taxonID	type	description	language	source
03BC87AAFFDAFFC9B83ABF33318FFDCC.taxon	type_taxon	Type species Epibrithus pustulatus Marshall, 1955, by original description.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDAFFC9B83ABF33318FFDCC.taxon	description	Redescription (♂ ♀) MEASUREMENTS. Medium-sized weevils, body length 6.6 – 9.44 mm. COLOR END VESTITURE. Integument dark red to black; integument of abdomen red-brown to black, ventrite 5 (♂) usually lighter red than 1 – 4. Dorsal vestiture (head, pronotum, elytra) forming a dense cover of small, round to elliptical (slightly longer than wide) appressed brown to light grey scales, not completely covering the integument; callosities on elytra and tubercles of pronotum with tufts (elytra) or single (pronotum) suberect elongate scales generally darker than surrounding vestiture; base of interstriae 1 with scale cover less dense, revealing integument, scales on legs generally more elongate (approx. twice as long as wide), similar in colour to vestiture on dorsum. Ventral vestiture (head, thorax, coxa, ventrites) similar to dorsum, scales generally 2 – 3 times as large, and more elongate (longer than wide); ventrites with sparse, stout, elongate, suberect scales, similar colour to surrounding vestiture but without pearly sheen. HEAD. Rostrum longer than wide (w / l ratio: 0.84 – 0.9), in dorsal view sides wider at antennal insertions than at base (♂ 1.12 – 1.31 ×; ♀ 1.21 – 1.27 ×), slightly concave in the basal ½ of length and strongly conveX in apical ½, in lateral view slightly conveX dorsally and concave ventrally (Figs 1 – 3); epifrons narrower than forehead, widening apicad, with a median longitudinal carina extending from frons to forehead mostly concealed by scales; frons Y-shaped, about ¾ as wide as width of eye in dorsal view, integument punctate, bearing 2 pairs of elongate setae; epistome widest at apex, semi-circular posteriorly with a carinate hind margin not extending backwards beyond the apex of pterygia, bilobate, bare of scales, micropunctate, with or without a pair of setae. Scrobes superficial, directed below middle of eye, densely covered with scales (Figs 1 – 3); mandibles trisetose, bare of scales; mentum bisetose; forehead flat or somewhat depressed in the middle, with an elongate median fovea. Vestiture of head made of dense appressed imbricate scales (as for elytra) and sparse elongate semi-erect flattened scales; lateral and ventral regions basal to frons with a line of very elongate, 5 × longer than wide flattened scales. Eyes convex, exceeding the lateral margin of the head capsule in dorsal view. Antennae robust; scape 0.73 – 0.83 × as long as funicle, slightly curved to nearly straight, cylindrical, moderately widening from base to apical ⅔, more so in apical ⅓, eXceeding anterior margin of pronotum in repose; integument densely covered with elongate imbricate scales and stout downcurved scales. Funicle with segments 1 – 2 elongate, 1 longer than 2 (1.14 – 1.68 ×), segments 3 – 6 subequal in length, longer than wide, conical, segment 7 subequal or slightly longer than 6, with elongate scales and stout erect setae; club spindle-shaped, segment 1 longer than 2, margins slightly sinuous, posterior 1 / 2 of segment 1 with vestiture as for funicle, rest of vestiture of club finely and densely setose. THORAX. Pronotum slightly wider than long (w / l ratio: ♂ 1.18 – 1.25, ♀ 1.30 – 1.32) in dorsal view, widest slightly beyond middle of length, apical margin 0.83 – 0.87 × as wide as base, sides moderately to strongly convex; in lateral view convex dorsally, highest point at or just before to middle of length; integument closely set with small subcontiguous tubercles, the apex of which is glabrous or more or less covered with scales, and bearing a short, stout, suberect scale, median line with a groove not reaching base and apex of pronotum; noto-sternal suture distinct, sinuous, tubercles reduced or absent at and below the level of this suture. Prosternum and prosternellum forming low obtuse spine-like process on either side of base of procoxae, covered with scales on prosternum and glabrous on prosternellum; mesosternum with raised intercoxal process covered with scales; metasternum with carinate margins around base of mesocoxae, may also be raised into a low ridge between basal half of mesocoxae; mesepimera narrowly triangular posteriorly, mesoventrite reaching elytral margin at apex; metanepisternal suture distinct in apical ⅓ – ½, superficial in posterior ½ – ⅔. SCUTELLUM. Scutellar shield not exposed. ELYTRA. Broadly ovate in dorsal view, (w / l ratio 0.63 – 0.73), sides sub-parallel or convex and widest beyond middle of length, apex ovately to broadly rounded; humeral angles obtuse, poorly distinct from lateral conveXity, located at 0.12 – 0.19 of elytral length; width at humeral angles 1.59 – 1.79 × (♂), 1.99 – 2.02 × (♀) as wide as base of prothoraX; in lateral view, dorsal line conveX, reaching highest point in basal ⅓ or about apical ⅓ beyond start of declivity; declivity steep in male, almost vertical in female; base of interstriae 1 with integument distinct, sparsely covered with scales or glabrous; each elytron with 10 shallow, hardly distinct striae, punctures partly irregular, almost or entirely concealed with scales; the interstriae with rows of somewhat irregular obtuse callosities which sometimes coalesce longitudinally or laterally with those on adjoining intervals, each callosity with scales more erect on posterior declivity than on base of elytra. METATHORACIC WINGS. Absent. LEGS. Robust. ProcoXae contiguous, mesocoXae separated by approXimately 1 / 5 × the width of a mesocoxa; femora clavate, unarmed, metafemora not reaching apex of elytra; tibiae slightly downcurved to nearly straight, internal margin bisinuate with a row of dark red to black denticles and generally with a row of 1 – 5 black suberect spines; external margin with dark semi-appressed setiform scales, internal margin with longer, more erect light setae; mucro concealed in a tuft of elongate curved golden-brown setae, distinctly longer and black on metatibiae in male (Figs 1 A, 2 A, 3 A – B); metatibiae without corbel, bears comb of black spines. Tarsi with segment 1 slightly wider than 2, 2 approximately ¾ the length of 1, 3 bilobed, slightly wider than 1 and 2; claws simple, strongly curved, weakly divaricate but not mobile, equal or subequal in length. ABDOMEN. Ventrite 1 with intercoXal process weakly (♀) or strongly (♂) concave in middle, ♂ 1.37 – 1.75 ×; ♀ 1.47 – 1.81 × the metacoXal width, apical edge bilobate to almost straight; in male ventrites 2 + 3 + 4 weakly concave in middle of apical ridge, concavities with integument poorly concealed by scales, apical edges slightly bilobate to almost straight in middle; ventrite 5 wide (w / l ratio: ♂ 1.49 – 1.78; ♀ 1.40 – 1.44), in female slightly conveX with a very shallow apical depression, rounded apically, with a homogeneous cover of scales, apical margin with thin, brown, elongate setae forming a hook shape at apeX, in males with a shallow cavity in apical ½ – ¾ of length, bearing a median carina, apical margin slightly to strongly bilobate, apex with two brushes of black setae. MALE TERMINALIA. Body of penis well sclerotized, 1.58 – 2.44 mm in length, elongate (w / l ratio: 0.27 – 0.37), sides in basal ⅓ – ½ subparallel or slightly concave in dorsal view, widest in basal ⅓ – ⅔, converging abruptly or regularly in apical ½ – ⅓, apeX acuminate; in lateral view strongly downcurved, curvature stronger near middle of length, with tip weakly or strongly downcurved; temones slightly shorter (0.91 ×) to slightly longer (1.04 – 1.14 ×) than body of penis. Copulatory sclerite with a left arm refleXed slightly outward or strongly inward, setose at apex. Tegmen slender, parameres distinct with fused bases divided by a median notch, manubrium more sclerotized than rest of tegmen, longer than diameter of ring. Spiculum gastrale well sclerotized, posteriorly curved and flattened to form a concave plate.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	description	Figs 1, 4 – 5	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	diagnosis	Diagnosis A morphologically variable species, even within populations. Can be distinguished from E. longicarinatus sp. nov. and E. boroveci sp. nov. by the cavity in ventrite 5 of male specimens that are covered across its area by scales (Fig. 1 E). The penis is shorter than in E. longicarinatus and E. boroveci (Fig. 1 C) with an abrupt taper in apical ⅓ (tapering regularly in E. longicarinatus and E. boroveci). Females of this species can be distinguished from those of E. longicarinatus by their generally slightly smaller size and the more rounded elytra in lateral view (which reach their highest point in the middle vs elytra in E. pustulatus, with the highest point being situated more posteriorly vs close to the elytral declivity in E. longicarinatus). It is the only species of Epibrithus amongst material examined where some specimens have a distinct darker ovate band on the elytra, rather than a more homogeneous grey vestiture (Fig. 1 A).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	etymology	Etymology The species was probably named after the callosities (‘ pustules’, Marshall, 1955) present on the interstriae on the elytra of adults.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	materials_examined	Material examined Holotype REPUBLIC OF SOUTH AFRICA – Western Cape Province • ♂; “ Olifants River bet; Citrusdal & Clanwilliam C. P. ” “ Epibrithus pustulatus TYPE ♂ ” “ HOLOTYPE ” “ Type SAM / Ent 4179 ” “ IMAGED LEICA LAS 4.9 SAMC 2024 ”; SAMC. Paratypes REPUBLIC OF SOUTH AFRICA – Western Cape Province • 4 ♂♂, 4 ♀♀; same collection data as for holotype; type SAM / Ent 4180; SAMC • 3 ♂♂, 3 ♀♀; same collection data as for holotype; NHMUK. Other material examined REPUBLIC OF SOUTH AFRICA – Western Cape Province • 5 ♂♂, 1 ♀ (1 specimen preserved in ethanol); The Baths; 32 ° 44 ′ 31 ″ S, 19 ° 02 ′ 6 ″ E; 3 Nov. 2019; J. Haran coll.; beating vegetation; JHAR 03020; CBGP • 1 ♀, Olifants River Mountains; O. Bokkeveld leg.; SAM-COL-A 045235, SAMC.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	description	Redescription (♂ ♀) BODY LENGTH. 6.6 – 9.1 mm. COLOUR AND VESTITURE. Body integument dark red to black; elytral vestiture forming a dense cover of small, round to elliptical (slightly longer than wide) appressed brown-to-light grey scales, not completely covering the integument; callosities on elytra with tufts of suberect elongate scales, generally darker than surrounding vestiture; each elytron with an oblique dark band from humeral angle to apical ⅔ of suture in approximately 10 – 100 % of specimens, depending on population; base of interstriae 1 with scale cover less dense, revealing integument. HEAD. Rostrum longer than wide (w / l ratio: ♂ 0.84 – 0.86; ♀ 0.89), in dorsal view sides wider at antennal insertions than at the base (1.12 – 1.24 × ♂; 1.27 × ♀); apical to middle dorsal edges of pterygia raised abruptly above the level of frons in some specimens, causing pterygial edges to appear ridge-like; frons with scattered scales in basal part, epistome bearing 1 pair of setae (lacking setae in single specimens). Forehead with a distinct median fovea between eyes, distance between dorsal posterior margin of eyes> than width of eye; eyes moderately conveX. Scape 0.73 – 0.82 × as long as funicle, slightly curved to nearly straight; funicle with segments 1 – 2 elongate, longer than wide (w / l ratio 1: 0.35 – 0.48; w / l ratio 2: 0.46 – 0.58), 1 subequal to longer (1.14 – 1.36 ×) than 2, segments 3 – 6 subequal in length, 7 subequal in length to slightly longer than 6, longer than wide (w / l ratio 3: 0.70 – 0.91), conical; club spindle-shaped, segment 1 longer than 2, margins slightly sinuous. PROTHORAX. Slightly wider than long (w / l ratio: ♂ 1.18 – 1.25, ♀ 1.32) in dorsal view, apical margin 0.83 – 0.87 × as wide as at base, sides moderately conveX, in lateral view highest point at the middle of length; median line with a groove not reaching base and apex of pronotum. ELYTRA. Sides sub-parallel in the middle (w / l ratio: 0.67 – 0.73), apex broadly rounded; humeral angles located at 0.16 – 0.19 of elytral length; in lateral view, dorsal line conveX, reaches highest point in basal ⅓. Size of elytral callosities differs between populations. LEGS. Tibiae nearly straight, internal margin often with row of 3 – 4 black, suberect spines; mucro concealed in a tuft of elongate curved golden-brown setae, distinctly longer and black on metatibiae in male. ABDOMEN. Ventrites with pearly white rounded scales, contiguous but not concealing the integument; ventrite 1 intercoXal process slightly wider than the metaXocal width (♂ 1.67 – 1.75 ×; ♀ 1.81 ×), apical edge slightly bilobate in middle; in male ventrites 2 + 3 + 4 apical edges almost straight in middle; ventrite 5 (♂ 0.66 – 0.70 ×; ♀ 0.77 ×) as long as 2 + 3 + 4, ventrite 5 wide (ratio w / l: ♂ 1.64 – 1.78; ♀ 1.44), male with a shallow cavity in apical ⅔ of length, bearing an indistinct short apical median carina shorter than ¼ the length of cavity, margin slightly bilobate, vestiture of ventrite 5 with a homogeneous cover scales, revealing integument but covering the bottom of cavity, apex with two short brush of black setae (Fig. 1 E). MALE TERMINALIA. Body of penis short (1.58 – 1.65 mm), elongate (w / l ratio: 0.33 – 0.37), widest in basal ⅔, sides of basal ⅔ subparallel in dorsal view, converging abruptly at apical ⅓, then regularly apicad, apeX acuminate, in lateral view downwards curvature strongest in middle ⅓, apeX with very slight downwards curvature; temones 1.04 – 1.12 × as long as body of penis (Fig. 1 C). Copulatory sclerite with left arm refleXed outward and curving slightly upward apicad in dorsal view, setae at base about a ⅓ as long as arm, left area of body wider than right (Fig. 1 D). Spiculum gastrale posteriorly curved. Life history Specimens were observed in large numbers during daytime in a small patch of forest in the Citrusdal area (JHAR 3020). Adults were found on the leaves of trees and on the lower branches, where vegetation was very dense (Fig. 4 A – B). The absence of specimens in the fynbos surrounding this patch (intensively sampled) suggests that E. pustulatus has a preference for forest habitat. This species was sampled on Galenia africana (Aizoaceae) by R. Borovec, but no specific host could be identified by the second author while sampling. Adults were collected in November.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	distribution	Distribution This species appears to occur naturally around the Citrusdal area and north towards the Olifants River (Fig. 4 D).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFDCFFC4B854BF213788FCBC.taxon	discussion	Remarks The species is morphologically variable, in size, relative size of callosities, colour and morphological ratios, even within populations. Larger scale genetic analysis will be necessary to determine whether cryptic species occur within this species concept.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	description	urn: lsid: zoobank. org: act: 00 B 4 FCAE- 2 A 33 - 4531 - AE 28 - 4 EDB 5266 ADB 0 Figs 2, 4 C – D, 5	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	diagnosis	Diagnosis Epibrithus longicarinatus sp. nov. can be distinguished from E. pustulatus and E. boroveci sp. nov. by the apical median carina in ventrite 5 of male being ½ the length of the ventrite (being proportionally shorter in the other two species). Unlike E. boroveci males, the apical median carina does not bear a brush of erect setae. From the specimens examined, this appears to be the largest species of Epibrithus, and has the proportionally largest eyes. The distance between the dorsal posterior margin of the eyes is less than the width of an eye in male specimens.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	etymology	Etymology This species is named for the elongated median carina in the cavity of ventrite 5 in males.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	materials_examined	Material examined Holotype REPUBLIC OF SOUTH AFRICA – Western Cape Province • ♂; “ REP. OF SOUTH AFRICA; WC. Algeria; 1 Xi. 2022; J. Sudre coll. ” “ JHAR 05672 - 01; - 32.394 19.086 [32 ° 23 ′ 38.4 ″ S, 19 ° 05 ′ 09.6 ′ ″ E]; Beating vegetation; SAMC ” “ HOLOTYPE; Epibrithus longicarinatus; Hansen & Haran 2025 ”; SAMC. Paratype REPUBLIC OF SOUTH AFRICA – Western Cape Province • 1 ♀; same collection data as for holotype; SAMC.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	description	Description (♂ ♀) BODY LENGTH. 9.0 – 9.5 mm. COLOUR AND VESTITURE. Body integument dark red to black, elytral vestiture consists of a dense cover of small, round to elliptical (slightly longer than wide) appressed light grey scales, callosities on elytra with tufts of suberect elongate scales, generally darker than surrounding vestiture; base of interstriae 1 with scale cover less dense, revealing integument. HEAD. Rostrum longer than wide (w / l ratio: 0.85), in dorsal view sides wider at antennal insertions than at the base (♂ 1.31 ×; ♀ 1.21 ×); frons with scattered scales in basal part only, and bearing 2 pairs of elongate setae; epistome bare of setae; forehead with a distinct median fovea between eyes, distance between dorsal anterior margin of eyes dorsally <than width of eye (♂), eyes large, conveX. Scape 0.78 – 0.82 × as long as funicle, nearly straight; funicle with segments 1 – 2 elongate, longer than wide (w / l ratio 1: 0.32 – 0.38; w / l ratio 2: 0.43 – 0.53), 1 longer (♂ 1.45 ×; ♀ 1.68 ×) than 2, segments 3 – 6 subequal in length, longer that wide (w / l ratio 3: 0.75 – 0.89), conical, segment 7 slightly longer than segment 6; club spindle-shaped, segment 1 longer than 2, margins slightly sinuous. PROTHORAX. Slightly wider than long (w / l ratio: ♂ 1.23, ♀ 1.30) in dorsal view, apical margin 0 × as wide as at base, sides moderately convex in male and more strongly convex in female, in lateral view highest point just before middle of length; tubercles are sparsely covered in scales at apex (revealing integument), median line with a groove not reaching base and apex of pronotum. ELYTRA. Sides sub-parallel (♀) or conveX and widest just beyond middle of length (♂) (w / l ratio: ♂ 0.63; ♀ 0.73) apeX ovately rounded (♂), broadly rounded (♀); humeral angles located at 0.13 – 0.16 of elytral length; in lateral view, dorsal line conveX, reaches highest point in about apical ⅓ beyond start of the declivity, callosities on apical ½ of posterior declivity with hook-shaped scales more elongate than on basal portion of elytra. LEGS. Tibiae slightly downcurved, internal margin often with row of 2 – 5 black, suberect spines; mucro concealed in a tuft of elongate curved golden-brown setae, distinctly longer and black on metatibiae in male. ABDOMEN. Ventrites with pearly white rounded to elongate scales, contiguous but not concealing the integument; ventrite 1 intercoXal process slightly wider than the metaXocal width (♂ 1.43 ×; ♀ 1.47 ×), apical edge almost straight in middle; in male ventrites 2 + 3 + 4 apical edges almost straight in middle; ventrite 5 (♂ 0.77 ×; ♀ 0.91 ×) as long as 2 + 3 + 4, ventrite 5 wide (w / l ratio: ♂ 1.49; ♀ 1.40), male with a shallow cavity in apical ¾ of length, bearing an elongate median carina in apical ½ of length of cavity, margin strongly bilobate, vestiture of ventrite 5 with a bare patch revealing integument either side of the median carina in apical ½ of cavity, median carina with sparse scales and erect setae, apeX with brushes of black setae (Fig. 2 E). MALE TERMINALIA. Body of penis long (2.28 mm), elongate (w / l ratio: 0.33), widest in basal ½, sides of basal ½ subparallel in dorsal view, converging regularly apicad from apical ½ of length, apeX acuminate, in lateral view downwards curvature strongest just before middle and again at apex, apex with strong downwards curvature, temones 1.14 × longer than body of penis. Copulatory sclerite with all structures strongly refleXed inward, giving a rounded appearance, left area of body wider than right. Parameres diverging strongly apically; spiculum gastrale posteriorly curved. Life history This species was collected in the daytime from vegetation in a small patch of forest during November.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	distribution	Distribution Limited information available. This species is known from a limited number of specimens collected at Algeria in the Cederberg mountains (Fig. 4 D).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD1FFC7BB89BEF237E0FDC3.taxon	discussion	Remarks Only one male (the holotype) and one female (the paratype) were collected, therefore knowledge about intraspecific variability is still unknown.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	description	urn: lsid: zoobank. org: act: 30 F 43113 - CB 52 - 400 E- 8 D 82 - 224 DD 432 DC 53 Figs 3, 4 C – D, 5	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	diagnosis	Diagnosis Epibrithus boroveci sp. nov. appears to be somewhat intermediate morphologically between E. pustulatus and E. longicarinatus sp. nov. and can be distinguished from the latter two species by the median carina of ventrite 5 carrying a brush of setae and a penis that is longer than the temones (Fig. 3 C). The median carina on ventrite 5 is relatively shorter, and the apical margin of this ventrite less strongly bilobed, than in male E. longicarinatus (Figs 2 E, 3 E).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	etymology	Etymology This species is named after our colleague and friend Roman Borovec, a weevil taxonomist who has contributed a huge body of knowledge on South African entimine weevils; and collected the paratypes for this species.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	materials_examined	Material examined Holotype REPUBLIC OF SOUTH AFRICA – Western Cape Province • ♂; “ REP. OF SOUTH AFRICA; WC. Piketberg; 26 viii. 2023; S. Hansen coll. ” “ JHAR 06376 - 01; - 32.820 18.665 [32 ° 49 ′ 12.0 ″ S, 18 ° 39 ′ 54.0 ″ E]; Beating fynbos; SAMC ” “ HOLOTYPE; Epibrithus boroveci; Hansen & Haran 2025 ”; SAMC. Paratypes REPUBLIC OF SOUTH AFRICA – Western Cape Province • 15 ♂♂, 4 ♀; Aurora Mts., 32 ° 41 ′ 638 ″ S, 18 ° 32 ′ 350 ″ E; 715 m a. s. l.; 10 Nov. 2016; R. Borovec coll.; beating trees at night and morning; CBGP (1 ♂, 1 ♀); CMNC (1 ♂); NHMUK (1 ♂); RB (9 ♂♂, 2 ♀); SAMC (1 ♂, 1 ♀); SANC (1 ♂); TMSA (1 ♂).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	description	Description (♂ ♀) BODY LENGTH. 7.7 mm. COLOUR AND VESTITURE. Body integument dark red to black; elytral vestiture forming a dense cover of small, round to elliptical (slightly longer than wide) appressed light grey scales; callosities on elytra with tufts of suberect elongate scales, generally darker than surrounding vestiture, base of interstriae 1, prothorax, and head of some specimens, with scale cover less dense, revealing integument. HEAD. Rostrum longer than wide (w / l ratio: ♂ 0.9), in dorsal view sides wider at antennal insertions than at the base (♂ 1.25 ×); frons with scattered scales in basal part; epistome bearing no or 1 pair of setae; forehead with indistinct to distinct median fovea between eyes, distance between dorsal anterior margin of eyes dorsally> width of eye; eyes moderately conveX. Scape 0.77 × as long as funicle, nearly straight; funicle with segments 1 – 2 elongate, longer than wide (w / l ratio 1: ♂ 0.39; w / d ratio 2: ♂ 0.46) longer (♂ 1.22 ×) than 2, segments 3 – 7 longer than wide (w / l ratio 3: ♂ 0.72) subequal in length, or segment 3 and 7 being slightly longer than 4 – 6, conical; club spindle-shaped, segment 1 longer than 2, margins slightly sinuous. PROTHORAX. Slightly wider than long (w / l ratio: ♂ 1.21) in dorsal view, apical margin 0.85 × as wide as at base, sides moderately convex, in lateral view highest point just before middle of length; tubercles have sparse to contiguous scales at base and are largely bare at the apex, revealing integument; median line with a groove not reaching base and apex of pronotum. ELYTRA. Widest anterior to middle of length (w / l ratio: ♂ 0.63), apeX ovately rounded (♂), broadly rounded (♀); humeral angles located at 0.12 of elytral length; in lateral view, dorsal line conveX, reaches highest point just beyond middle of length to about apical ⅓ beyond the start of the declivity. LEGS. Tibiae nearly straight, internal margin of some legs with row of 1 – 3 black, semi erect spines; apical mucro concealed in a tuft of elongate curved golden-brown setae, distinctly longer and black on metatibiae in male. ABDOMEN. Ventrites with pearly white rounded to elongate scales, contiguous but not concealing the integument; ventrite 1 intercoXal process 1.37 × metacoXal width (♂), in some male specimens bearing a weak median carina visible through scales in apical ½, apical edge bilobate (♂) or straight (♀) in middle; in male ventrites 2 + 3 + 4 apical edges almost straight to slightly bilobate in middle; ventrite 5 0.77 × as long as 2 + 3 + 4 (♂); ventrite 5 wide (ratio w / l: ♂ 1.62) with a shallow cavity in apical ½ of length, bearing a short median carina approXimately ¼ to ½ of length of cavity with a brush of erect grey setae, margin strongly bilobate, vestiture of male ventrite 5 with a bare patch revealing integument either side of the median carina in apical ⅓ of cavity, apeX with brushes of black setae (Fig. 3 E). MALE TERMINALIA. Body of penis long (2.44 mm), elongate (w / l ratio: 0.27), widest at base, sides of middle ⅓ slightly concave in dorsal view, converging regularly apicad from apical ⅓ of length, in lateral view downwards curvature strongest just before middle and again at apex, apex with strong downwards curvature, temones 0.91 × as long as body of penis (Fig. 3 C). Copulatory sclerite with left arm refleXed outward, curving slightly downward apicad in dorsal view, setae at base approximately ¾ as long as arm, right and left area of body of approXimately equal width (Fig. 3 D). Parameres diverging weakly apically; spiculum gastrale posteriorly curved strongly dorsally. Life history Collected from fynbos shrub during daytime in August, and from trees during night and morning in November.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	distribution	Distribution Mountains around Piketberg and Aurora (Fig. 4 D).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	discussion	Remarks In the two populations examined, the depth of the median fovea on forehead, the presence or absence of a median carina on ventrite 1 of males, the proportional length of the cavity and apical median carina of ventrite 5 of males, and proportional length vs width of elytra differ. These differences between populations may represent cryptic speciation. Morphological ratio measurements are based on the holotype. Other species	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	materials_examined	Material examined REPUBLIC OF SOUTH AFRICA – Western Cape Province • 1 ♂; Lemoens Hoek, Heidelberg, C. P.; Nov 1927; K. H. Barnard coll.; Epibrithus sp. n. det. R. Oberprieler, 198 *; SAM-COL-A 051991; SAMC.	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	discussion	This specimen does not match with the genus description above, with fewer and more elongate elytral callosities; truly free claws, a raised dorsal process on pronotum and no elongate metatibial setae in male. The specimen was also collected considerably to the east of other known Epibrithus spp. in CFR. The specimen represents a different and possibly novel genus that can be placed in Oosomini (open corbels, trisetose mandibles, dorsally placed antennal scrobes, and lack of ocular lobes, vibrissae and a humeral callus).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
03BC87AAFFD2FFC3B865BF13317EFBBA.taxon	description	Genetic analysis Interspecific variation in the 658 region of the COI gene supports morphological species divisions. Uncorrected p- distances of genetic variation in this region are 7.4 % and 13.2 % between E. pustulatus and E. longicarinatus sp. nov. and E. boroveci sp. nov. respectively, and 14.4 % between E. longicarinatus and E. boroveci. The preliminary maximum likelihood tree of the COI region suggests that E. boroveci is a sister clade to E. pustulatus and E. longicarinatus (Fig. 5). This pattern possibly refers to the isolation of the Piketberg mountain range from the Cederberg range (Fig. 4 C – D).	en	Hansen, Steffan P., Haran, Julien M. (2025): Revision of the South African genus Epibrithus Marshall (Curculionidae, Entiminae) with description of two new species. European Journal of Taxonomy 1012: 201-221, DOI: 10.5852/ejt.2025.1012.3043, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3043/13645
