3 strong dorsal setae (MSSC); tibia I with row of 8–10 ad brown setae on basal half, decreasing in size towards apex (MSSC). Leg II. Femur II with ad row of strong brown setae along apical 2/3, decreasing in length towards apex (Fig. 4E); ventral surface completely bare, except for av row of white setae on basal 3/4, decreasing in length towards apex, ending in one sinuous, brown av seta (Fig. 4D); tibia II with one anterior row of long, conspicuous brown setae, strongly curved medially (Fig. 4D, E). Relative podomere ratios: I: 20.9/ 18.1/ 11.3/ 2.5/ 1.4/ 1/ 1.4; II: 6.3/ 5/ 5.8/ 1/ 1/ 1.1/ 1; III: 7.4/ 9.9/ 4/ 5/ 3/ 1.5/ 1. Abdomen. Mostly brown, with some dark blue to green reflections. Hypopygium. As in M. altamiro sp. nov. (Fig. 2) (see Discussion). Female. Unknown.</p><p>Type material. HOLOTYPE ♂, “ BRAZIL, DF [Distrito Federal], TAGUATINGA | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.07122&amp;materialsCitation.latitude=-15.829166" title="Search Plazi for locations around (long -48.07122/lat -15.829166)">Chácara Frutos da Terra [organic vegetable farm]</a> | 15°49′45.0″S, 48°04′16.4″W | Malaise T 3 – 10/08/2012 [08.x.2012] | msp 09.26 | Harterreiten-Souza, E.S col.”, “HOLOTYPE | Mberu ericae | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Distrito Federal. Same data as holotype (1 ♂, MZUSP) . Brasília, Fazenda Água Limpa-UnB, Córrego da Capetinga, 4–12.ix.2024, Malaise Gressit &amp; Gressit, F. S. P. Godoi e equipe col. (4 ♂, MZUSP) . Goiás. Mineiros, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.996418&amp;materialsCitation.latitude=-17.902222" title="Search Plazi for locations around (long -52.996418/lat -17.902222)">Parque Nacional das Emas</a>, Malaise - Parcela 2 Mata, 17°54′08.0″S, 52°59′47.1″W, 04.ix–08.x.2017, Cols. Oliveira &amp; Lopes (1 ♂, dissected, ZUFG) .</p><p>Etymology. Named after Érica Sevilha Harterreiten-Souza, who collected the holotype of this species and studied Dolichopodidae in the Brazilian Cerrado, especially in agricultural areas.</p><p>Remarks. Mberu ericae sp. nov. is similar to M. altamiro sp. nov. in general habitus and chaetotaxy, but they can be differentiated by wing membrane completely hyaline in M. ericae sp. nov. (Fig. 4F), the lower surface of occiput is covered by one row of pocl s and one row of ocp s, both white, sinuous, dense, regularly spaced, increasing in size ventrally, and the lowermost two times as long as oc s (Fig. 4B), and the anterior surface of coxa I covered with setae, the basal ones are sinuous and conspicuously longer than the width of coxa I (Fig. 4C). In M. altamiro sp. nov., the wing membrane is light brown, slightly darker along the anterior margin and at apex (Fig. 1F); the lower surface of the occiput is covered by one complete row of pocl s and one row of ocp s, both straight, sparse, and irregularly spaced, increasing in size ventrally, as long as vt s (Fig. 1B), the anterior surface of coxa I is covered by setae shorter than the width of coxa I (Fig. 1D). Mberu ericae sp. nov. is one of the seven morphospecies collected by Harterreiten-Souza et al. (2020), but only M. ericae sp. nov. and M. jawara sp. nov. have males in good enough condition to be described.</p><p>Distribution (Fig. 24). Brazil: Distrito Federal and state of Goiás (Cerrado biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFF95FFCD5AAD4CA6FBBAF9C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF95FFCA5AAD4A8DFEDCF8ED.text	03B18E5CFF95FFCA5AAD4A8DFEDCF8ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu jawara Silva & Capellari & Oliveira 2025	<div><p>Mberu jawara sp. nov.</p><p>(Fig. 5)</p><p>Description. Male. Body length: 2.55–3.15 mm. Wing length: 2.38–2.91 mm long, 0.84–1.00 mm wide (n = 10). Similar to M. altamiro sp. nov. in both general habitus and chaetotaxy, except as noted. Head (Figs 5B, C). Complete row of white pocl s, increasing in size ventrally, ventralmost dense and sinuous, as long as anterior setae at base of coxa I; lower occiput covered by numerous white, sinuous ocp s, increasing in size ventrally, as long as lower pocl s. Pedicel completely yellow; postpedicel about 1.6 times longer than basal width; stylus about 4.9 times longer than postpedicel. Thorax (Fig. 5C). Brown, with some dark green reflections. Wing (Fig. 5G). Membrane hyaline, veins brown, venation greatly modified, reticulated, with 9–11 short spurious veins arising perpendicularly from R 4+5, 13–15 from M 1, 6–8 from dm-m and 9–11 from M 4, those at apex of R 4+5 and M 1 high anastomosed, with anastomoses forming spots touching each other or nearly so, with most spurious veins brownish at apex, so that pattern of spurious veins consistent among specimens (MSSC). RMx ratio = 1.7, CuAx ratio = 2.1. Legs. Mostly brown, yellow at apex of femur I, apex of tibia I, apical 3/4 of tibia II, basal 3/4 of IIt1, apical 3/4 of tibia III and basal 3/4 of IIIt1. Femora covered by black setae dorsally, some white setae laterally. Leg I. Coxa I largely covered with white, fine setae, decreasing in length towards apex, basalmost ones longer, sinuous and conspicuously longer than width of coxa I (MSSC); base of femur I with 2–4 long pv white setae, sometimes bunched together, with 2–3 tiny hairs posteriad, apex of femur I with 2–3 strong, dorsal setae (MSSC); tibia I with row of 8–10 brown ad setae at basal half, decreasing in size towards apex (MSSC); It 1 ventrally swollen at basal 1/3 (Fig. 5D) (MSSC). Leg II. Femur II with ad row of strong, conspicuous brown setae along apical 2/3, decreasing in length towards apex, ventral surface completely bare, except av row of pale setae along basal 3/4, decreasing in length towards apex, ending in one sinuous, brown av seta (Fig. 5E) (all MSSC); tibia II with anterior row of long, conspicuous brown setae, strongly curved medially (MSSC). Leg III. Coxa III with one conspicuous, long, white seta near base; tibia III with dorsal row of small white setae (MSSC). Relative podomere ratios: I: 12.5/ 8.8/ 7/ 2.1/ 1.3/ 1/ 1; II: 10/ 8.9/ 8.7/ 3.3/ 1.9/ 1/ 1; III: 9.9/ 10.4/ 4.3/ 4.8/ 2.6/ 1.4/ 1. Abdomen. As in M. altamiro sp. nov., except in some specimens with tergite 1 and tergite 2 yellow on basal third. Hypopygium. As in M. altamiro sp. nov. (Fig. 2) (see Discussion).</p><p>Female. Body length: 2.6–3.3 mm. Wing: 2.5–2.9 mm long, 0.9–1.1 mm wide (n = 3). Habitus similar to male, except for MSSCs and as noted. Head. Face slightly broader (about as broad as ocellar tubercle), slightly wider towards apex, clypeal suture evident. Wing (Fig. 5H). Spurious veins on R 4+5 and M 1 significantly shorter, fewer in number and restricted to distal parts of veins, variable in quantity and size among specimens; no spurious veins on dm-m; R 4+5 only slightly concave on distal half, not as close to wing margin; M 1 bent anteriorly on distal third; RMx ratio = 2.6; CuAx ratio = 0.5. Legs. Mostly yellow. Coxa II and III dark brown, sclerotized; femora yellow to pale brown; darker at apex of tibia I, apex of It1, It2–5, apex of tibia II, IIt1 at apex, IIt2–5, apex of tibia III, IIt1 at apex and IIt2–5; coxa I covered by short scattered pale setae anteriorly; tibia II with 1 anterior seta at 1/4, 2 ad at 1/3 and 2/3, 1 pd at apex, 1 ad, 1 av, and 1 ventral setae at apex. Relative podomere ratios: I: 13.8/ 8.1/ 4.8/ 2.8/ 1.3/ 1/ 1; II: 8.8/ 9/ 6.9/ 3.6/ 2/ 1.2/ 1; III: 9.3/ 11.2/ 3.4/ 4.2/ 2.2/ 1.1/ 1. Abdomen. Tergite 1–5 with short vestiture of black setae. Sternites light brown, covered by few black setae. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “ BRAZIL, GO [Goiás], Varjão | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.554276&amp;materialsCitation.latitude=-17.032528" title="Search Plazi for locations around (long -49.554276/lat -17.032528)">Chácara [Farm] N. Sra. Aparecida</a> | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.554276&amp;materialsCitation.latitude=-17.032528" title="Search Plazi for locations around (long -49.554276/lat -17.032528)">MalaiseMata [Semideciduous Seasonal Forest</a>] | 17°01′57.1″S 49°33′15.4″W | 12.viii.2018 - 09.ix.2018 | Col. Lopes W.R. ”; “HOLOTYPE | Mberu jawara | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Goiás. Same data as holotype (2 ♂, 1 ♀, MZUSP). Same data except, 27.vi–08.viii.2021 (2 ♂, INPA). Same data except, 14.vii–11.viii.2019 (1 ♂, ZUFG). Goianápolis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.113804&amp;materialsCitation.latitude=-16.53489" title="Search Plazi for locations around (long -49.113804/lat -16.53489)">Parque Estadual Altamiro de Moura Pacheco-PEAMP</a>, Malaise Mata, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.113804&amp;materialsCitation.latitude=-16.53489" title="Search Plazi for locations around (long -49.113804/lat -16.53489)">margem do córrego Carapina – Trilha da Onça</a>, 16°32′05.6″S 49°06′49.7″W, 29.vi–02.viii.2022, cols. Oliveira, S.S. &amp; Lopes, W.R. (2 ♂, one dissected, 2 ♀, MNRJ) . Mato Grosso do Sul. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.07122&amp;materialsCitation.latitude=-15.829166" title="Search Plazi for locations around (long -48.07122/lat -15.829166)">Aquidauana. Res. Ecol. UEMS</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.07122&amp;materialsCitation.latitude=-15.829166" title="Search Plazi for locations around (long -48.07122/lat -15.829166)">Mata Ciliar, Córrego Fundo</a>, 20º26′03.7″S 55º39′20.8″W, 27.viii– 11.ix.2011, Malaise 7. Lamas, Nihei e eq. Col (1 ♂, dissected, MZUSP). Distrito Federal. Taguatinga, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.07122&amp;materialsCitation.latitude=-15.829166" title="Search Plazi for locations around (long -48.07122/lat -15.829166)">Chácara Frutos da Terra</a>, 15°49′45.0″S, 48°04′16.4″W, Malaise T3, 24.viii.2012, msp 09.50, Harterreiten-Souza, E.S. col. (1 ♂ MZUSP) .</p><p>Etymology. From the Tupi-Guarani, one of the native Brazilian linguistic families, “jawára ” (jaguar), treated as a noun in apposition, alluding to the conspicuous wing vein pattern similar to the jaguar coat spots.</p><p>Remarks. Mberu jawara sp. nov. is similar to M. pepocatu, and both share a number of male secondary sexual characters, such as the conspicuously modified wing venation (Figs 5G, 7D). They can be differentiated by wing membrane hyaline in males of M. jawara sp. nov., with veins at the apex of R 4+5 and M 1 greatly anastomosed, with anastomoses forming spots that touch each other or almost so (Fig. 5G), while in M. pepocatu the wing membrane is light brown, slightly darker along anterior margin, those at the apex of R 4+5 and M 1 slightly anastomosed, and the membrane around the apex of most spurious veins is brownish (Fig. 7D).</p><p>Distribution (Fig. 24). Brazil: Distrito Federal and states of Goiás and Mato Grosso do Sul (Cerrado biome).</p><p>4+5</p></div>	https://treatment.plazi.org/id/03B18E5CFF95FFCA5AAD4A8DFEDCF8ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF91FFC95AAD4CA6FC52F9C3.text	03B18E5CFF91FFC95AAD4CA6FC52F9C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu lopesi Silva & Capellari & Oliveira 2025	<div><p>Mberu lopesi sp. nov.</p><p>(Figs 6, 22B–D)</p><p>Description. Male. Body length: 1.79–2.26 mm. Wing length: 1.87–2.11 mm long, 0.49–0.72 mm wide (n = 10). Similar to M. altamiro sp. nov. in both general habitus and chaetotaxy, except as noted. Head (Fig. 6B, C, D). Eyes touching on face (Fig. 6B); one pair of oc s as long as vt s. Scape brown; postpedicel about 1.6 times longer than basal width; stylus about 5.6 times longer than postpedicel. Thorax (Fig. 6C). Six dc setae, increasing in size posteriorly, posteriormost three conspicuously longer than anteriormost setae. Wing (Fig. 6F). Membrane hyaline. R 4+5 and M 1 converging beyond dm-m, parallel near wing apex. RMx ratio = 4.7, CuAx ratio = 0.04. Legs (Fig. 6A–E). Mostly yellow, brown at anterior surface of coxa I, basal 2/3 of femur I, tibia II at apex, IIt1 at apex, IIt2–5, tibia III at apex, IIIt1 at apex and IIIt2–5. Leg I. Apex of coxa I with ventral comb of white seta; base of femur I with 4–5 small, white pv setae, decreasing in size towards apex, with anterior bush-like comb of white setae (Fig. 6D), apex of femur I with 2–3 strong, dorsal setae, apex of femur I with 2–3 strong, dorsal setae (all MSSC); tibia I with row of three brown ad setae on basal half, decreasing in size towards apex (MSSC). Leg II. Coxa II with 4–5 white setae anteriorly, increasing in size towards apex, bare laterally (Fig. 6C); femur II without ad row of setae along apical third (Fig. 6E), ventral surface completely bare, except anteroventral row of 12–13 pale setae, strongly decreasing in length towards apex, ending in one sinuous, brown av (all MSSC); tibia II without row of ventral setae (Fig. 6E). Leg III. Femur III without dorsal row of dark setae, with ad row of pale setae decreasing in size towards apex. Relative podomere ratios: I: 14.3/ 11.2/ 7.7/ 2/ 1.3/ 1/ 1.1; II: 10.1/ 10.3/ 8.7/ 3.3/ 1.2/ 1; III: 7.1/ 9/ 3.4/ 3.3/ 1.9/ 1.1/ 1. Abdomen. As in M. altamiro sp. nov. Hypopygium. As in M. altamiro sp. nov. (Fig. 2) (see Discussion).</p><p>Female. Body length: 2.03–2.50 mm. Wing: 1.79–2.05 mm long, 0.58–0.72 mm wide (n = 10). Habitus similar to male, except for MSSCs and as noted. Head. Eyes not touching on face; face slightly broader (about as broad as ocellar tubercle), slightly wider towards apex, clypeal suture evident. Wing. S-shaped curve of M 1 less pronounced (Fig. 6G). RMx ratio = 3, CuAx ratio = 0.2. Legs. Coxa I covered by short, scattered white setae anteriorly; tibia II with two ad seta at 1/3 and 2/3, one pd, one ad, one av and one ventral seta at apex. Relative podomere ratios: I: 12.6/ 10.1/ 6.1/ 3.2/ 1.7/ 1/ 1.2; II: 10.5/ 10.2/ 7.4/ 3.3/ 2/ 1.4/ 1; III: 12.9/ 14/ 4.9/ 4.6/ 2.4/ 1.5/ 1. Abdomen (Fig. 22 B–D). Tergites 1–5 with short vestiture of black setae. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “ BRAZIL, GO [Goiás], Varjão | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.55428&amp;materialsCitation.latitude=-17.032534" title="Search Plazi for locations around (long -49.55428/lat -17.032534)">Chácara [Farm] N. Sra. Aparecida</a> | Malaise - <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.55428&amp;materialsCitation.latitude=-17.032534" title="Search Plazi for locations around (long -49.55428/lat -17.032534)">Mata [Semideciduous Seasonal Forest]</a> | -17.032533, -49.554281 [17°01′57.1″S 49°33′15.4″W] | 27.vi.2021 - 08.viii.2021 | Col. Lopes W.R. ”; “HOLOTYPE | Mberu lopesi | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Goiás. Same data as holotype (3 ♂, 1 ♀, MNRJ; 1 ♂, 1 ♀, ZUFG). Mato Grosso do Sul. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-55.65578&amp;materialsCitation.latitude=-20.433054" title="Search Plazi for locations around (long -55.65578/lat -20.433054)">Aquidauana. Res. Ecol</a>. UEMS, 20º25′59″S 55º39′20.8″W, Floresta Estacional Decidual, malaise 4, 27.viii–11.ix.2011, Lamas, Nihei et al. Leg. (9 ♂, 15 ♀, MZUSP) .</p><p>Etymology. Named after the Brazilian dipterist Welinton Ribamar Lopes (in memoriam), who collected the type series of this species and was one of the professors who most helped the first author in his career at UFG.</p><p>Remarks. Mberu lopesi sp. nov. is distinguished from all other species of the pepocatu group by its small body size (about 2 mm while other species of the group have body size greater than 2.30 mm), vein M 1 S-shaped, parallel to R 4+5 at apex (Fig. 6F) (vein M 1 gently bent and converging to R 4+5 near wing apex in other species of group) and by lacking the ad row of setae on femur II and anterior on tibia II (present in other species of group) (Fig. 6E).</p><p>Distribution (Fig. 24). Brazil: states of Goiás and Mato Grosso do Sul (Cerrado biome).</p><p>Comments. Mberu lopesi sp. nov. is one of the morphospecies collected in Mato Grosso do Sul (Brazil) during the SISBIOTA-Diptera Brazilian Network (see Lamas et al. 2023).</p></div>	https://treatment.plazi.org/id/03B18E5CFF91FFC95AAD4CA6FC52F9C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF8EFFD55AAD4CEEFBDFFD7B.text	03B18E5CFF8EFFD55AAD4CEEFBDFFD7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu amorimi Silva & Capellari & Oliveira 2025	<div><p>Mberu amorimi sp. nov.</p><p>(Figs 8, 9)</p><p>Description. Male. Body length: 2.32–2.50 mm. Wing length: 2.10–2.44 mm long, 0.67–0.73 mm wide (n = 3). Body mostly brown, with slight blue or green reflections. Head (Fig. 8B, C). Frons width 1/3 head width; at narrowest part, slightly wider at ventral and dorsal ends; eyes separated on face, converging at middle, narrowest distance as broad as diameter of mid ocellus; face and clypeus with dense silver pruinosity; palpus brown, whitish on edges and apex, covered by 1–2 setulae on apex; proboscis light brown, with short whitish hairs; one proclinate, convergent vt s, one reclinate, slightly divergent, strong oc s, nearly two as long as vt s; pair of poc s, 0.5 times length of vt s; occiput distinctly concave above, shiny dark brown, lower surface of occiput covered by one complete row of white pocl s, increasing in size ventrally, ventralmost sparse and straight, as long as vt s; lower occiput covered by some white, straight ocp s, increasing in size ventrally, 1.3 times as long as lower pocl s. Scape short, yellow, dorsal surface bare; pedicel yellow, with crown of apical dark setae; postpedicel dark yellow, triangular, pubescent, about 1.5 times longer than basal width; stylus about 4.3 times longer than postpedicel, dorsally inserted, bare, displaced laterally, bi-articulated at base. Thorax (Fig. 8C). Brown, with some green or blue reflections, 0.7 times as long as abdomen, posterior slope of mesonotum distinctly flattened, postpronotal lobe with lower pale spot. Short scattered setae from anterior end of mesonotum to slightly beyond transverse suture; two rows of 13–15 small ac, ending at mesonotal slope, last two diverging; 6–7 dc, increasing in size posteriorly, posterior five conspicuously longer than anteriormost; one psut ial s, one presut spal s, one large psut spal s; two white pprn s, one strong and one tiny; two npl s; two pal s, one strong and one setula anteriad; one pair of strong l sctl s, one pair of smaller laterad setae, about half as long as l sctl s; two white prepst s. Wing (Fig 8A). Membrane hyaline, veins brown. C ending on M 1, before wing apex; Sc ending at basal third of R 1; R 1 ending at about basal third of wing; R 2+3 and R 4+5 strongly displaced towards anterior margin as compared to R 1, anterior cells strongly compressed, R 2+3 ending in C at apical fourth of wing; R 4+5 and M 1 convergent beyond dm-m, parallel at wing apex; M 4 complete, reaching wing margin; CuA+CuP strong, not reaching wing margin. Cilia of calypter pale, long. RMx ratio = 3.2, CuAx ratio = 0.3. Lower calypter and halter whitish. Legs. Completely yellow, except coxa II and III brown, sclerotized. Claws unmodified. Pulvilli small. Leg I. Coxa I (Fig. 8B, C) covered with white setae increasing in length towards apex, but smaller than ocp s; femur I with one ventral row of small white setae, posterior surface bare, except one white preapical seta; tibia I covered by small, fine setae; tarsus I with ventral surface entirely bare, dorsal surface covered by dark setae; tarsomere 1 smaller than remaining tarsomeres combined. Leg II. Coxa II with a few white setulae anteriorly, bare laterally; femur II covered by small setulae, ventral surface completely bare and one strong brown pv seta near apex; with ventral excavation at apex (all MSSC); tibia II with major setae at dorsal 1/2 and 3/4 and one apical seta dorsally; IIt1–4 with one av seta, one ventral and one pv small black seta at apex. Leg III. Coxa III with one conspicuous, long, white seta near base; femur III with one dorsal row of setae, decreasing in length towards apex and av row of 7–8 pale setae increasing in size towards apex, last four conspicuously longer than basalmost (MSSC) (e.g. Fig. 16D); tibia III with one row of dorsal pale setae and one ventral seta at apex; tarsomere 1 with one row of ventral setae, without ventral excavation at base. Relative podomere ratios: I: 6.1/ 6.8/ 3.4/ 2.2/ 1.6/ 1/ 0.8; II: 7.7/ 7.7/ 4.7/ 2.6/ 2/ 1.2/ 1; III: 6.9/ 8.1/ 2/ 2.5/ 1.7/ 1/ 1. Abdomen. Mostly light brown, with some dark blue to green reflections. Tergites 1–5 covered by short setae. Segment 6 partially hidden under tergite 5, tergite 6 bare; segment 7 forming short peduncle, tergite and sternite 7 bare; sternite 8 small, triangular. Sternites 1–3 visible, sclerotized, sternite 4 membranous. Hypopygium (Fig. 9). Epandrium elongate, twice longer than high, dark (Fig. 9B). MEp stout, approximately 2.5 times less than VSur height, with dorsal face rough, with 2 small dorsal setae near apex (Fig. 9C, F). DSur and VSur covering internal appendages (Fig. 9A); VSur (Fig. 9C) enlarged, twice higher than DSur, sclerotized at apex; DSur (Fig. 9D) elongated, twice wider than height. Hypandrium basally fused to epandrium. Phallus straight, without apical projection (Fig. 9G). Cercus (Fig. 9E) divided in two parts, ventral part curved and stout, with 3–4 spines ventrally and one long, anterior seta, dorsal part rounded, covered by fine setae, with two long setae apically. Female. Body length: 2.72–2.93 mm. Wing length: 2.6–2.8 long, 0.92–0.94 mm wide (n = 3). Habitus similar to male, except for MSSCs and as noted. Head. Face slightly broader (about as broad as ocellar tubercle), slightly wider towards apex, clypeal suture evident. Thorax. Two strong, white prepst s. Wing. RMx ratio = 4.5, CuAx ratio = 0.34. Legs. Leg I. Coxa I covered with white setae increasing in length towards apex; femur I without row of setae. Leg II. Tibia II with major setae at ad 1/4 and 1/2, one pv at 2/3, one av, one ventral and one pv dark seta at apex; It1–4 with one av, one ventral and one pv small black seta at apex. Relative podomere ratios: I: 6/ 6.9/ 3.5/ 2.3/ 1.6/ 1/ 1; II: 18.3/ 16.9/ 12.9/ 6.9/ 4.5/ 2.7/ 1; III: 10.8/ 11.5/ 3/ 4.1/ 2.5/ 1.7/ 1. Abdomen. Mostly light brown, with some dark blue to green reflections. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “ BRASIL, RS [Rio Grande do Sul], São Lourenço do Sul | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-31.4816&amp;materialsCitation.latitude=-31.481611" title="Search Plazi for locations around (long -31.4816/lat -31.481611)">Foz do Antônio Grande: Mata nas dunas próximo à Lagoa dos Patos</a> | Malaise - 31.x-8-xi.2011 | S31.4816 W52.00817 [31°28′53.8″S 52°00′29.4″W] | F.D. Kirst &amp; R.F. Kruger col.”; “HOLOTYPE | Mberu amorimi | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Rio Grande do Sul. Same data as holotype (2 ♂, one dissected, 3 ♀, MZUSP) .</p><p>Etymology. Named after the Brazilian entomologist Dalton de Souza Amorim, one of the authors of the genus Mberu and who provided the material for the study.</p><p>Remarks. Mberu amorimi sp. nov. is similar to M. britoi sp. nov. but can be differentiated by legs completely yellow in M. amorimi sp. nov. (Fig. 8A), and MEp stout, approximately 2.5 times less than VSur height, with dorsal face rugose (Fig. 9C, F). In M. britoi sp. nov., the legs are completely dark brown (Fig. 10A) and MEp is narrow, approximately 4.5 times less than VSur height, with dorsal face straight (Fig. 11C, E).</p><p>Distribution (Fig. 24). Brazil: state of Rio Grande do Sul (Pampa biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFF8EFFD55AAD4CEEFBDFFD7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF8CFFD15AAD4AABFA9CFDD6.text	03B18E5CFF8CFFD15AAD4AABFA9CFDD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu britoi Silva & Capellari & Oliveira 2025	<div><p>Mberu britoi sp. nov.</p><p>(Figs 10, 11)</p><p>Description. Male. Body length: 2.11–2.47 mm. Wing length: 2.11–2.33 mm long, 0.59–0.75 mm wide (n = 2). Similar to Mberu amorimi sp. nov. in both general habitus and chaetotaxy, except as noted. Body mostly metallic black. Head (Figs 10B, C). Palpus brown, yellowish on edges and apex; proboscis dark brown; postpedicel dark brown, triangular and pubescent, about 1.3 times longer than basal width; stylus about 5.2 times longer than postpedicel. Thorax (Fig. 10B). Metallic black, with some green or blue reflections. One presut spal s and two large psut spal s; two dark pprn s, one strong and one tiny. Wing. RMx ratio = 4.5, CuAx ratio = 0.4. Legs (Fig. 10A). Mostly brown; coxa I, II and III sclerotized. Yellow at apex of femur I, base of tibia I, apex of femur II and apex of femur III. Leg I. It1–5 with ventral surface covered by small dark setae. Leg II. Femur II covered by small setulae, ventral surface completely bare without pv; tibia II with major setae at ad 1/4 and 1/2 and one ad apical seta. Leg III. Coxa III with one conspicuous, long and dark seta near base; femur III with one av row of 3–4 setae decreasing in size towards apex. Relative podomere ratios: I: 6.9/ 7.3/ 3.8/ 2.6/ 1.6/ 1.3/ 1; II: 9.4/ 9.3/ 6.4/ 3.8/ 2.2/ 1.3/ 1; III: 8.7/ 10.4/ 2.9/ 3.5/ 2.1/ 1.3/ 1. Abdomen. Mostly black, with some dark blue to green reflections; tergite 2–5 with anterior shiny green bands laterally. Hypopygium (Fig. 11A–H). Epandrium elongate, twice longer than high, dark (Fig. 11A, B); MEp elongate, digitiform, with two small, dorsal setae near apex (Fig. 13C, E); DSur and VSur covering internal appendages (Fig. 11A, B); VSur (Fig. 11C) enlarged, twice higher than DSur, sclerotized at apex; DSur (Fig. 11D) elongated, twice wider than height. Hypandrium basally fused to epandrium. Phallus straight, without apical projection (Fig. 11G). Cercus (Fig. 11F) divided in two parts, ventral part stout, rounded, with 3–4 spines ventrally and one long anterior seta, dorsal part rounded and covered by fine setae, with one long, flattened seta apically. Female. Body length: 2.55–3.32 mm. Wing length: 2.52–3.24 mm long, 0.78–1.04 mm wide (n = 5). Similar to male, except as noted. Head. Face broader (about as broad as ocellar tubercle). Wing. RMx ratio = 2.7, CuAx ratio = 0.4. Legs. Leg II. Tibia II with av at 1/4, 1/2, 3/4, and at apex, one anterior seta at apex, two ad at 1/3 and 2/3, and two dorsal setae at 2/3 and at apex. Relative podomere ratios: I: 7.1/ 7.9/ 3.6/ 2.8/ 1.9/ 1.1/ 1; II: 6/ 7.2/ 4.5/ 2.8/ 1.8/ 1/ 1; III: 9.5/ 10.8/ 2.6/ 3.7/ 2.5/ 1.3/ 1. Abdomen. Tergites 1–5 covered by short setae. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “ BRAZIL, SP [São Paulo], Conchas, | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.102444&amp;materialsCitation.latitude=-22.983833" title="Search Plazi for locations around (long -48.102444/lat -22.983833)">Juquiratiba, Faz. Boa Vista.</a> | 22º59′1.8″ S 48º6′8.8″ W | Malaise. Pomar. [Orchard] | 17-19.I.2022, Capellari leg.”; “HOLOTYPE | Mberu britoi | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia dissected and stored in microvial with glycerin, pinned beneath the specimen. PARATYPES. Brazil. São Paulo. Same data as holotype (4 ♀, MZUSP; 1 ♀, ZUFG) . Goiás. Varjão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.55483&amp;materialsCitation.latitude=-17.031122" title="Search Plazi for locations around (long -49.55483/lat -17.031122)">Chácara Nossa Sra.Aparecida</a>, Malaise – Pomar, 17º01′52.04″S 49º33′17.39″W, 12.vi.– 08.vii.2018 Col. Lopes, W.R. (1 ♂, MZUSP) .</p><p>Etymology. Named after Dr. Pedro Vale de Azevedo Brito (UFG), an attentive professor and advisor of the first author during the past years.</p><p>Remarks. Mberu britoi sp. nov. is similar to M. amorimi sp. nov. but can be differentiated by legs mostly dark brown (Fig. 10A), and medial epandrial lobe narrow, approximately 4.5 times less than VSur height, with dorsal face straight (Fig. 11C, E). In M. amorimi sp. nov., the legs are completely yellow (Fig. 8A) and MEp stout, approximately 2.5 times less than VSur height, with dorsal face rough (Fig. 9C, F).</p><p>Distribution (Fig. 24). Brazil: states of Goiás (Cerrado biome) and São Paulo (Atlantic Forest Biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFF8CFFD15AAD4AABFA9CFDD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF89FFDF5AAD4E96FC8CFEAB.text	03B18E5CFF89FFDF5AAD4E96FC8CFEAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu lamasi Silva & Capellari & Oliveira 2025	<div><p>Mberu lamasi sp. nov.</p><p>(Figs 12, 13)</p><p>Description. Male. Body length: 2.66–2.87 mm. Wing length: 2.60–2.74 mm long, 0.70–0.84 mm wide (n = 10) (Fig. 12A). Similar to Mberu amorimi sp. nov. in both general habitus and chaetotaxy, except as noted. Body mostly black to dark brown, with slight blue or green reflections. Head (Fig. 12B). Frons width 1/3 head width; vertex slightly excavated dorsally; face with dense silver pruinosity; clypeus dark brown, with silver pruinosity on edges; palpus dark brown, yellowish on edges and apex; postpedicel brown, triangular and pubescent, about 1.4 times longer than basal width; stylus about 4.1 times longer than postpedicel. Thorax. Dark brown, with some green or blue reflections, as long as abdomen; 4–5 dc s, increasing in size posteriorly; one presut spal s and two large psut spal s; two dark pprn s, one strong and one tiny. Wing. RMx ratio = 4.5, CuAx ratio = 0.3. Legs. Completely brown, coxa II and III sclerotized. Leg I. Coxa I with dorsal hook-shaped projection at apex (Fig. 12C) (MSSC); femur I with ventral excavation at apex; tarsomere 1 elongated, as long as remaining tarsomeres combined, with 5–6 small dark av, similar to spines (Fig. 12D). Leg II. Femur II with row of dark setae on anterior surface; ventral surface bare, except av row of dark setae on apical half and one strong, brown pv seta near apex; with ventral excavation at apex (all MSSC). Leg III. Femur III with apical pv and ventral excavation at apex; tibia III (Fig. 12E) with dorsal swelling at apical half, row of dorsal and ventral white setae and comb of posterior pale setae projecting into IIIt1; IIIt1 with comb of posterior white setae projecting into IIIt2, with ventral excavation at base (MSSC). Relative podomere ratios: I: 8.7/ 8.6/ 4.8/ 2.3/ 1.6/ 1.3/ 1; II: 8.6/ 8/ 5.7/ 3.1/ 2.1/ 1.3/ 1; III: 7.9/ 8.1/ 2.8/ 3/ 2/ 1.3/ 1. Abdomen. Mostly black to dark brown, with some dark blue to green reflections. Hypopygium (Fig. 13). Epandrium elongate, twice longer than high, dark (Fig. 13A, B). MEp digitiform, with two small dorsal setae near apex (Fig. 13C, E). DSur and VSur covering internal appendages (Fig. 13A, B); VSur (Fig. 13C) enlarged, twice higher than DSur, with one internal seta at apex, probably being LEp fused with VSur (see Quevedo et al., 2025); DSur (Fig. 13D) irregularly shaped and robust, with curved ventral margins and truncated, rounded distal lobe, 3.2 times wider than height. Hypandrium basally fused to epandrium. Phallus with short dorsal projection near apex (Fig. 13G). Cercus (Fig. 13F) divided in two parts, ventral part rounded and stout, with 4–5 spines ventrally, dorsal part rounded and covered by fine setae, with flattened seta apically. Female. Body length: 2.83–2.90 mm. Wing length: 2.89 mm long, 0.95–0.97 mm wide (n = 5). Similar to male, except as noted. Head. Face broader (about as broad as ocellar tubercle). Wing. RMx ratio = 5.3, CuAx ratio = 0.3. Legs. Leg II. Tibia II with av at 1/4, 1/2, 3/4 and at apex, one anterior seta at apex, two ad at 1/3 and 2/3, and two dorsal setae at 2/3 and at apex. Relative podomere ratios: I: 8.6/ 8.3/ 4.8/ 3/ 1.6/ 1/ 1.1; II: 13.6/ 13.2/ 8.9/ 4.9/ 3.3/ 2/ 1; III: 9.4/ 11.1/ 3.2/ 4.1/ 2.3/ 1.5/ 1. Abdomen. Tergites 1–5 covered by short setae. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “SISBIOTA - | CNPQ/FAPESP [Vertical line] | BRASIL, MS [Mato Grosso do Sul], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.880386&amp;materialsCitation.latitude=-20.69886" title="Search Plazi for locations around (long -56.880386/lat -20.69886)">Bodoquena</a> | 20º41′55.9″S 56º52′49.4″W | Malaise 6, 5-20.ii.2012, | Lamas, Nihei et al. leg.”; “HOLOTYPE | Mberu lamasi | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Mato Grosso do Sul. Same data as holotype (6 ♂, 4 ♀, MZUSP) . Same data, 21.iii–06.iv.2012 (1 ♀, MZUSP) . Same data except, 20º41′49.9″S 56º52′54″W, Malaise 4 - 6–21.i.2012 (6 ♂, one dissected, 2 ♀, MZUSP) . Rio Grande do Sul. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.139503&amp;materialsCitation.latitude=-31.56436" title="Search Plazi for locations around (long -52.139503/lat -31.56436)">Arroio Corrientes, Fazenda Corrientes, Paleodunas</a>, Malaise - 29.x– 6.xi.2011, 31°33′51.7″S 52°08′22.2″W, F.D. Kirst &amp; R.F. Kruger col. (1 ♂, INPA) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.254223&amp;materialsCitation.latitude=-31.72175" title="Search Plazi for locations around (long -52.254223/lat -31.72175)">Arroio Pelotas</a>, 27.x–04.xi.2011, Malaise Areal, 31°43′18.3″S 52°15′15.2″W, F.D. Kirst &amp; R.F. Kruger col. (1 ♂, dissected, MZUSP) . Paraguay. Caaguazú. Coronel Oviedo, 1971, L Peña leg. (2 ♂, one dissected, MZUSP) .</p><p>Additional material. Argentina. Corrientes. Corrientes City, 27°25′26.6″S 58°43′55.4″W, 09.xi.2022, Matias Ignacio Dufek col., hand net, Universidad Nacional del Nordeste (1♂, 1♀, UNNE) .</p><p>Etymology. Named after the Brazilian dipterist Carlos Lamas (MZUSP), one of the collectors of the type series of this species.</p><p>Remarks. Mberu lamasi sp. nov. is similar to M. soaresi sp. nov., both have tibia III with dorsal swelling on apical half, comb of posterior pale setae projecting into IIIt1, projecting towards tarsal excavation (Figs 12E, 18E), and the hypopygial morphology is similar to the pepocatu group, but can be differentiated by coxa I with conspicuous hook-shaped projection in M. lamasi sp. nov. (Fig. 12C) and It1 with distinct row of setae similar to spines (Fig. 12D) (coxa I without conspicuous projection (Fig. 18C) and It1 without distinct row of setae (Fig. 18D) in M. soaresi sp. nov.).</p><p>Matias Dufek (pers. comm.) reported that he collected this species emerging from a cactus in a region of the Argentinian Chaco, and the identification was confirmed with photos sent by him. Parent (1933) described a female from San José (Northern Bolivia) as an uncertain genus (“Genre?”) suggesting it would be placed in Medeterinae . Photos of the specimen (currently in the Staatliches Museum für Naturkunde Stuttgart, Germany) proved that it belongs to Mberu and is probably conspecific with M. lamasi sp. nov.</p><p>Distribution (Fig. 24). Argentina: Corrientes; Paraguay: Coronel Oviedo (Chaco biome); Brazil: states of Mato Grosso do Sul (Cerrado biome) and Rio Grande do Sul (Pampa biome).</p><p>Comments. Mberu lamasi sp. nov. is one of the morphospecies collected in Mato Grosso do Sul (Brazil) during SISBIOTA-Diptera Brazilian Network (see Lamas et al. 2023).</p></div>	https://treatment.plazi.org/id/03B18E5CFF89FFDF5AAD4E96FC8CFEAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF87FFDC5AAD4B37FC7CFF4F.text	03B18E5CFF87FFDC5AAD4B37FC7CFF4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu murtiniensis Silva & Capellari & Oliveira 2025	<div><p>Mberu murtiniensis sp. nov.</p><p>(Figs 14, 15)</p><p>Description. Male. Body length: 1.57 mm. Wing length: 1.70 mm long, 0.52 mm wide (n = 1). Similar to M. amorimi sp. nov. in both general habitus and chaetotaxy, except as noted. Head (Fig. 14B, C). Eyes touching on face; region of face anterior to antenna brown, lower surface of face and clypeus with slight silver pruinosity. Postpedicel dark yellow, triangular, pointed and pubescent; about 1.6 times longer than basal width; stylus about 2.5 times longer than postpedicel. Thorax (Fig. 14C). Brown, except for black spot below wing. 4–5 dc s, increasing in size posteriorly, posteriormost three conspicuously longer than anteriormost; one brown pprn s. Wing. R 1 ending near middle of wing; RMx ratio = 2.3, CuAx ratio = 0.2. Legs. Completely yellow, including coxae II and III. Leg I. Femur I bare of major setae; dorsal surface of tarsus I covered by white setae. Leg II. Femur II with row of small ventral setae, without major setae; without ventral excavation; tibia II without major setae at dorsal 1/2 and 3/4, but with one av, one ventral, one pv and one dorsal small black seta at apex. Leg III. Femur III without conspicuous row of dorsal setae, av row with 5–6 white setae; tibia III with one av and one ad white seta at apex; IIIt1 with row of ventral setae, with ventral excavation at base. Relative podomere ratios: I: 7.3/ 6/ 3.9/ 2.4/ 1.5/ 1/ 1; II: 7.6/ 7.3/ 4.5/ 2.6/ 1.7/ 1/ 1; III: 7/ 7.9/ 2.6/ 3.1/ 1.7/?/?. Abdomen. As in M. amorimi sp. nov. Hypopygium (Fig. 15). Epandrium elongate, twice longer than high, dark. MEp narrow, branched, without setae at apex (Fig. 15C, G). DSur and VSur covering internal appendages (Fig. 15A, B); VSur (Fig. 15C) enlarged, twice higher than DSur, with one internal seta at apex, probably being LEp fused with VSur (see Quevedo et al. 2025); DSur (Fig. 15D) with proximal and distal widths subequal, dorsal margin slightly concave, twice wider than height, with ventral projection near apex. Hypandrium basally fused to epandrium. Phallus (lost during dissection) straight, without apical projection. Cercus (Fig. 15F) divided in two parts, ventral part curved and stout, with 3–4 spines ventrally, dorsal part rounded and covered by fine setae, with one short seta apically. Female. Unknown.</p><p>Type material. HOLOTYPE ♂, “SISBIOTA - | CNPQ/FAPESP [Vertical line] | BRAZIL, MS [Mato Grosso do Sul], Porto Murtinho | Faz. Retiro Conceição - Trilha | da Espinhadeira | S21°40′59.7″ W57°46′42.5″ | Malaise 31 | 18.iv-15.v.2012 | Lamas, Nihei &amp; eq. Col.”; “HOLOTYPE | Mberu murtiniensis | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia dissected and stored in microvial with glycerin, pinned beneath the specimen.</p><p>Etymology. Toponymic, referring to the type-locality.</p><p>Remarks. Mberu murtiniensis sp. nov. is similar to M. takwajasu sp. nov., and both share the eyes touching on face (Figs 14B, 20B) and MEp branched (Figs 14G, 21H). Mberu murtiniensis sp. nov. is distinguished by its reduced body length (about 1.60 mm), postpedicel about 1.6 times longer than its basal width (Fig. 14C), DSur slight concave dorsally (Fig. 15D), and hypopygium not elongated, reaching forward abdominal segment 4 (Fig. 15A). In M. takwajasu sp. nov., the body length is longer than 2 mm, postpedicel about 2.3 times longer than its basal width (Fig. 20C), DSur subtriangular (Fig. 21D) and hypopygium elongated, reaching abdominal segment 2 (Fig. 21A).</p><p>Distribution (Fig. 24). Brazil: state of Mato Grosso do Sul (Chaco biome).</p><p>Comments. Mberu murtiniensis sp. nov. is one of the morphospecies collected in Mato Grosso do Sul (Brazil) during SISBIOTA-Diptera Brazilian Network (see Lamas et al. 2023).</p></div>	https://treatment.plazi.org/id/03B18E5CFF87FFDC5AAD4B37FC7CFF4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF84FFDA5AAD4A1BFC61FE43.text	03B18E5CFF84FFDA5AAD4A1BFC61FE43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu periotoi Silva & Capellari & Oliveira 2025	<div><p>Mberu periotoi sp. nov.</p><p>(Figs 16, 17)</p><p>Description. Male. Body length: 2.21–2.28 mm. Wing length: 2.02–2.19 mm long, 0.68–0.72 mm wide (n = 3). Similar to M. amorimi sp. nov. in both general habitus and chaetotaxy, except as noted. Head (Fig. 16B, C). Pair of reclinate, slightly divergent, strong oc s, almost 1.5 as long as vt s; pair of proclinate, convergent vt s; postpedicel brown, about 1.2 times longer than basal width; stylus about 3.9 times longer than postpedicel; face with dense silver pruinosity. Thorax (Fig. 16C). As in M. amorimi sp. nov. Wing. RMx ratio = 2.6, CuAx ratio = 0.3. Legs. Leg I. It1 smaller than remaining tarsomeres combined. Leg II. Tibia II with major setae at dorsal 1/4 and 1/2, one dorsal, one ventral, and one anterior apical seta; posterior surface of IIt1 with row of 4–5 setae. Relative podomere ratios: I: 6.7/ 7.1/ 3.6/ 2.1/ 1.5/ 1/ 0.9; II: 8/ 8/ 5/ 2.8/ 1.8/ 1/ 1; III: 6.9/ 8.4/ 2.1/ 2.4/ 1.6/ 1/ 1. Abdomen. As in Mberu amorimi sp. nov. Hypopygium (Fig. 17). Epandrium elongate, twice longer than high, dark (Fig. 17A, B). MEp digitiform, ventrally swollen, approximately four times less than VSur height, with dorsal face straight, (Fig. 17C). DSur and VSur covering internal appendages (Fig. 17A, B); VSur (Fig. 17C) enlarged, twice higher than DSur, sclerotized at apex, with three apical setae, probably being LEp fused with VSur (see Quevedo et al. 2025); DSur (Fig. 17D) elongated and curved, apically rounded, three times wider than height, conspicuously projecting beyond VSur. Hypandrium basally fused to epandrium. Phallus straight, without apical projection (Fig. 17F). Postgonite absent. Cercus (Fig. 17E) divided in two parts, ventral part curved and stout, with 6–7 spines ventrally and 1 long and flattened anterior seta, dorsal part rounded, covered by fine setae. Female. Body length: 2.28–2.53 mm. Wing length: 2.30–2.46 mm long, 0.74–0.82 mm wide (n = 5). Similar to male, except as noted. Head. Face broader (about as broad as ocellar tubercle). Thorax. Three strong white prepst s. Wing. RMx ratio = 4.3, CuAx ratio = 0.3. Legs. Leg II. Tibia II with major setae at dorsal 1/3 and 2/3, and av at 2/3; IIt1–4 with one av, one ventral and one pv small black seta at apex; posterior surface of IIt1 without row of setae. Relative podomere ratios: I: 6.8/ 7.3/ 3.4/ 2.4/ 1.5/ 1/ 1; II: 8.8/ 8.7/ 5.5/ 3/ 1.8/ 1/ 1; III: 8.5/ 9.4/ 2.6/ 3.4/ 1.9/ 1.3/ 1. Abdomen. Tergites 1–5 covered by short setae. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “ BRASIL, SP [São Paulo], Luiz Antônio, | E.E. [Ecological Station] Jataí, Cerrado [Savanna], malaise I | 07.vii.2006, Perioto, N. col.”; “HOLOTYPE | Mberu periotoi | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. São Paulo. Same data as holotype (1 ♂, dissected, 2 ♀, one dissected, MZUSP). Conchas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.102444&amp;materialsCitation.latitude=-22.983833" title="Search Plazi for locations around (long -48.102444/lat -22.983833)">Juquiratiba</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-48.102444&amp;materialsCitation.latitude=-22.983833" title="Search Plazi for locations around (long -48.102444/lat -22.983833)">Faz. Boa Vista</a>, 22°59′1.8″S 48°6′8.8″W, Malaise. Pomar, 17–19.i.2022, Capellari leg.” (1 ♂, dissected, 3 ♀, MZUSP) .</p><p>Etymology. Named after Brazilian entomologist Nelson Perioto, who collected the holotype of the species.</p><p>Remarks. Mberu periotoi sp. nov. is similar to M. amorimi sp. nov. but can be easily separated by DSur elongated and curved, apically rounded, three times wider than height, conspicuously projecting beyond the VSur (Fig. 17A, B, D). In M. amorimi sp. nov., the size of both surstyli are similar (Fig. 9A, B, D).</p><p>Distribution (Fig. 24). Brazil: state of São Paulo (Cerrado biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFF84FFDA5AAD4A1BFC61FE43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFF81FFD95AAD4CA6FE24F8C7.text	03B18E5CFF81FFD95AAD4CA6FE24F8C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu soaresi Silva & Capellari & Oliveira 2025	<div><p>Mberu soaresi sp. nov.</p><p>(Figs 18, 19)</p><p>Description. Male. (Fig. 18A) Body length: 2.49–2.94 mm. Wing length: 2.76–3.01 mm long, 0.84–0.95 mm wide (n = 9). Similar to Mberu amorimi sp. nov. in both general habitus and chaetotaxy, except as noted. Body mostly black to dark brown, with slight blue or green reflections. Head (Fig. 18B). Frons width 1/3 head width; vertex slightly excavated dorsally; face with dense silver pruinosity; clypeus dark brown, with silver pruinosity on edges; palpus dark brown, yellowish on edges and apex; postpedicel brown, triangular and pubescent, about 1.9 times longer than basal width; stylus about 2.6 times longer than postpedicel. Thorax. Brown, with some green or blue reflections, 1.1 times as long as abdomen; 4–5 dc s, increasing in size posteriorly; one presut spal s and two large psut spal s; two dark pprn s, one strong and one tiny. Wing. RMx ratio = 3.3, CuAx ratio = 0.3. Legs. Completely brown, coxae II and III sclerotized. Leg I. Coxa I without conspicuous projection (Fig. 18C); femur I with ventral excavation at apex; It1elongated, as long as remaining tarsomeres combined (Fig. 18D). Leg II. Femur II with one row of dark setae on anterior surface; ventral surface bare, except for av row of dark setae on apical half and one strong brown pv near apex; with ventral excavation at apex (all MSSC). Leg III. Femur III with apical pv and ventral excavation at apex; tibia III (Fig. 18E) with dorsal swelling at apical half, row of dorsal and ventral white setae, and comb of posterior white setae projecting into IIIt1, projecting towards tarsal excavation; IIIt1 with comb of posterior white setae projecting into IIIt2, with ventral excavation at base (MSSC). Relative podomere ratios: I: 8.2/ 8.5/ 4.9/ 2.1/ 1.5/ 1/ 1.2; II: 8.4/ 8.2/ 6/ 3.3/ 2.1/ 2.1/ 1; III: 7.7/ 8.4/ 2.8/ 2.9/ 1.7/ 1.2/ 1. Abdomen. Mostly black to dark brown, with some dark blue to green reflections. Hypopygium (Fig. 19). Epandrium elongate, twice longer than high, dark (Figs 19A, B). MEp elongate, digitiform, with two small dorsal setae near apex (Fig. 19C). DSur and VSur covering internal appendages (Fig. 19A, B); VSur (Fig. 19C) enlarged, twice higher than DSur, with one internal seta at apex, probably being LEp fused with VSur (see Quevedo et al. 2025); DSur (Fig. 19D) irregularly shaped and robust, with curved ventral margins and truncated, rounded distal lobe, 2.1 times wider than height. Hypandrium basally fused to epandrium. Phallus with short dorsal projection near apex (Fig. 19F). Cercus (Fig. 19E) divided in two parts, ventral part rounded and stout, with 4–5 spines ventrally, dorsal part rounded and covered by fine setae, with flattened seta apically. Female. Body length: 3.09 mm. Wing length: 2.99 mm long, 1.13 mm wide (n = 1). Similar to male, except as noted. Head. Face broader (about as broad as ocellar tubercle). Wing. RMx ratio = 4.6, CuAx ratio = 0.4. Legs. Leg II. Tibia II with av at 1/4, 1/2, 3/4 and at apex, one anterior seta at apex, two ad at 1/3 and 2/3, and two dorsal setae at 2/3 and at apex. Relative podomere ratios: I: 7.7/ 7.5/ 4.6/ 2.3/ 1.4/ 1/ 1; II: 9.5/ 8/ 6.2/ 3.3/ 1.9/ 1.2/ 1; III: 9/ 10.6/ 3.5/ 3.9/ 2.2/ 1.4/ 1. Abdomen. Tergite 1–5 covered by short setae. Oviscapt as in M. bodoquena sp. nov. .</p><p>Type material. HOLOTYPE ♂, “ BRASIL, RS [Rio Grande do Sul], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.143364&amp;materialsCitation.latitude=-31.555056" title="Search Plazi for locations around (long -52.143364/lat -31.555056)">Arroio [Stream] Corrientes</a>, | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.143364&amp;materialsCitation.latitude=-31.555056" title="Search Plazi for locations around (long -52.143364/lat -31.555056)">Fazenda [Farm] Corrientes – Mata</a> | Malaise – 28.x-5.xi.2011 | S31.55506 W52.14336 [31°33′18.2″S 52°08′36.1″W] | F.D. Kirst &amp; R.F. Kruger col.; “HOLOTYPE | Mberu soaresi | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia not dissected. PARATYPES. Brazil. Rio Grande do Sul. Same data as holotype (4 ♂, two dissected, MZUSP) . Santa Catarina. Nova Teutônia, x.1970, 27°11′S 52°23′W, Fritz Plaumann Col. (2 ♂, one dissected, MZUSP) . Paraná. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.132423&amp;materialsCitation.latitude=-25.37942" title="Search Plazi for locations around (long -49.132423/lat -25.37942)">Bairro Santa Rita</a>, 25°22′45.91″S 49°07′56.73″W, Malaise – 913 m, iv.2012, Savaris, M. col. (1 ♂, 1 ♀, MZUSP) . São Paulo. iv.1960, Forattini col. (1 ♂, MZUSP) .</p><p>Etymology. Named after Matheus Mickael Mota Soares (INPA), a prominent Brazilian dipterist specializing in Brachycera and a good friend who helped the first author during his stay at INPA.</p><p>Remarks. Mberu soaresi sp. nov. is similar to M. lamasi sp. nov., and both have tibia III with dorsal swelling at apical half, comb of posterior white setae projecting into IIIt1, projecting towards tarsal excavation (Figs 12E, 18E), and the hypopygial morphology similar to the pepocatu group, but can be differentiated by coxa I without conspicuous projection in M. soaresi sp. nov. (Fig. 18C), and It1 without modified setae (Fig. 18D). Mberu lamasi sp. nov. has coxa I with conspicuous posterior hook-shaped projection (Fig. 12C), and It1 with distinct row of spine-like setae (Fig. 12D).</p><p>Distribution (Fig. 24). Brazil: states of Paraná, São Paulo, Santa Catarina (Atlantic Forest biome), and Rio Grande do Sul (Pampa biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFF81FFD95AAD4CA6FE24F8C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
03B18E5CFFBFFFE75AAD4CA6FC51F80F.text	03B18E5CFFBFFFE75AAD4CA6FC51F80F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mberu takwajasu Silva & Capellari & Oliveira 2025	<div><p>Mberu takwajasu sp. nov.</p><p>(Figs 20, 21, 22A)</p><p>Description. Male. Body length: 2.20 mm. Wing length: 2.15 mm long, 0.73 mm wide (n = 1). Similar to M. amorimi sp. nov. in both general habitus and chaetotaxy, except as noted. Body mostly brown, with slight blue reflections. Head (Fig. 20B, C). Eyes touching on face, forming triangle at upper and lower face; palpus yellow, with one brown seta on apex; pair of tiny poc s. Postpedicel dark brown, triangular and pointed, covered by many pale setulae, elongate, about 2.3 times longer than basal width; stylus about two times longer than postpedicel. Thorax (Fig. 20C). Dark brown, with some green or blue reflections, 1.7 times as long as abdomen. 6–7 dc s, increasing in size posteriorly, posteriormost three conspicuously longer than anteriormost; three pprn s, one strong dark seta and two white setulae below; one npl s, one large pal s; one pair of strong l sctl s, laterad setae absent. Wing. Sc ending at middle of R 1; R 1 ending at about basal third of wing; R 2+3 ending in C at apical fourth of wing. Lower calypter white, long, increasing in size towards apex. RMx ratio = 2.2, CuAx ratio = 0.2. Legs. Completely yellow, except coxa I with anterior surface light brown, coxae II and III light brown, sclerotized. Leg I. Femur I with av row of small white setae; one dark posterior preapical seta; It1 with ventral surface covered by short dark setae less dense than dorsal region, dorsal surface covered by dark setae; It1 as long as remaining tarsomeres combined, with row of small av and pd setae on apical third. Leg II. Coxa II with a few pale setae anteriorly, decreasing in size towards apex, bare laterally; femur II with ventral surface with sparse dark setulae, row of pv setae on apical fourth and one strong brown pv seta near apex; It1–4 without conspicuous setae at apex. Leg III. Femur III with av row of 10–12 white setae increasing in size towards apex; tibia III with apical comb of small pale setae projecting towards tarsal excavation; IIIt1–2 with row of ventral setae, IIIt1 with ventral excavation at base. Relative podomere ratios: I: 6.1/ 6.9/ 3.6/ 1.4/ 1.3/ 1.2/ 1; II: 7.7/ 8.1/ 2.6/ 2.6/ 1.7/ 1.1/ 1; III: 7.4/ 10/ 2.7/ 3.1/ 1.9/ 1.2/ 1. Abdomen. Mostly brown, lighter than thorax. Segment 6 not hidden under Tergite 5; sternite 8 enlarged, globular. Hypopygium (Fig. 21). Epandrium elongate, three times longer than high, dark; MEp narrow, about six times wider than height, dorsally branched, without setae at apex (Fig. 21C, H); DSur and VSur covering internal appendages (Fig. 21A, B); VSur (Fig. 21C) narrow, as high as DSur, four times wider than high, with apicoventral projection and one seta at apex (probably being LEp fused with VSur (see Quevedo et al. 2025); DSur (Fig. 21D) narrow, about four times wider than high, subtriangular, increasing in height near apex. Hypandrium basally fused to epandrium. Phallus (Fig. 21E) straight, without apical projection. Postgonite (Fig. 21G) slender, slight curved near apex. Cercus (Fig. 21F) elongate, about three times wider than height, divided in two parts, ventral part elongate, with slender, pointed branch dorsally, rounded at apex, with one apicodorsal seta, dorsal part elongate, base covered by fine setae and apex narrow. Female (Fig. 22A). Body length: 2.39–2.42 mm. Wing length: 2.13–2.59 mm long, 0.83–0.91 mm wide (n = 3). Similar to male except as noted. Head. Eyes not touching on face; postpedicel brown, triangular, pointed, covered by many pale setulae. Wing. RMx ratio = 2.1, CuAx ratio = 0.2. Legs. Leg I. Femur I without posterior preapical seta; Leg II. Femur II without row of pv; IIt1–4 with one av, one ventral and one pv small black seta at apex. Leg III. Femur III without anteroventral row of setae; tibia III without comb of setae; IIIt1 without ventral excavation. Relative podomere ratios: I: 9.3/ 10.4/ 6.9/ 3.3/ 2.2/ 1.3/ 1; II: 7.5/ 7.9/ 4.4/ 2.5/ 1.7/ 1.2/ 1; III: 9/ 11.1/ 2.9/ 3.2/ 1.7/ 1.3/ 1. Abdomen. Tergites 1–5 covered by short setae. Oviscapt as in M. bodoquena sp. nov.</p><p>Type material. HOLOTYPE ♂, “ BRAZIL, GO [Goiás], Mineiros | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.99641&amp;materialsCitation.latitude=-17.902225" title="Search Plazi for locations around (long -52.99641/lat -17.902225)">Parque Nacional [National Park] das Emas</a> | Malaise – Parcela [Point] 2 Mata [<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.99641&amp;materialsCitation.latitude=-17.902225" title="Search Plazi for locations around (long -52.99641/lat -17.902225)">Semideciduous Seasonal Forest</a>] | -17.9022253779 -52.9964093068 [17°54′08.0″S 52°59′47.1″W] | 12.vii.2017 - 16.viii.2017 | Cols. Oliveira. S.S. &amp; Lopes, W.R. ”; “HOLOTYPE | Mberu takwajasu | Silva, Capellari &amp; Oliveira [red label]” (MZUSP) . Holotype in good condition, genitalia dissected and stored in microvial with glycerin, pinned beneath the specimen. PARATYPES. Brazil. Goiás. Same data as holotype (3 ♀, MZUSP) .</p><p>Etymology. From the Tupi-Guarani, one of the native Brazilian linguistic families, “ takwãj ” (penis) + “ asu” (intensive), referring to the large size of the male genital capsule in relation to the other species of the genus.</p><p>Remarks. Mberu takwajasu sp. nov. is easily distinguished from other Mberu by having an elongate postpedicel, about 2.3 times longer than its basal width (Fig. 20B, C) (usually equal to 1.3 times longer in other species), and by the large size of the male hypopygium, reaching forward to abdominal segment 2 (Fig. 21A) (usually reaching abdominal segment 4 in other species).</p><p>Distribution (Fig. 24). Brazil: state of Goiás (Cerrado biome).</p></div>	https://treatment.plazi.org/id/03B18E5CFFBFFFE75AAD4CA6FC51F80F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Silva, Paulo Cesar;Capellari, Renato S.;Oliveira, Sarah S.	Silva, Paulo Cesar, Capellari, Renato S., Oliveira, Sarah S. (2025): Revision and morphological phylogeny of the Neotropical genus Mberu Capellari & Amorim (Diptera: Dolichopodidae), with description of twelve new species. Zootaxa 5637 (3): 469-514, DOI: 10.11646/zootaxa.5637.3.3, URL: https://doi.org/10.11646/zootaxa.5637.3.3
