taxonID	type	description	language	source
054E5D3A1170FFCEFFB3FD1316F82F90.taxon	materials_examined	Material. Holotype, male: “ 12. - 30.4.2007 Malaysia / Ringlet 926 m. n. m. / N 04 ° 23 E 101 ° 22 / Kremitovský ”, “ Holotype / Amphimenes (s. str.) / kmecoi sp. n. / des. Anichtchenko A. 24 [red label, handwritten] ” (DUBC).	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1170FFCEFFB3FD1316F82F90.taxon	diagnosis	Diagnosis. This new species is the most southern known representative of the genus, and seems to present transitory features between the subgenus Amblops Andrewes, 1931 and the “ medius – group ” of the subgenus Amphimenes s. str. (in sense of Fedorenko 2019). With Amblops it shares the indistinct temples, and with the nominotypical subgenus shares the transversally rugose intervals of elytra. By the small body size and the aedeagus strongly dilated in ventral view the new species is closely related to A. (s. str.) medius Fedorenko 2010, known from Southern Vietnam, but can be easily distinguished from it by the indistinct temples, the very narrow margins of pronotum and the occurrence of three discal pores on elytra.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1170FFCEFFB3FD1316F82F90.taxon	description	Description. Body length 5.0 mm. Dorsum black, mouthparts, legs and antennae light brown, clypeus and labrum brown (Fig. 1). Gula, trochanters and coxa brownish-red. 260 Eyes moderately convex, temples indistinct; posterior supraorbital seta situated in distance equal to radius of eye back. Labrum incised anteriorly. Frontal foveae shallow. Microreticulation isodiametric, smoothed on the vertex. Antennae moderately long, surpassing base of pronotum by last joint only. Pronotum 1.19 times as wide as long, 1.41 times as wide as head, with front angles protruding but slightly rounded apically; sides slightly angulate in front of lateral pore, slightly sinuate before hind angles; not reflexed between lateral pore and anterior angle, very slightly reflexed towards hind angles; lateral margin very narrow, especially so anteriorly. Hind angles straight. Base of pronotum is almost straight. Mid-line reaching anterior border, superficial on the disc, much deeper where adjoining basal transverse depression, latter distinctly separated from a very convex disc; lateral basal foveae indistinct, paramedian foveae flat and almost indistinct. Microreticulation on the disc strong, consist of polygonal meshes, towards the sides meshes becomes almost isodiametric, and towards hind angles meshes turns into small tubercles. Elytra oval, 1.37 times as long as wide, 1.56 times as wide as pronotum, broadest at about middle, with shoulders strongly rounded; fused along midline; apical truncature hardly sinuate between a rounded outer angle and a point- ed apex. Three discal setigerous pores: D 1 / EL = 0.15, D 2 / EL = 0.73 and D 3 / EL = 0.95. All elytral striae deep, crenulato-punctate, intervals strongly transversally striate. Microreticulation strongly transverse, surface slightly iridescent. Hind wings reduced. Ventral segments smooth, impunctate. Metepisternum short, sub square. Last metatarsomere with one distal pair of ventral setae only. Claws with four denticles, basal one much smaller. Basal third of male profemur without ventral tubercle. Aedeagus (Figs. 3 - 4): median lobe robust, abruptly bent near basal orifice, strongly in- flated on left side in ventral view; apical lamella New species of genus Amphimenes Bates, 1873 (Coleoptera, carabidae, Lebiinae) from Laos and Malaysia spatulate, with a rounded tip. Endophallus without distinct structures.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1170FFCEFFB3FD1316F82F90.taxon	etymology	Etymology. This species is named after my friend and colleague Rudolf Kmeco (Czechia), who kindly provided a lot of interesting material for determination.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1170FFCEFFB3FD1316F82F90.taxon	distribution	Distribution. Known only from the type locality in Malaysia. Figures 1 - 2. Habitus of Amphimenes in dorsal view. 1 – A. (s. str.) kmecoi sp. n., Holotype. 2 – A. (s. str.) hartmanni sp. n., holotype. Images courtesy A. Anichtchenko.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1173FFC8FF55FAD3113D2BE7.taxon	materials_examined	Material. Holotype, male: “ NE – Laos, Pr. Hua Phan / Ban Saleui, Phou Pan (Mt.) / ~ 20 ° 12 N, 104 ° 01 E / 31. V. 2011, 1300 - 1900 m, / local collector ”, “ collection / NATURKUNDE - / MUSEUM ERFURT [yellow label] ”, “ Holotype / Amphimenes (s. str.) / hartmanni sp. n. / des. Anichtchenko A. 24 [red label, handwritten] ” (NME).	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1173FFC8FF55FAD3113D2BE7.taxon	diagnosis	Diagnosis. This new species belongs to the “ piceolus ” - group sensu Fedorenko (2019). In general appearance, by the reduced hind-wings and the shape of aedeagus it is closely related to A. (s. str.) asahinai Nakane, 1957, known from Taiwan, from which it can be distinguished by the small size, the narrow lateral margins of pronotum and the shape of median lobe of aedeagus, curved before the middle in lateral view, and wide in dorsal view.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1173FFC8FF55FAD3113D2BE7.taxon	description	Description. Body length 4.5 mm. Dorsum brown, mouthparts, legs and antennae light brown, clypeus and labrum brown (Fig. 2). 261 Alexander Anichtchenko Gula, trochanters and coxa brownish-red. Eyes flattened, temples about half as long as eye; posterior supraorbital seta situated in distance equal to radius of eye back. Labrum slightly incised. Frontal foveae shallow. Microreticulation consists of polygonal meshes. Antennae long, surpassing base of pronotum by two and half last joints. Pronotum 1.25 times as wide as long, 1.42 times as wide as head, with front angles slightly protruding and rounded apically; sides slightly angulate in front of lateral pore, weakly sinuate before hind angles; not reflexed between lateral pore and anterior angle, very slightly reflexed towards hind angles; lateral margin narrow, gradually expanding from anterior angles towards hind angles. Hind angles obtuse. Base of pronotum slightly protruding backwards. Mid-line reaching anterior border, superficial on the disc. Basal foveae flat and indistinct. Microreticulation on the disc very strong, almost isodiametric, towards the sides meshes becomes stronger, and towards hind angles meshes turns into small tubercles. Elytra oval, 1.32 times as long as wide, 1.66 times as wide as pronotum, broadest at about middle, with shoulders strongly rounded; not fused along midline; apical truncature moderately sinuate between a rounded outer angle and a pointed apex. Three discal setigerous pores: D 1 / EL = 0.32, D 2 / EL = 0.78 and D 3 / EL = 0.94. All elytral striae deep, gently punctate, intervals moderately transversally striate, on the disc of elytra transverse striation becomes weak. Microreticulation strongly transverse. Hind wings reduced. Ventral segments smooth, impunctate. Metepisternum short, subsquare. Last metatarsomere with two distal pairs of ventral setae. Claws with four denticles, basal one much smaller. Basal third of male profemur with obtuse and wide ventral tubercle. Aedeagus (Figs. 5 - 6): median lobe elongate, in lateral view evenly rounded dorsally and bend- ed at an obtuse angle before the middle ventrally; apical lamella elongate, with a rounded tip. Endophallus without distinct structures.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1173FFC8FF55FAD3113D2BE7.taxon	etymology	Etymology. This new species is named for Matthias Hartmann, Coleoptera specialist, and Managing Director at Naturkundemuseum Erfurt (Germany).	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1173FFC8FF55FAD3113D2BE7.taxon	distribution	Distribution. Known only from type locality in Laos. Figures 3 - 6. Aedeagus structure of Amphimenes in lateral and dorsal view. 3 - 4 – A. (s. str.) kmecoi sp. n. 5 - 6 – A. (s. str.) hartmanni sp. n. Images courtesy A. Anichtchenko. 262 New species of genus Amphimenes Bates, 1873 (Coleoptera, carabidae, Lebiinae) from Laos and Malaysia	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1175FFC8FF55FF3816712928.taxon	materials_examined	Material. 1 male: “ China, Yunnan / Honghe, Qibaofeng, 1350 m, 23 ° 4 ’ 4.2 N 103 ° 23 ’ 46.9 E, 17. vi. 2018, A. Weigel leg. ” (NME); 1 male: “ China, Yunnan / Honghe, Qibaofeng, 1350 m, 23 ° 4 ’ 4.2 N 103 ° 23 ’ 47 E, 13. v. 2018, EKL, leg. L. Z. Meng (QBF 13) ” (NME); 1 female: “ China, Yunnan / Honghe, Gulinqin, 585 m, 22 ° 43 ’ 51 N, 103 ° 59 ’ 35 E, 07. V. 2019, leg. L. Z. Meng FIT 5 ” (NME).	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1175FFC8FF55FF3816712928.taxon	diagnosis	Diagnosis. Species easily recognizable by the unique combination of red pronotum (except for narrow dark stripe along mid-line) with black head and elytra (Fig. 7). The structure of the aedeagus in this species is unique. The median lobe is greatly expanded in the middle, weakly sclerotized in this place on right side and has a membranous window.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
054E5D3A1175FFC8FF55FF3816712928.taxon	distribution	Distribution. Previously, this species was known from type locality only (N Vietnam, Nghe An Prov.). This is the first country record for China.	en	Anichtchenko, Alexander (2024): New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia. Acta Biologica Universitatis Daugavpiliensis 24 (2): 259-264, DOI: 10.59893/abud.24(2).010, URL: https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/
