taxonID	type	description	language	source
03BE3C58FFA3C0450D87FB6031A4FE34.taxon	description	(Figures. 4, 8)	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA3C0450D87FB6031A4FE34.taxon	materials_examined	Type material examined. Neotype (herein designated): male (CL 26.60 mm, CW 40.63 mm) (CNCR 35390), El Ojito Spring, pool in San Luis Apizaquito, municipality of Apizaco, 19 ° 26 ’ 16.28 ” N, 98 ° 07 ’ 00.71 ” W, 2,418 m asl, coll. E. Moreno & Y. Montiel, 17 August 2019. Other material examined. Mexico –– Puebla. 2 males (CL 19.41 – 25.73 mm, CW 28.74 – 38.41 mm), 2 females (CL 16.92 mm, CW) 24.80 mm) (CNCR 355), small dam in Chignahuapan, municipality of Chignahuapan, 19 ° 50 ’ 18.16 ” N 98 ° 01 ’ 52.35 ” W, 2256 m asl, coll. A. Villalobos-Figueroa, May 1994; 5 females (CL 22.02 – 26.10 mm, CW 33.63 – 40.59 mm) (CNCR 13409), San Isidro River, municipality of Zacatlán de las Manzanas, 19 ° 56 ‘ 06.29 ” N 97 ° 57 ‘ 40.15 ” W, 2047 m asl, coll. H. Brailovsky & E. Barrera, 18 July 1991; 7 males (CL 12.35 – 20.69 mm, CW 17.84 – 30.75 mm), 3 females (CL 16.28 – 26.40 mm, CW 24.27 – 40.68 mm) (CNCR 25527), Ajajalpa River in the Quetzalapan cascades, municipality of Zacatlán de las Manzanas, 19 ° 52 ’ 22.44 ” N 97 ° 58 ’ 53.98 ” W, 2193 m asl, coll. J. L. Villalobos & J. Segura, 15 October 2000; 1 male (CL 22.36 mm, CW 36.45 mm), 1 female (CL 16.82 mm, CW 29.06 mm) (CNCR 26108), stream between Tizcatlacoyan and San Antonio Juárez, municipality of Tizcatlacoyan, 18 ° 50 ’ 21.53 ” N 98 ° 02 ’ 43.89 ” W, 1998 m asl, coll. M. Olson & A. Páez-García, 20 June 2010; 1 male (CL 13.56 mm, CW 20.26 mm) (CNCR 31700), stream in La Magdalena Cuauxixtla, Tecali de Herrera, 18 ° 52 ’ 44.43 ” N 97 ° 57 ’ 23.65 ” W, 2068 m asl, coll. R. Hernández-Austria, 15 November 2019; 5 males (CL 22.43 – 27.29 mm, CW 32.63 – 41.16 mm), 2 females (CL 21.67 – 26.45 mm, CW 31.68 – 40.96 mm) (CNCR 36154), Quetzalapan cascade, municipality of Zacatlán de las Manzanas, 19 ° 52 ’ 18.70 ” N 97 ° 58 ’ 53.89 ” W, 2193 m asl, coll. K. J. Madrigal, E. Moreno & J. L. Villalobos, 11 November 2021; 1 female (CL 17.02 mm, CW 25.36 mm) (CNCR 36157), spring in Michac, municipality of Chignahuapan, 19 ° 51 ’ 3.00 ” N 98 ° 3 ’ 13.00 ” W, 2295 m asl, coll. E. Moreno, K. J. Madrigal & J. L. Villalobos, 12 November 2021; 4 males (CL 18.92 – 22.32 mm, CW 31.92 – 37.07 mm), 2 females (CL 15.05 – 18.31 mm, CW 25.60 – 32.40 mm) (CNCR 36550), stream in Cuahutinchán, municipality of Cuahutinchán, 18 ° 57 ’ 23.00 ” N 98 ° 01 ’ 53.00 ” W, 2128 m asl, coll. K. J. Madrigal, J. L. Villalobos & J. Santillán, 29 May 2022. –– Tlaxcala. 1 male (CL 15.03 mm, CW 24.78 mm) (CNCR 10460), spring at Apizaquito, municipality of Apizaco, 19 ° 26 ’ 16.28 ” N 98 ° 07 ’ 00.71 ” W, 2418 m asl, coll. E. Bonilla, 2 December 1987; 2 males (CL 3.95 – 11.11 mm, CW 7.48 – 19.06 mm) (CNCR 33848), stream in La Ascensión Huitzcolotepec, municipality of Xaltocan, 19 ° 22 ’ 44.27 ” N 98 ° 14 ’ 55.31 ” W, 2338 m asl, coll. R. Robles, 12 January 1997; 1 male (CL 23.31 mm, CW 37.37 mm) (CNCR 35900), tributary stream in San Miguel La Presa Dam, municipality of Ixtacuixtla, 19 ° 22 ’ 6.87 ” N 98 ° 23 ’ 42.16 ” W, 2347 m asl, coll. K. J. Madrigal, C. F. Vergara, J. Jiménez, A. Hernández, G. Ugarte & R. Rugerio, 16 May 2021; 1 male (CL 11.29 mm, CW 19.19 mm), 2 females (CL 18.74 – 21.06 mm, CW 31.45 - 35.44 mm) (CNCR 35901), same locality, date, collectors and identifier as previous record; 4 males (CL 15.40 – 17.77 mm, CW 27.07 – 30.44 mm), 1 female (CL 15.55 mm, CW 26.29 mm) (CNCR 35902), Palo Huérfano Spring, barrio de Atotonilco, municipality of Amaxac de Guerrero, 19 ° 21 ’ 44.96 ” N 98 ° 10 ’ 55.71 ” W 2283 m asl, coll. K. J. Madrigal & R. Rugerio, 25 May 2021; 1 male (CL 17.44 mm, CW 30.59 mm) (CNCR 35904), tributary stream in San Miguel La Presa Dam, municipality of Ixtacuixtla, 19 ° 22 ’ 6.87 ” N 98 ° 23 ’ 42.16 ” W, 2347 m asl, coll. K. J. Madrigal, C. F. Vergara, J. Jiménez, A. Hernández, G. Ugarte & R. Rugerio, 16 May 2021; 3 males (CL 13.90 – 17.22 mm, CW 22.94 – 28.35 mm), 1 female (CL 16.12 mm, CW 27.45 mm) (CNCR 35970), same locality as previous record, coll. K. J. Madrigal, C. Vergara & J. Jiménez, 2 September 2021; 1 male (CL 8.89 mm, CW 15.56 mm) (CNCR 36158), El Ojito Spring, pool in San Luis Apizaquito, municipality of Apizaco, 19 ° 26 ’ 16.28 ” N 98 ° 07 ’ 00.71 ” W, 2418 m asl, coll. E. Moreno, K. J. Madrigal & J. L. Villalobos, 10 November 2021; 1 male (CL 19.45 mm, CW 32.09 mm), 4 females (CL 16.89 – 23.21 mm, CW 29.37 – 39.28 mm) (CNCR 36508), stream in La Ascensión Huitzcolotepec, municipality of Xaltocan, 19 ° 25 ’ 24 ” N 98 ° 12 ’ 34 ” W 2338 m asl, coll. K. J. Madrigal & C. F. Vergara, 15 April 2022. –– Hidalgo. 4 males (CL 9.92 - 19.27 mm, CW 16.96 – 32.88 mm), 4 females (CL 8.16 – 18.14 mm, CW 13.90 – 29.85 mm) (CNCR 340), spring in Ventoquipa, municipality of Santiago Tulantepec de Lugo, 20 ° 02 ’ 03.99 ” N 98 ° 20 ’ 17.06 ” W, 2204 m asl, coll. A. E. Rabling, 16 April 1967; 1 male (CL 14.51 mm, CW 24.51 mm) (CNCR 12925), small springs tributaries to Tula River in Mixquiahuala, municipality of Mixquiahuala de Juárez, 20 ° 14 ’ 18.16 ” N 99 ° 12 ’ 42.67 ” W, 1969 m asl, coll. B. Barrera, 10 July 1994; 9 males (CL 8.50 – 16.73 mm, CW 15.13 – 27.75 mm), 8 females (CL 12.23 – 19.20 mm, CW 20.51 – 31.66 mm) (CNCR 36155), small springs in aquatic park Baño Grande, municipality of Mixquiahuala de Juárez, 20 ° 14 ’ 41.63 ” N 99 ° 12 ’ 18.62 ” W, 1914 m asl, coll. K. J. Madrigal, J. L. Villalobos & E. Moreno, 13 November 2021; 2 males (CL 12.81 – 15.78 mm, CW 21.36 – 26.69 mm), 4 females (CL 10.68 – 14.04 mm, CW 18.04 – 23.27 mm) (CNCR 361550), spring in Ventoquipa, municipality of Santiago Tulantepec de Lugo, 20 ° 02 ’ 03.99 ” N 98 ° 20 ’ 17.06 ” W, 2204 m asl, coll. E. Moreno, K. J. Madrigal & J. L. Villalobos, 12 November 2021.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA3C0450D87FB6031A4FE34.taxon	description	Redescription. Carapace slightly convex in dorsal view; surface finely punctuated / reticulated, punctuation grouped between postfrontal lobes and frontal border (Fig. 4 B). Superior frontal border absent; front gently curved downwards; frontal border bilobed, continuous, slender, slightly punctuated laterally, giving crenated appearance, central portion with shallow, narrow depression (Fig. 4 A, B). Postfrontal lobes visible, faintly elevated; frontal median groove poorly defined, wide; cervical grooves gently marked, straight, shallow, not reaching anterolateral margin (Fig. 4 B). Gastric region well defined, convex, slightly higher than branchial region; cardiac region marked by 3 well delineated depressions, 2 near urogastric area, 1 close to intestinal region; urogastric sutures Y-shaped, horizontally disposed, middle faintly curved, elongated. External orbital angle without notch (Fig. 4 B). Anterolateral margin ornamented with low granules, most conspicuous in meso-branchial section, decreasing progressively in size towards orbital external angle; posterior margin straight (Fig. 4 B). Frontal border inclined towards central portion in frontal view, convex at level of antennular fossae, without widening in lateral angle (Fig. 4 A). Orbits subquadrangular; inferior orbital margin gently crenated, horizontally displayed; external orbital angle subacute; internal orbital tooth subtriangular, like grooved plate, separated from lateral angle of frontal border, leaving wide hiatus (Fig. 4 A). Antennae not separated from front; antennal basal article not covered by lateral angle of frontal border (Fig. 4 A). Interantennular septum small, sharp, inserted under frontal border. Operculum of antennal gland ovoid, with tuft of bristles on external portion. Efferent channel aperture rectangular, its width / length ratio 1.92. Epistomal tooth at septum level, triangular, downwards directed, apex slightly projected anteriorly (Fig. 4 A). Pterygostomial region and epistome pubescent. Third maxillipeds completely cover buccal cavity; ischium of third maxilliped subtrapezoidal, width / length ratio 0.89, gnathal margin ornamented with small triangular teeth decreasing progressively in size distally, bristle tufts between teeth; merus of third maxilliped with anterolateral border rounded, does not extend beyond buccal cavity, width / length ratio 1.25; anterior notch present, well defined; gnathal border gently excavated to permit palp insertion, inferior portion ornamented with small teeth and long bristles; palp mesially pubescent, denser towards dactylus; third maxilliped surface punctuated, each punctuation with setae, distributed along gnathal margin; exognath wider at base, ending in acute apex, exognath / ischium length ratio 0.83 (Fig. 4 C). Chelipeds asymmetrical. Internal border of merus ornamented with small tubercles; carpus internal border granulated, small, rounded, with wide granules, distal portion armed with triangular spine, wide, with rounded apex. Major chela with palm surface punctuated; dactylus and fixed finger separated by broad gap; dactylus slender, slightly incurved; cutting borders dentated (Fig. 4 D). G 1 straight, robust, distal third twisted towards median axis of sternal surface of body. In caudal view, distal third straight; distal crest of caudo-marginal projection higher than lateral crest; distal end of caudal surface with broad concavity, moderately deep (Fig. 4 G). Caudal surface of mesial process with basal lobe rounded, poorly developed. In mesial view, caudo-marginal projection with subapical surface of caudal margin slightly excavated; crest of distal lobe smooth, widely rounded; distal lobe small, triangular, gently directed downwards, apex subacute, ornamented at tip with thin spine, acute and inconspicuous, separated from proximal lobe by V-shaped notch; proximal lobe U-shaped, elongated, length / width ratio 1.85, directed proximally, cephalic margin smooth, acutely curved distally (Fig. 4 H). In cephalic view, distal third straight; distal and proximal lobes of caudo-marginal projection not disposed in the same plane, former one slightly mesially directed, latter one with inner surface moderately swollen, with raised median carina reaching middle portion (Fig. 4 F). Mesial process subtriangular, perpendicular to G 1 principal axis, ending laterally in acute spine; superior border with sinusoidal short distal crest, rising in middle portion, extending to inner lobe; inferior border convex, ending in lateral spine; inner lobe subtriangular, subacute, in contact with inner surface of proximal lobe (Fig. 4 F). In lateral view, mesial process slightly projected caudo-laterally (Fig. 4 I). In distal view, apical cavity U-shaped; widening cephalad; central crest of apical cavity subtriangular, deep, broad, ending well before reaching inner lobe of mesial process; apical spines field deep, adjacent to inner surface of lateral crest, with approximately 27 moderately long spines, disposed in 3 – 4 irregular caudo-cephalic rows, cephalic spines partially covered by superior border of mesial process (Fig. 4 E). Aperture of spermatic channel in caudal position. Mesial process with lateral spine proximally broadened, distally straight, narrowing towards tip, faintly directed latero-caudally; basal lobe of caudal surface scarcely noticeable, rounded, poorly developed (Fig. 4 E.)	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA3C0450D87FB6031A4FE34.taxon	materials_examined	Type locality. El Ojito Spring, pool in San Luis Apizaquito town, municipality of Apizaco, Tlaxcala, Mexico (see Discussion, Table 5).	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA3C0450D87FB6031A4FE34.taxon	discussion	Remarks. The general morphological pattern of this species is constant throughout its distribution range. Nevertheless, some variations were found in the opening of the efferent branchial channel (subquadrangular to subrectangular), the elevation of the gastric region relative to the branchial region and a different degree of development of the internal orbital tooth. The differences in the G 1 comprise the different inclination of the distal third in cephalic view; the extension of the inner lobe of mesial process, that could stay in touch or not with the inner surface of the proximal lobe of the caudo-marginal projection; the presence of the small serration on the border of the distal lobe of the caudo-marginal projection is more evident in the population of San Miguel La Presa and absent in other localities; and the variation in the size of the basal lobe of the caudal surface of the mesial process. However, all these variations are not consistent with any geographic or population pattern, thus they may be representing the natural specific variation.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA3C0450D87FB6031A4FE34.taxon	distribution	Distribution. Pseudothelphusa americana s. s. is distributed in Mexico along the central portion of the TVB in the states of Hidalgo, Puebla, and Tlaxcala (Villalobos-Hiriart, 2005). This species has been collected in springs, streams, and rivers. The localities in Michac in Puebla, as well as San Miguel La Presa, Amaxac de Guerrero, Apizaco and La Ascención Huitzcolotepec in Tlaxcala, are new records.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	description	http: // zoobank. org / 2 C 9 A 52 D 9 - 6992 - 41 A 8 - 8 CCC-BED 2810 E 9 FCD (Figures. 5, 8)	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	materials_examined	Type material examined. Holotype (herein designated): male (CL 21.73 mm, CW 34.14 mm) (CNCR 29077), Ajolotero River, ~ 5 km SW of Chilapa, municipality of Chilapa de Álvarez, Guerrero, Mexico, 17 ° 34 ’ 30.00 ” N 99 ° 12 ’ 24.60 ” W, 1527 m asl, coll. J. L. Villalobos, A. Celis, O. Vázquez & G. García, 9 June 2000. Paratypes: 9 males (CL 14.42 – 21.44 mm, CW 22.29 – 33.62 mm), 4 females (CL 10.06 – 19.99 mm, CW 16.10 – 30.42 mm) (CNCR 29078), same data as holotype.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	description	Description. Carapace slightly convex in dorsal view; surface finely punctuated / reticulated, punctuation concentrated near anterolateral margin, between postfrontal lobes and on frontal border (Fig. 5 B). Superior frontal border absent; front gently curved downwards; frontal border bilobed, continuous, thick, smooth, central portion with shallow, narrow depression (Fig. 5 A, B). Postfrontal lobes well marked, slightly elevated; frontal median groove well defined, wide, shallow; cervical grooves notably marked, wider, deeper at proximal end, not reaching anterolateral margin (Fig. 5 B). Gastric region well defined, convex, slightly higher than branchial region, cardiac region marked by 2 shallow lateral depressions near urogastric region; urogastric sutures Y-shaped, horizontally arranged, slightly inclined, curved at middle. Notch on external orbital angle absent (Fig. 5 B). Anterolateral margin ornamented with low denticles, most conspicuous on meso-branchial section, decreasing progressively in size towards external angles; posterior margin straight (Fig. 5 B). Frontal border straight in frontal view, convex at level of antennular fossae, without widening at lateral angle (Fig. 5 A). Orbits subrectangular; orbital margin with inferior and lateral borders crenated; inferior border slightly inclined; external orbital angle subacute with lateral notch; internal orbital tooth subtriangular, like grooved plate, well-developed, separated from lateral angle of frontal border, leaving moderately narrow hiatus (Fig. 5 A). Antennae not separated from front; antennal basal article not covered by lateral angle of frontal border (Fig. 5 A). Interantennular septum small, sharp, inserted under inferior frontal border. Operculum of antennal gland ovoid, tuft of bristles on external portion. Branchial efferent channel aperture subquadrangular, width / length ratio 1.2. Epistomal tooth at septum level, triangular, downwards directed. Pterygostomial region, epistome pubescent. Third maxillipeds covering buccal cavity, ischium subtrapezoidal, width / length ratio 0.90, gnathal margin ornamented with small subtriangular teeth decreasing in size distally, bristle tuft placed between teeth; merus of third maxilliped with anterolateral border rounded, not extending beyond buccal cavity, width / length ratio 1.21; anterior notch present, shallow; gnathal border gently excavated to permit the palp insertion, inferior portion ornamented with small teeth and long bristles; palp mesially pubescent, denser towards the dactylus; third maxilliped surface punctuated, each punctuation with setae, distributed longitudinally in the gnathal margin; exognath wider at base, ending in acute apex, exognath / ischium length 0.65 (Fig. 5 C). Chelipeds asymmetrical; internal border of merus with small tubercules; internal border of carpus with low, wide granules, armed with triangular, acute spine. Major chela with palm surface punctuated; dactylus and fixed finger separated by moderately broad gap; dactylus some widened, faintly curved; cutting borders with low teeth dentated (Fig. 5 D). G 1 straight, robust, with distal third twisted towards median axis of sternal surface of body. In caudal view, distal third straight; distal crest of caudo-marginal projection higher than lateral crest; distal end of caudal surface with wide, shallow concavity (Fig. 5 G). Caudal surface of mesial process with basal lobe well-developed, conical, wide at base. In mesial view, caudo-marginal projection with subapical surface of caudal margin slightly excavated; crest of distal lobe smooth, rounded caudally, straight cephalically; distal lobe small, triangular, slightly directed downwards, with acute apex, faintly projected towards mesial side, separated from proximal lobe by wide V-shaped notch; proximal lobe U-shaped, broad, short, length / width 1.66, moderately oblique to principal axis, distal margin rounded (Fig. 5 H). In cephalic view, distal third moderately tilted laterally; distal and proximal lobes of caudo-marginal projection disposed in same plane, proximal lobe oblique to G 1 principal axis, inner surface moderately swollen, with low median carina reaching beyond middle portion (Fig. 5 F). Mesial process subtrapezoidal, perpendicular to G 1 principal axis, ending laterally in triangular subacute spine; superior border straight with long and sinusoidal crest covering two thirds of process reaching inner lobe; inferior border with proximal portion convex, distal portion gently concave, extending to lateral spine; inner lobe subquadrangular, laminar, in contact with inner surface of proximal lobe (Fig. 5 F). In lateral view, mesial process latero-caudally curved; basal lobe of caudal surface with conical apex (Fig. 5 I). In distal view, apical cavity U-shaped, of uniform width; central crest of apical cavity moderately high, straight, reaching inner lobe of mesial process; apical spines field deep, in central position, with approximately 17 moderately short spines, disposed in 2 irregular caudo-cephalic rows, cephalic spines partially covered by superior border of mesial process (Fig. 5 E). Aperture of spermatic channel in caudal position. Mesial process with lateral spine moderately slender at base, curved, directed latero-caudally; basal lobe of caudal surface noticeable, conical, developed (Fig. 5 E).	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	materials_examined	Type locality. Ajolotero River (17 ° 34 ’ 30.00 ” N 99 ° 12 ’ 24.60 ” W, 1527 m asl), ~ 5 km SW of Chilapa, municipality of Chilapa de Álvarez, Guerrero, Mexico.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	etymology	Etymology. The species epithet is taken from the municipality of Chilapa de Álvarez where the species was collected.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	discussion	Remarks. The G 1 of P. chilapaensis sp. nov. is similar to that of P. americana s. s., particularly in mesial view. However, it is possible to recognize certain differences between them: in caudal view, P. chilapaensis sp. nov. has the basal lobe of the caudal surface of the mesial process well-developed, and it is discernible laterally and caudally (Fig. 5 G, I) (versus basal lobe inconspicuous in P. americana s. s.; Fig. 4 G, I); in cephalic view, the lobes of the caudo-marginal projection are moderately inclined away from the main axis of the gonopod in P. chilapaensis sp. nov. (Fig. 5 F) (versus straight and parallel to the main axis in P. americana s. s.; Fig. 4 F); and in mesial view, the distal crest of the caudo-marginal projection of P. chilapaensis sp. nov. is rounded caudally and straight cephalically, and the proximal lobe is short and broad, its length / width ratio is 1.66, and is moderately oblique to the principal axis, and rounded distally (Fig. 5 H) (versus the distal crest of the caudo-marginal projection is widely rounded, the proximal lobe is slender and elongated, its length / width ratio is 1.85, and is proximally directed, and acutely curved distally in P. americana s. s.; Fig. 4 H). Additionally, some somatic structures display morphometric differences, e. g., the exognath length / ischium length ratio is 0.65 in P. chilapaensis sp. nov. (versus 0.83 in P. americana s. s). Bott (1968) revised freshwater crabs that came from Xochitempa near Chilapa, Guerrero. He considered those individuals to belong to Pseudothelphusa (Pseudothelphusa) americana americana without further comments. Villalobos-Hiriart (2005) identified the crabs from Río Ajolotero as a different morphological form of P. americana s. s., but just as Bott (1968), did not make any remarks about them. Our phylogenetic analysis retrieved P. chilapaensis sp. nov. as sister group to P. belliana + P. hoffmanae clade and completely separated from P. americana s. s. populations (Fig. 2). In summary, the molecular, morphological, and geographical evidence presented support the description of P. chilapaensis sp. nov. as a new taxon.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFA7C0580D87FB1F3756FD80.taxon	distribution	Distribution. Pseudothelphusa chilapaensis sp. nov. was collected from the type locality, Ajolotero River, and at Xochitempa (17 ° 31 ’ 36.73 ” N 99 ° 10 ’ 10.47 ” W; 1900 m asl), near Chilapa, Guerrero, Mexico (Bott, 1968).	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	description	http: // zoobank. org / 76 ED 3203 - 4225 - 4 E 3 B- 8 DF 7 - B 84 C 22 F 7 EF 76 (Figures. 6, 8)	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	materials_examined	Type material examined. Holotype (herein designated): male (CL 24.32 mm, CW 37.39 mm) (CNCR 36592), underground water channel, used for water supply to town San Marcos Necoxtla, municipality of Tehuacán, Puebla, 18 ° 24 ’ 37.44 “ N 97 ° 22 ’ 5.16 “ W, coll. K. J. Madrigal, E. F. Huerta, M. Hermoso & A. Estradas, 31 July 2022. Paratypes: 2 males (CL 17.72 – 17.92 mm, CW 26.34 – 27.05 mm), 2 females (CL 20.69 – 22.94 mm, CW 31.30 – 35.89 mm) (CNCR 36593), same data as holotype. Other material examined. 3 males (CL 19.85 – 21.47 mm, CW 29.82 – 32.33 mm), 2 females (CL 16.38 – 19.54 mm, CW 23.8 – 29.72 mm) (CNCR 36591), same locality as holotype, coll. E. F. Huerta, M. Hermoso & L. Hermoso, 24 June 2022.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	description	Description. Carapace slightly convex; surface finely punctuated, reticulated, punctuation denser near frontal border (Fig 6 B). Superior frontal border absent; front gently curved downwards; frontal border bilobed, continuous, thick, lightly punctuated, giving it crenated appearance, central portion with shallow depression (Fig 6 A, B). Postfrontal lobes barely distinguishable; frontal median groove poorly defined, wide; cervical grooves well marked, straight, deeper anteriorly, not reaching anterolateral margin (Fig 6 B). Gastric region well defined, convex, slightly higher than branchial region; cardiac region marked by 4 deep depressions, 2 near urogastric area, 2 close to gastric region; urogastric sutures Y-shaped, slender, horizontally disposed, straight in middle (Fig 6 B). External orbital angle without notch (Fig 6 B). Anterolateral margin lightly granulated, granules inconspicuous, widening towards orbital external angle; posterior margin straight (Fig 6 B). Frontal border straight in frontal view, convex at level of antennal fossae, with conspicuous widening at lateral angle (Fig 6 A). Orbits subquadrangular; margin completely crenated; inferior margin slightly inclined; external angle rounded; internal tooth subtriangular, as grooved plate, extends until meeting lateral angle of frontal border, closing the orbital hiatius (Fig 6 A). Antennae not separated from front; antennal basal article partially covered by widening of lateral angle (Fig 6 A). Interantennular septum small, blunt, not visible, inserted under frontal border. Operculum of antennal gland ovoid, with tuft of bristles on external portion. Efferent branchial channel aperture subquadrangular, width / length ratio 1.13. Epistomal tooth at septum level, triangular, downwards directed. Pterygostomial region and epistome pubescent. Third maxillipeds covering buccal cavity; ischium subtrapezoidal, width / length ratio 0.84, gnathal margin with small subquadrangular teeth decreasing in size distally, long bristle tufts in between teeth; merus with anterolateral border rounded, not extending beyond buccal cavity, width / length ratio 1.2, anterior notch well defined; gnathal border gently excavated to permit palp insertion, inferior portion with small teeth, long bristles; palp mesially pubescent, decreasing in thickness towards dactylus; third maxilliped surface punctuated, each punctuation with setae, distributed along gnathal margin; exognath wide, short, ending in rounded apex, exognath / ischium length ratio 0.56 (Fig. 6 C). Chelipeds asymmetrical. Merus internal border with small tubercles; carpus with internal border granulated, small, wide granules; distal portion with triangular spine, wide, with acute apex. Major chela with palm surface punctuated; fingers separated by broad gap; dactylus slender, curved; cutting borders with well-developed triangular teeth (Fig 6 D). G 1 strong, distal third twisted towards median axis of sternal surface of body. In caudal view, distal third straight; distal crest of caudo-marginal projection higher than lateral crest; distal end of caudal surface with shallow concavity (Fig 6 G). Caudal surface of mesial process with basal lobe poorly developed, barely noticeable. In mesial view, caudo-marginal projection with subapical surface of caudal margin slightly concave; crest of distal lobe smooth, widely rounded caudally, straight cephalically; distal lobe small, with acute apex, tip with fine spine, acute, inconspicuous, separated from proximal lobe by narrow V-shaped notch; proximal lobe U-shaped, short, broad, length / width ratio 1.11, oblique to principal axis, with widened, rounded apex (Fig 6 H). In cephalic view, distal third slightly inclined laterally, distal and proximal lobes of caudo-marginal projection disposed in same plane; short carina on inner face of proximal lobe, reaching one third of its surface, notably widened (Fig 6 F). Mesial process subtriangular, perpendicular to G 1 principal axis, ending laterally in triangular spine with subacute apex; superior border with straight, long crest, reaching inner lobe, middle portion elevated; inferior border gently curved; inner lobe widely triangular, with slightly rounded borders, wide, separated from inner surface of proximal lobe (Fig 6 F). In lateral view, mesial process caudally projected; basal lobe of caudal surface inconspicuous (Fig 6 I). In distal view, apical cavity wide U-shaped; central crest high, straight narrow; apical spine field deep, arranged in central stripe with approximately 17 long spines, cephalic spines partially covered by superior border of mesial process (Fig 6 E). Aperture of spermatic channel in caudal position, wide. Mesial process short, lateral spine faintly widened proximally, straight, laterally directed (Fig 6 E).	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	materials_examined	Type locality. Underground water channel used for water supply in town San Marcos Necoxtla (18 ° 24 ’ 37.44 ” N 97 ° 22 ’ 05.16 ” W), Tehuacán, Puebla, Mexico.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	etymology	Etymology. This species is dedicated to the Hermoso siblings, Ana Margarita, Luis and José who notified us about the existence of a Pseudothelphusa population in San Marcos Necoxtla, and helped us find the necessary crabs to carry out the analysis. The specific epithet is formed by using as root the last name ‘ Hermoso’ and adding to the suffix ‘ - orum’.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	discussion	Remarks. This species presents clear differences when compared to P. americana s. s. Pseudothephusa hermosorum sp. nov. shows lateral thickenings of the frontal border (Fig. 6 A) (versus no lateral thickenings, and a more slender frontal border in P. americana s. s.; Fig. 4 A); the exognath length / ischium length ratio in P. hermosorum sp. nov. is 0. 65 (Fig. 6 C) (versus 0.83 in P. americana s. s.; Fig. 4 C). The caudo-marginal projection of the male G 1 is notably different between the two species. The proximal lobe in P. hermosorum sp. nov. is oblique, short and broad, and with a length / width ratio of 1.11 (Fig. 6 H) (versus parallel to the principal axis, elongated, and with a length / width ratio of 1.85 in P. americana s. s.; Fig. 4 H). Additionally, the number of spines in the apical cavity is less in P. hermosorum sp. nov. (17) (Fig. 6 E) than in P. americana s. s. (21) (Fig. 4 E). Villalobos-Hiriart (2005) considered P. hermosorum sp. nov. as the Tehuacán form of P. americana s. s. and offered a formal description. In the present survey, molecular, morpho-geometric, and morphological evidence support the description of a new species. One ovigerous female of the new species was examined. The female (CL 20.69 mm, CW 31.3 mm) was carrying 79 eggs, each with a diameter of 2.12 – 2.35 mm.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFBAC05A0D87FD7A371CF9EC.taxon	distribution	Distribution. Pseudothelphusa hermosorum sp. nov. has only been collected in the type locality.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	description	http: // zoobank. org / CE 78 FBCC- 00 E 3 - 41 B 7 - BE 7 C-BE 3 A 39 A 4699 F (Figures. 7, 8)	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	materials_examined	Type material examined. Holotype (herein designated): male (CL 23.33 mm, CW 38.77 mm) (CNCR 342), Querétaro-state of México, Mexico, coll. anonymous. Paratype: male (CL 21.16 mm, CW 34.02 mm) (CNCR 344), same data as holotype.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	description	Description. Carapace slightly convex; surface finely punctuated / reticulated, punctuation clustered on branchial and gastric regions, and between frontal border and postfrontal lobes (Fig. 7 B). Superior frontal border present; front abruptly curved downwards, almost vertical; frontal border bilobed, continuous, thick, completely punctuated, with crenated appearance, central portion with shallow, wide depression (Fig. 7 A, B). Postfrontal lobes marked, somewhat elevated; frontal median groove ill defined, moderately deep, narrow; cervical grooves well defined, slightly curved proximally, deep, not reaching anterolateral margin (Fig. 7 B). Gastric region poorly delineated, convex; cardiac region marked by three shallow depressions; urogastric sutures Y-shaped, horizontally disposed, faintly inclined towards central portion, notably short (Fig. 7 B). Notch on external orbital angle (Fig. 7 B). Anterolateral margin with small, rounded granules, decreasing progressively in size anteriorly; posterior margin notably concave (Fig. 7 B). Superior frontal border forms an obsolete line in frontal view, slightly oblique, central portion interrupted by median groove; frontal border straight, convex at level of antennal fossae, wider at lateral angle (Fig. 7 A). Orbits subquadrangular; inferior margin gently crenated, horizontally displayed; external angle subacute; internal tooth subtriangular, as grooved plate, separated from lateral angle of frontal border, leaving a narrow hiatus (Fig. 7 A). Antennae not separated from front; antennal basal article partially covered by widening of lateral angle of frontal border (Fig. 7 A). Interantennular septum visible, sharp, inserted under frontal border. Operculum of antennal gland ovoid, with tuft of bristles on external portion. Efferent channel aperture subquadrangular, width / length ratio 1.07. Epistomal tooth at septum level, triangular, downwards directed (Fig. 7 A). Pterygostomial region and epistome pubescent. Third maxillipeds covering buccal cavity; ischium of third maxilliped subtrapezoidal, width / length ratio 0.91, gnathal margin with small subtriangular teeth decreasing in size distally, long bristle tufts in between teeth; merus of third maxilliped with anterolateral border rounded, extending beyond buccal cavity, width / length ratio 1.27; anterior notch well defined; gnathal border gently excavated for palp insertion, inferior margin with teeth and bristles; palp mesially pubescent, denser towards dactylus; surface punctuated, each punctuation with setae ,, distributed longitudinally along gnathal margin; exognath wider at base, ending in acute apex, exognath / ischium ratio 0.65. Chelipeds asymmetrical. Merus internal border with small tubercles, internal border of carpus with small, rounded granules, distally with triangular spine. Major chela with palm surface punctuated; fingers leaving narrow gap; dactylus slender, slightly curved; cutting borders with well-developed triangular teeth (Fig. 7 D). G 1 strong, distal third twisted towards median axis of sternal surface of body. In caudal view, distal third slightly inclined mesially; distal crest of caudo-marginal projection almost at same level as lateral crest; distal end of caudal surface with wide and shallow concavity (Fig. 7 G). Caudal surface of mesial process with basal lobe, rounded, inconspicuous. In mesial view, caudo-marginal projection with subapical surface of caudal margin slightly excavated; crest of distal lobe broadly rounded; distal lobe broad, triangular, cephalically directed, apex acute, without spine, separated from proximal lobe by wide V-shaped notch; proximal lobe U-shaped notch, length / width ratio 1.4, oblique to principal axis, constrained at base, wider at middle portion, rounded distally (Fig. 7 H). In cephalic view, distal third slightly inclined laterally, distal and proximal lobes of caudo-marginal projection disposed in same plane, long carina on inner face of proximal lobe, not reaching apex, notably widened (Fig. 7 F). Mesial process subtriangular, perpendicular to G 1 principal axis, ending laterally in acute spine; superior border with long, straight crest reaching inner lobe; inferior border gently convex proximally, concave towards lateral spine; inner lobe subtriangular, subacute, in contact with inner surface of proximal lobe of caudo-marginal projection (Fig. 7 F). In lateral view, mesial process latero-caudally curved; basal lobe of caudal surface rounded, inconspicuous (Fig. 7 I). In distal view, apical cavity wide U-shaped; central crest moderately high, slender, reaching inner lobe of mesial process; apical spine field deep, with approximately 30 long spines, disposed in irregular rows; cephalic setae partially covered by superior border of mesial border (Fig. 7 E). Aperture of spermatic channel in caudal position. Mesial process slightly proximally widened, lateral spine gently curved caudolaterally, long, slender, acute (Fig. 7 E).	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	materials_examined	Type locality. The precise locality is still unknown, but according to the original label it is somewhere between the states of Querétaro and México.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	etymology	Etymology. The specific epithet corresponds to the demonym of Querétaro’s inhabitants “ queretano ” and “ queretana ”; given that Pseudothelphusa is gender femenine, the female form of the demonym is used. The name must be taken as a noun in apposition.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	discussion	Remarks. The age of the original specimens and the absence of new collection of this taxon mean that DNA evidence for this species is not available. Pseudothelphusa queretana sp. nov. exhibits morphological affinities with P. purhepecha Ojeda-Escoto, Villalobos & Álvarez, 2017. However, both their disjunct distributions and differences in the G 1 morphology between these species (i. e, a mesial process with one lateral spine in P. queretana sp. nov. (Fig. 7 F, G) versus two lateral spines in P. purhepecha (Ojeda-Escoto et al., 2017) are enough evidence to recognize them as different. Villalobos-Hiriart (2005) recognized the new species as different from P. americana s. s., however, he did not make any further comments.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
03BE3C58FFB8C05D0D87F9563730FE10.taxon	distribution	Distribution. Pseudothelphusa queretana sp. nov. has not been collected since the last century. Unfortunately, the original label does not include clear information about neither the locality nor the date of the collection, but as far as we can know this species might be found somewhere between the states of Querétaro and México.	en	Madrigal, Kevin J., Villalobos-Hiriart, José Luis, Álvarez, Fernando (2025): Integrative taxonomic analysis of the Pseudothelphusa americana de Saussure, 1857 species complex with the description of three new species (Decapoda, Pseudothelphusidae, Pseudothelphusinae). Zootaxa 5659 (1): 1-28, DOI: 10.11646/zootaxa.5659.1.1, URL: https://doi.org/10.11646/zootaxa.5659.1.1
