taxonID	type	description	language	source
4180A25A0C285B6AB48EC0820DF3D4A9.taxon	description	Figs 4, 5	en	Liu, Fengjing, Wu, Yayong, Zhang, Jindong, Yang, Guang, Liu, Shuo, Chen, Xue, Chang, Jiang, Xie, Qiang, Cai, Bo (2025): A new species of Diploderma Hallowell, 1861 (Squamata, Agamidae) discovered in the upper Dadu River valley of the Hengduan Mountains, Sichuan, China. ZooKeys 1251: 17-38, DOI: 10.3897/zookeys.1251.153705
4180A25A0C285B6AB48EC0820DF3D4A9.taxon	discussion	Japalura flaviceps authority, date: Zhao 2003: 84, partim in the Aba Tibetan and Qiang Autonomous Prefecture	en	Liu, Fengjing, Wu, Yayong, Zhang, Jindong, Yang, Guang, Liu, Shuo, Chen, Xue, Chang, Jiang, Xie, Qiang, Cai, Bo (2025): A new species of Diploderma Hallowell, 1861 (Squamata, Agamidae) discovered in the upper Dadu River valley of the Hengduan Mountains, Sichuan, China. ZooKeys 1251: 17-38, DOI: 10.3897/zookeys.1251.153705
4180A25A0C285B6AB48EC0820DF3D4A9.taxon	diagnosis	Diagnosis. Diploderma bifluviale sp. nov. can be diagnosed from other Diploderma species by a combination of the following morphological characteristics: (1) body size short, SVL 62.51 ~ 64.47 mm, mean 63.62 mm (sample standard deviation 1.01) in adult males, 63.13 ~ 72.55 mm, mean 68.41 mm (3.53) in adult females; (2) tail short, TAL / SVL 1.41 ~ 1.53, mean 1.48 (0.06) in adult males, 1.37 ~ 1.56, mean 1.46 (0.08) in adult female; (3) head relatively long, HW / HL 0.71 ~ 0.78, mean 0.75 (0.03) in adult males, 0.68 ~ 0.74, mean 0.72 (0.02) in adult females; (4) limbs moderately long, FLL / SVL 0.42 ~ 0.44, mean 0.43 (0.01) in adult males, 0.41 ~ 0.42, mean 0.41 (0.01) in adult females, HLL / SVL 0.64 ~ 0.69, mean 0.66 (0.03) in adult males, 0.64 ~ 0.69, mean 0.65 (0.02) in adult females; (5) MD 49 ~ 54; (6) F 4 S 15 ~ 17, T 4 S 20 ~ 23; (7) tympanum concealed; (8) nuchal and dorsal crests almost continuous, scales of nuchal and dorsal crests enlarged, moderately erected skin fold under nuchal in males in life, weakly erected skin fold under nuchal crest and no skin fold under dorsal crest in females in life; (9) distinct transverse gular fold present; (10) ventral scales of head almost uniform in size, posterior and side ones smaller, all strongly keeled; (11) ventral scales of body strongly keeled; (12) gular spot absent in both sexes; (13) dorsolateral stripes distinct in males, strongly jagged and the upper and lower edges are almost touching each other, pale yellow in life; (14) a series of dark spots or a dark stripe like large wavy between dorsolateral stripes on dorsum; (15) a distinct wide black stripe on shoulder fold region on each side; (16) stripes around eye absent or very indistinct; (17) tongue wheat color in life; (18) fourth toe with claw reaching either the tympanum or the area between shoulder and tympanum when hind limbs adpressed forward (Table 2).	en	Liu, Fengjing, Wu, Yayong, Zhang, Jindong, Yang, Guang, Liu, Shuo, Chen, Xue, Chang, Jiang, Xie, Qiang, Cai, Bo (2025): A new species of Diploderma Hallowell, 1861 (Squamata, Agamidae) discovered in the upper Dadu River valley of the Hengduan Mountains, Sichuan, China. ZooKeys 1251: 17-38, DOI: 10.3897/zookeys.1251.153705
4180A25A0C285B6AB48EC0820DF3D4A9.taxon	description	Description of holotype. Adult male, body relatively small-sized, SVL 62.5 mm; tail short, TAL 88.2 mm; head longer (19.9 mm) than wide (15.6 mm); head depth 12.3 mm; length of tallest nuchal crest 0.5 mm; snout – eye length 8.6 mm; foreleg length 26.2 mm; hindleg length 40.4 mm; toe IV length 10.5 mm; trunk length 28.5 mm; snout moderately long, SEL / HL 0.43. Rostral flat, bordered by six small postrostral scales; dorsal head scales heterogeneous in size, most of them keeled. Number of scales between supraoculars 16; nasal scale approximately oval; internasals 9; loreals 7 / 6, small, unkeeled; mental pentagonal; supralabials 10 / 10, infralabial scales 11 / 13, supratemporals 4 / 3, and middorsal crest scales 54, keeled; gular scales 30 and ventralscales 64; finger IV subdigital lamellae 16 / 17; toe IV subdigital lamellae 21 / 22; nasal supralabial scale rows 1 / 1; suborbital scale rows 3 / 4. Transverse gular fold present; gular pouch present in life; tympanum concealed, covered with small keeled scales; well-developed skin fold under nuchal crest present, vertebral crest continuous between nuchal and dorsal sections; axillary scales much smaller than remaining dorsals; dorsal and ventral scales distinctively keeled exclude scales around eyes and lips; dorsal scales of head, trunk, limbs, and tail heterogeneous in size, ventral scales of head almost ventral scales of head almost uniform in size with few larger scales, posterior and side ones smaller, all strongly keeled; fold present in front of shoulder; fourth toe with claw reaching at tympanum when hindlimbs adpressed forward. Upper and lower edges of dorsolateral stripes are strongly serrated and separated only by a lemon-chiffon dorsal scale at the serrated tip; tail scales all strongly keeled, ventral tail scales slightly larger than dorsal tail scales; fourth toe with claw reaching tympanum when hind limbs adpressed forward (Table 2).	en	Liu, Fengjing, Wu, Yayong, Zhang, Jindong, Yang, Guang, Liu, Shuo, Chen, Xue, Chang, Jiang, Xie, Qiang, Cai, Bo (2025): A new species of Diploderma Hallowell, 1861 (Squamata, Agamidae) discovered in the upper Dadu River valley of the Hengduan Mountains, Sichuan, China. ZooKeys 1251: 17-38, DOI: 10.3897/zookeys.1251.153705
4180A25A0C285B6AB48EC0820DF3D4A9.taxon	distribution	Distribution and natural history. Diploderma bifluviale sp. nov. is currently known to inhabit the semi-arid region of warm-dry valley in the upper reaches of the Dadu River. It is primarily concentrated around Shuangjiangkou, located at the confluence of the Chuosijia River and the Jiaomuzu River, spanning Jinchuan County and Barkam City (Ma’erkang City) within the Aba Tibetan and Qiang Autonomous Prefecture, Sichuan Province, China (Fig. 1). This area is characterized by long hours of sunlight, frequent clear days, distinct wet and dry seasons, and significant diurnal temperature variations (Zhang 1992). The known distribution range of Diploderma bifluviale sp. nov. exhibits higher humidity and greater vegetation coverage compared to that of D. danbaense. The species is known to inhabit altitudes ranging from 2187 to 2314 meters, residing in arid shrublands with small leaves and scattered rock piles, where the shrubs can reach heights of 0.5 – 2 meters (Fig. 5). All specimens were collected between 09: 00 and 18: 00 hr from June to August. During the investigation, it was observed that this species preys on valley insects, this species represents one of the few secondary consumers inhabiting this arid river valley ecosystem. This species is oviparous. Specimens CB 23 JC 16, CIB 119369, CB 23 JC 07, CB 23 JC 03, and CB 23 JC 20 were found to contain 3, 5, 6, 6, and 7 eggs, respectively.	en	Liu, Fengjing, Wu, Yayong, Zhang, Jindong, Yang, Guang, Liu, Shuo, Chen, Xue, Chang, Jiang, Xie, Qiang, Cai, Bo (2025): A new species of Diploderma Hallowell, 1861 (Squamata, Agamidae) discovered in the upper Dadu River valley of the Hengduan Mountains, Sichuan, China. ZooKeys 1251: 17-38, DOI: 10.3897/zookeys.1251.153705
4180A25A0C285B6AB48EC0820DF3D4A9.taxon	etymology	Etymology. The specific epithet bifluviale is derived from the Latin words bi - meaning two, and fluviale, relating to rivers. This name refers to the species’ discovery location near Shuangjiangkou (双江口), which denotes the area around the confluence of the Chuosijia River (绰斯甲河, Chuosi River) and the Jiaomuzu River (脚木足河, Kyom-kyo River) of Sichuan Province, China. We suggest Upper Dadu Mountain Lizard as its English common name and 双江口攀蜥 (Chinese phonetic alphabet: Shuāng jiāng kǒu Pānxī) as its Chinese common name.	en	Liu, Fengjing, Wu, Yayong, Zhang, Jindong, Yang, Guang, Liu, Shuo, Chen, Xue, Chang, Jiang, Xie, Qiang, Cai, Bo (2025): A new species of Diploderma Hallowell, 1861 (Squamata, Agamidae) discovered in the upper Dadu River valley of the Hengduan Mountains, Sichuan, China. ZooKeys 1251: 17-38, DOI: 10.3897/zookeys.1251.153705
