taxonID	type	description	language	source
03BAF97BFFC1FF90FF46F97EC31F7C92.taxon	materials_examined	Type material. Ochodaeus montanus Fuente, 1912. NEOTYPE ♀ [erroneously considered a male!], designated by Alonso-Zarazaga and López-Colón (1996) (MNCN). Labels: (a) white rectangular paper, handwritten: Molinicos, 29. V. 1938; (b) white label, rectangular, handwritten: Ochodaeus / pocadioides Mots., ♂ / J. J. L. Colón. det., 1980; (c) white label, rectangular, framed in black, handwritten and printed: Ochodaeus / montanus / Fuente, 1912 / F. M. Piera det. 1988; (d) red label, rectangular, handwritten: Ochodaeus montanus / FUENTE, 1912 / M. A. Alonso Zarazaga / & J. I. López-Colón des. 97 / – NEOTYPUS –; (e) red label, rectangular, framed in black, printed: MNCN / Cat. Tipos Nº / 8380; (f) white label, rectangular, framed in black, printed: MNCN _ Ent / 2957; (g) white label, rectangular, handwritten and printed: Ochodaeus / montanus Fuente ♀ / J. B Huchet det. 2024. Other specimens (3 ex.): SPAIN: ALBACETE: River Guadalmena (Villapalacios), UTM: 30 SWH 2871, 730 m., 18 - V- 2003, attracted by artificial light in a meadow on the riverbank, in a holm oak forest environment with Quercus ilex L. ssp. ballota (Desf.) Samp., J. L. Lencina & F. Lencina leg. (1 ♂ / CJLL); JAEN: Sierra de Cazorla: Santiago-Pontones, Coto Ríos, 38.045688 - 2.852325 (650 mts), Light trap, Ribera, 7. VI. 2021, F. J. Campillo leg. (1 ♀ / CJBH). MURCIA: Collado de las Cabellas, Sierra de Villafuerte (Moratalla), UTM: 30 SWH 7422, 1650 m., 9 - VI to 8 - VII- 2017, in a clearing of pine forest Pinus nigra Arnold, in pitfall trap, J. L. Lencina & J. Miñano leg. (1 ♂ / CJLL). To the best of our knowledge, besides these four specimens, no other specimens of O. montanus are known, highlighting the extreme uncommonness of this species. Additionally, three specimens previously identified as O. montanus in the collections of the Museum of Madrid, as listed below (MNCN _ Ent 2959, 211587, and 211588), were found to correspond to O. inermis.	en	Gutierrez, Jean-Bernard Huchet José Luis Lencina (2024): New insights and comparative morphology of the rare Western Palearctic species Ochodaeus montanus Fuente, 1912 and its closest relative, O. inermis Reitter, 1893 (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 2024 (79): 1-16, DOI: 10.5281/zenodo.14662556
03BAF97BFFC2FF9DFF46FD1AC3EE781A.taxon	materials_examined	SPAIN: ALBACETE: Molinicos, 29.5.1938. [Escalera], MNCN _ Ent 2959, Ochodaeus pocadioides Mots. J. I. L. Colón det., 1980, Ochodaeus montanus Fuente, 1912, F. M. Piera det. 1988, Ochodaeus inermis Reitter, J. B Huchet det. 2024 (1 ♀ / MNCN); Albacete: Molinicos, without data [Escalera], MNCN _ Ent 211587, Ochodaeus pocadioides Mots. J. I. L. Colón det., 1980, Ochodaeus montanus Fuente, 1912, F. M. Piera det. 1988 [erroneously considered a male!], Ochodaeus inermis Reitter, J. B Huchet det. 2024 (1 ♀ / MNCN); Molinicos, 17. V. 2022, F. Lencina and F. Albert leg. (1 ex. / CJLL); Ayna, 20. VI. 2003, J. L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); same data and collectors but 27. VIII. 2003 (2 ex. / CJLL); Férez, 13. V. 2006, E. Aistleitner leg. (1 ex. / CJLL); Hellín, 30. V. 2012, J. L. Lencina leg. (1 ex. / CJLL); same data but 30. V. 2009, J. L. Lencina et al. leg. (1 ex. / CJLL); same data but 28. VIII. 2007, J. L. Lencina and F. Lencina leg. (6 ex. / CJLL); same data but 18. V. 2007, J. L. Lencina and F. Lencina leg. (1 ex. / CJLL); same data but 19. IV. 2007, J. L. Lencina, F. Albert, and F. Lencina leg. (5 ex. / CJLL); E. Hellin, Saladar de Cordovilla, 18. V. 2007, J. L Lencina leg., Ochodaeus inermis Reitter, 1893, det. J. L Lencina 2007 (2 ex. / CJBH); E. Hellin, Embalse de Camarillas, 19. IV. 2007, luz, J. L Lencina leg. (1 ex. / CJBH); E. Tobarra, Saladar de Cordovilla, 11. V. 2007, J. L Lencina leg., Ochodaeus inermis Reitter, 1893, det. J. L Lencina 2007 (1 ♂ / CJBH); Letúr, 02. II. 2008, J. L. Lencina and A. Andújar leg., berlese (1 ex. / CJLL); Motilleja, 26. VI. 2004, J. L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); Nerpio, 11. IX. 2004, J. L. Lencina, F. Albert, and F. Lencina leg. (2 ex. / CJLL); Tobarra, 01. V. 2004, J. L. Lencina, F. Albert, and F. Lencina leg. (2 ex. / CJLL); same data but 11. V. 1999, J. L. Lencina and F. Lencina leg. (7 ex. / CJLL); same data but 11. V. 2007, J. L. Lencina, F. Lencina, and P. Abarca leg. (3 ex. / CJLL); same data but 28. VII. 2007, J. L. Lencina leg. (2 ex. / CJLL); Villapalacios, 04. VI. 2004, J. L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); Yeste, 19. VI. 2004, J. L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); CÁDIZ: Jerez, without data. L. Cepero, “ codocera ” MNCN _ Ent 211572, Ochodaeus pocadioides Mots. Jerez / L. Cepero – codocera (1 ex. / MNCN); CÓRDOBA: Córdoba, without data, Mora!. Col. del Sr. Perez Arcas. MNCN _ Ent 211575 (1 ex. / MNCN); CUENCA: Villalba de la Sierra, 01. VII. 1989, J. L. Lencina and M. C. Muñoz leg. (1 ex. / CJLL); GRANADA: Puebla de Don Fadrique, Nablanca, 22 - 23 VI 2012, Z. Laštùvka leg., Ochodaeus inermis Reitter, 1893, J. B Huchet det. 2017 (1 ex. / NMPC); JAÉN: Úbeda, Jaén, without data, Ceballos, MNCN _ Ent 211574 (1 ex. / MNCN); Cazorla, Coto Ríos, 7. VI. 2021, F. J. Campillo leg., trampa de luz, 38.045688 / - 2.852325 (650 mts), Ribera (1 ♂, 1 ♀ / CJBH); Santiago-Pontones, 07. VI. 2021, F. J. Campillo leg. (2 ex. / CJLL); MADRID: El Regajal, Aranjuez, 28 - 05 - 2001, M. G. París, T. Luz. MNCN _ Ent 211588, Ochodaeus pocadioides Mots. J. I. L. Colón det., 1980, Ochodaeus montanus Fuente, 1912, F. M. Piera det. 1988, Ochodaeus inermis Reitter, J. B Huchet det. 2024 (1 ♂ / MNCN); Alcalá, 10.5.94. [Escalera], MNCN _ Ent 2955 (1 ex. / MNCN); MURCIA: Jumilla, Monte de Santa Ana, 10. V. 2013, trampa de luz, 30 SXH 4754, 650 m, J. L Lencina leg., Ochodaeus inermis Reitter, 1893, det. J. L Lencina 2013 (1 unsexed, 1 ♂ / CJBH); Jumilla, 29. V. 2004, F. Lencina leg. (1 ex. / CJLL); same data but 10. V. 2013, J. L. Lencina and F. Lencina leg. (12 ex. / CJLL); same data but 23. VIII. 2016, J. L. Lencina and M. C. Muñoz leg. (1 ex. / CJLL); Moratalla, 22. VI. 2013, UMU (1 ex. / CJLL); SÉVILLE: Villamanrique, 10 - IV- 55, J. Baraud, Muséum Paris, 1993, Coll. J. Baraud (2 ex. / MNHN); TERUEL: Teruel, without data, B. Muñoz 935 m, MNCN _ Ent 211573 (1 ex. / MNCN); ZARAGOZA: Val de Boquero el Ciego, la Retuerta, 11. VI. 2013, UTM 30 TYL 2795, R. Macía leg. (1 ex. / CJBH).	en	Gutierrez, Jean-Bernard Huchet José Luis Lencina (2024): New insights and comparative morphology of the rare Western Palearctic species Ochodaeus montanus Fuente, 1912 and its closest relative, O. inermis Reitter, 1893 (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 2024 (79): 1-16, DOI: 10.5281/zenodo.14662556
03BAF97BFFC9FF98FF46FB0DC4C37C75.taxon	description	Color patterns: within the Ochodaeini tribe, which includes, among others, the genera Ochodaeus Dejean, 1821 and Parochodaeus Nikolajev, 1995, the coloration of the habitus is relatively variable within a single species and thus not highly discriminatory. However, O. inermis is typically yellowish-tan to light brown, sometimes darkened, whereas the four known specimens of O. montanus exhibit a dark reddish-brown coloration. Male genitalia Ochodaeus montanus: The phallobase is elongated and dorsally curved, tapering to a point at the apex. The parameres are short, symmetrical, and rounded apically. The internal sac contains a single, large, boomerang-shaped endophallite (Fig. 16). The genital segment (urite IX) is elongated, with the base being nearly straight (Fig. 17). Ochodaeus inermis: The phallobase resembles that of the previous species. The parameres are short, symmetrical, and only slightly pointed at the apex. The internal sac contains two endophallites: one medium-sized, arched, and serrated along its convex surface, and another smaller one located beside it. The surface of the internal sac is partially covered with raspulae and fine spicules (Fig. 18). The genital segment (urite IX) is relatively wide and transverse, with the base distinctly sinuate in the middle (Fig. 19). Female terminalia Ochodaeus montanus (Fig. 20): The segment is complete. Tergite IX is arch-shaped, narrow, poorly sclerotized dorsally, and glabrous. Gonopod IX is trimeric, consisting of the subcoxite, coxite, and stylus. The subcoxite is roughly triangular and concave, with four to five long setae aligned at the tip of the ventral edge. The coxite is elongated, roughly cylindrical, slightly flattened at the apex, and bears 10 long setae. The style is elongated, digitiform, and bears eight thin setae apically. The trimeric gonopod IX was shown by Huchet (2021) to be a character of Ochodaeini, in contrast to the same structure in Nothochodaeini Nikolajev, 2015 (Nothochodaeus Nikolajev, 2005 and Ceratochodaeus Huchet, 2017) that is dimeric and lacks a terminal stylus. Ochodaeus inermis (Fig. 22): The gonopod IX is trimeric. The subcoxite is roughly triangular and concave, with five to six long setae arranged in staggered rows at the tip of the ventral edge. The coxite is elongated, roughly cylindrical, and slightly flattened at the apex, bearing 11 long setae. The style is short, digitiform, and bears five thin setae apically. Sexual dimorphism In O. inermis, the anterior tibiae of males are significantly narrower with sharper teeth, whereas in females, the tibiae are enlarged, and the teeth are wider (Fig. 14). To our knowledge, these characteristics associated with sexual dimorphism encountered across multiple species within the genus Ochodaeus have not been documented in the literature. Additionally, the clypeus in males is typically more markedly sinuate on either side of the midline, whereas in females, the clypeus exhibits a more smoothly curved anterior margin.	en	Gutierrez, Jean-Bernard Huchet José Luis Lencina (2024): New insights and comparative morphology of the rare Western Palearctic species Ochodaeus montanus Fuente, 1912 and its closest relative, O. inermis Reitter, 1893 (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 2024 (79): 1-16, DOI: 10.5281/zenodo.14662556
