identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
75D5ED8658C55B8D96ADC383659F6D82.text	75D5ED8658C55B8D96ADC383659F6D82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina banbarica X.-M. Li 2025	<div><p>Pseudognaptorina banbarica X.-M. Li sp. nov.</p><p>Type materials.</p><p>Holotype: China • ♂ (MHBU HBU (E) 339867): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.6192&amp;materialsCitation.latitude=31.0235" title="Search Plazi for locations around (long 94.6192/lat 31.0235)">Marxog Township</a>, Banbar County, Xizang / 31°01.410'N, 94°37.152'E / Alt. 4400 m / 2019-VII-31 / Guo-Dong Ren, Ya-Lin Li &amp; Xing-Long Bai leg.  Paratypes: China • 3 ♂♂, 7 ♀♀ (MHBU HBU (E) 339868 –339877): same data as holotype .</p><p>Description.</p><p>Male (Figs 2, 3 A – C). Body length 10.5–11.0 mm, width 4.9–5.0 mm; shiny, black or brownish; antennae, palpi, and tarsi brown.</p><p>Head (Fig. 2 A, B). Anterior margin of clypeus slightly sinuate. Head widest at eye level. Lateral margin of head with pair of projections between antennal base and oculus, brownish red. Genal margin arcuately converging before eyes. Eyes barely protruding beyond contour of head. Vertex flat or slightly convex, with uniform punctures. Antennae (Fig. 2 D) slender and long, reaching beyond pronotal base when posteriorly extended, antennomeres III very long, 3.2 times as long as antennomeres II, antennomeres VIII – X oval, XI spindle-shaped. Length (width) ratio of antennomeres II – XI as follows: 10.9 (10.0): 27.6 (10.0): 13.9 (10.0): 14.3 (10.0): 15.0 (10.0): 15.7 (10.0): 12.6 (12.7): 11.3 (12.6): 11.7 (14.3): 18.0 (14.4).</p><p>Prothorax. Pronotum (Fig. 2 C) transverse, 1.28 times as wide as long, widest in middle, 1.78 times as wide as head. Ratio of width at anterior margin to its maximum width and posterior margin 0.60: 1.00: 0.97. Lateral margins of pronotum arcuately narrowing to anterior margin, bordered along entire length; posterior margin straight; anterior margin slightly emarginate; anterior angles widely obtuse-angled, posterior angles almost rectangular. Surface of pronotum slightly convex between lateral margins, covered with fine dense punctation. Hypomera covered shallow longitudinal wrinkles and granules. Prosternum before procoxae gently sloping. Prosternal process gently sloping behind procoxae, forming obtuse projection.</p><p>Pterothorax. Elytra oblong-oval and convex, 1.37–1.39 times as long as wide, 1.38–1.39 times as wide as pronotum, widest in apical third. Dorsal surface of elytra passing into outer (deflexed) surface without traces of humeral carina. Outer margin of epipleura visible in dorsal view at basal third and apex. Surface of elytra with dense, rather smooth punctation and wrinkles almost vanishing on apical declivity.</p><p>Legs (Fig. 2 E – J). Femora and tibiae moderately thickened. Ratio of length (width) of pro-, meso-, and metatibiae: 43.8 (6.6): 43.2 (7.1): 69.6 (8.0). Protibiae straight with shorter spurs, inner surface of protibiae slightly widened at basal third; mesotibiae slightly curved; metatibiae curved, narrow. Ventral surface of protarsomeres I – III with hairy brush; ventral surface of mesotarsomeres I – II with hairy brush. Ratio of length (width) of metatarsomeres I – IV: 24.0 (9.8): 18.7 (8.8): 15.6 (8.3): 30.8 (8.3).</p><p>Abdomen. Abdominal ventrites rather sparsely covered with minute, pale, recumbent setae.</p><p>Aedeagus. (Fig. 2 K – O) Length of aedeagus 2.31 mm, width 0.42 mm; length of parameres 0.87 mm, width 0.29 mm. Slightly curved to ventral side apically in lateral view. Parameres strongly elongate, widest at base, regularly narrowing towards apex; outer margins slightly curved to ventral side apically in lateral view. Spiculum gastrale as in Fig. 2 K. Posterior margin of abdominal sternite VIII sinuate (Fig. 2 O).</p><p>Female (Figs 3 A, B, 4 D – F). Body larger and wider than male, length 12.6–14.4 mm, width 6.9–7.4 mm. Antennae shorter than male, not posteriorly reaching base of pronotum when posteriorly extended. Pronotum 1.52 times as wide as long, widest in middle, lateral margins subparallel from base to middle and arcuately narrowing toward anterior angles, sides of pronotum slightly convex, 1.79 times as wide as head, with very dense punctation. Elytra oval, more convex than male, 1.34 times as long as wide. Protibial spurs rounded at apex. Distal gonocoxite (Fig. 3 A) rounded apically, densely covered with setae; spiculum ventrale as in Fig. 3 B.</p><p>Diagnosis.</p><p>This new species is morphologically similar to  P. oblonga, but can be distinguished from it by the following male character states: ventral surface of mesotarsomeres I – II with hair brushes, mesotarsomere III with small hairy tuft (ventral surface of mesotarsomeres I – III with hairy brush in  P. oblonga); surface of elytra with dense, rather smooth punctation and regular wrinkle (surface of elytra with fine punctures and irregular wrinkles in  P. oblonga); parameres strongly elongate, 3.0 times as long as wide (parameres elongate, 2.75 times as long as wide in  P. oblonga).</p><p>Etymology.</p><p>This species is named for Banbar County, where the type locality is located.</p><p>Distribution.</p><p>Banbar County, Xizang, China.</p></div>	https://treatment.plazi.org/id/75D5ED8658C55B8D96ADC383659F6D82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
A8DFE6E4244E5B3193101A4C23664A95.text	A8DFE6E4244E5B3193101A4C23664A95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina exsertogena Shi, Ren & Merkl 2005	<div><p>Pseudognaptorina exsertogena Shi, Ren &amp; Merkl, 2005</p><p>Type material examined.</p><p>Holotype: China • ♂ (MHBU XZ 04062848): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=90.5&amp;materialsCitation.latitude=30.1" title="Search Plazi for locations around (long 90.5/lat 30.1)">Yangbajain</a>, Damxung County, Xizang / 30°06'N, 90°30'E / 3700–4100 m / 2004-VI-28 / Yi-Bin Ba &amp; Ai-Min Shi leg.  Paratypes: China • 15 ♂♂, 23 ♀♀ (MHBU XZ 04062843 –04062847,04062849–04062871): same as holotype;  China • 4 ♂♂, 1 ♀ (MHBU XZ 02070511 –02070515): Maizhokunggar County, Xizang / 29°48'N, 91°48'E / 4100 m / 2002-VII-5 / Guo-Dong Ren leg.</p><p>Distribution.</p><p>Xizang, China.</p></div>	https://treatment.plazi.org/id/A8DFE6E4244E5B3193101A4C23664A95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
0675700D897656B6ADE47BC63F8065F4.text	0675700D897656B6ADE47BC63F8065F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina flata Liu & Ren 2009	<div><p>Pseudognaptorina flata Liu &amp; Ren, 2009</p><p>Type material examined.</p><p>Holotype: China • ♂ (MHBU XZ 08071698): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.03333&amp;materialsCitation.latitude=30.116667" title="Search Plazi for locations around (long 99.03333/lat 30.116667)">Batang</a>, Sichuan / 30°07'N, 99°02'E / 3850 m / 2008-VII-16 / Guo-Dong Ren leg.  Paratype: China • 1 ♀ (MHBU XZ 08071697): same data as the holotype .</p><p>Distribution.</p><p>Sichuan, China.</p></div>	https://treatment.plazi.org/id/0675700D897656B6ADE47BC63F8065F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
584201EB90EF5CD7818DFEB0E77574DB.text	584201EB90EF5CD7818DFEB0E77574DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina himalayana X.-M. Li 2025	<div><p>Pseudognaptorina himalayana X.-M. Li sp. nov.</p><p>Type materials.</p><p>Holotype: China • ♂ (MHBU HBU (E) 339877): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=92.318535&amp;materialsCitation.latitude=28.326033" title="Search Plazi for locations around (long 92.318535/lat 28.326033)">Qucho Mountain</a>, Lhunze County, Xizang / 28°19.562'N, 92°19.112'E / Alt. 4824 m / 2019-VII-30 / Xiu-Min Li &amp; Zhao Pan leg.</p><p>Description.</p><p>Male (Figs 5, 6 A – C). Body length 10.3 mm, width 4.9 mm; shiny, brownish; antennae, palpi, and tarsi brown.</p><p>Head (Fig. 5 A, B). Anterior margin of clypeus slightly sinuate. Head widest at eye level. Lateral margin of head with pair of projections between antennal base and oculus, brownish red. Genal margin arcuately converging before eyes. Eyes barely protruding beyond contour of head. Vertex flat or slightly convex, with uniform punctures. Antennae (Fig. 5 D) slender, long, and reaching beyond pronotal base when posteriorly extended; antennomeres VIII – X oval, XI spindle-shaped. Length (width) ratio of antennomeres II – XI as follows: 11.0 (10.0): 27.7 (10.0): 12.5 (10.0): 12.5 (10.0): 12.5 (10.0): 13.4 (10.0): 10.4 (12.5): 10.4 (14.8): 10.4 (15.6): 15.5 (15.8).</p><p>Prothorax. Pronotum (Fig. 5 C) transverse, 1.29 times as wide as long, widest in middle, 1.72 times as wide as head. Ratio of width on anterior margin to its maximum width and posterior margin 0.58: 1.00: 0.92. Lateral margins of pronotum arcuately narrowing in middle, bordered along entire length; posterior margin straight; anterior margin slightly emarginate; anterior angles widely obtuse, posterior angles slightly obtuse. Surface of pronotum very narrowly flattened along lateral margins from base nearly to anterior angles, covered with dense punctation. Hypomera covered shallow longitudinal wrinkles and granules. Prosternum before procoxae gently sloping. Prosternal process gently sloping behind procoxae, forming obtuse projection.</p><p>Pterothorax. Elytra oblong-oval and convex, 1.32–1.33 times as long as wide, 1.38–1.41 times as wide as pronotum, widest in apical third. Dorsal surface of elytra passing into outer (deflexed) surface without traces of humeral carina. Outer margin of epipleura visible in dorsal view at basal third and apex. Surface of elytra with dense, rather smooth punctation and wrinkles almost vanishing on apical declivity.</p><p>Legs (Fig. 5 E – J). Femora and tibiae moderately thickened. Ratio of length (width) of pro-, meso-, and metatibiae: 45.0 (6.5): 50.0 (7.5): 82.5 (8.0). Protibiae narrow, straight with shorter spurs; mesotibiae slightly arcuately curved; metatibiae arcuately curved, narrow. Ventral surface of protarsomeres I – III with hairy brush; ventral surface of mesotarsomeres I – II with hairy brush. Ratio of length (width) of metatarsomeres I – IV: 32.0 (9.5): 17.0 (10.0): 13.5 (9.0): 30.0 (8.5).</p><p>Abdomen. Abdominal ventrites rather sparsely covered with minute, pale, recumbent setae.</p><p>Aedeagus (Fig. 5 K – M). Length of aedeagus 2.30 mm, width 0.48 mm; length of parameres 0.83 mm, width 0.34 mm. Slightly curved to ventral side apically in lateral view. Parameres moderately elongate, regularly narrowing towards apex; outer margins slightly curved to ventral side apically in lateral view.</p><p>Diagnosis.</p><p>This new species is morphologically similar to  P. migana but can be distinguished from it by the following male character states: pronotum 1.34 times as wide as long, lateral margins arcuately narrowed at basal 2 / 3 (pronotum 1.29 times as wide as long, lateral margins of pronotum regularly arcuate in  P. migana); ventral surface of mesotarsomeres I – III with hair brushes (mesotarsomeres I – II with hair brushes, mesotarsomere III with small hairy tuft in  P. migana); parameres moderately elongate, regularly narrowing towards apex, more obtuse from basal half to apex (parameres strongly elongate, widest at base, regularly narrowing towards apex, more acute from basal half to apex in  P. migana).</p><p>Etymology.</p><p>This species is named for the Himalayas, where the type locality of the species is located.</p><p>Distribution.</p><p>Lhunze County, Xizang, China.</p></div>	https://treatment.plazi.org/id/584201EB90EF5CD7818DFEB0E77574DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
E8192B04C2A05158BCC89965DCE3D194.text	E8192B04C2A05158BCC89965DCE3D194.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina Kaszab 1977	<div><p>Genus  Pseudognaptorina Kaszab, 1977</p><p>Pseudognaptorina Kaszab, 1977: 250; Shi et al. 2005: 163; Liu and Ren 2009.</p><p>Type species.</p><p>Pseudognaptorina nepalica Kaszab, 1977, by original designation, by monotypy.</p><p>Generic diagnosis.</p><p>Antennomere VII narrower than VIII; epipleural carina visible in dorsal view at basal part and apex; all tibiae narrow, dilated apically, tarsi slender, ventral surface of male protarsomeres I – III and at least mesotarsomeres I – II with hair brushes; protibial spurs subequal in length; parameres moderately elongate, at least 1.8 times as long as wide; apical part of ovipositor short, 1–1.4 times as long as wide, lobes transverse.</p><p>Distribution.</p><p>Nepal and China.</p></div>	https://treatment.plazi.org/id/E8192B04C2A05158BCC89965DCE3D194	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
894F98F750CC5FA9A6235D3CC48509FE.text	894F98F750CC5FA9A6235D3CC48509FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina migana X.-M. Li 2025	<div><p>Pseudognaptorina migana X.-M. Li sp. nov.</p><p>Type materials.</p><p>Holotype: China • ♂ (MHBU HBU (E) 339879):  Miga Mountain pass, Gongbogyamda County, Xizang / 29°84.105'N, 92°33.422'E / Alt. 4775 m / 2023-VII-18 / Xiu-Min Li &amp; Tong-Yang Guo leg.  Paratypes: China • 10 ♂♂, 20 ♀♀ (MHBU HBU (E) 339880 –339909): same data as holotype;   China • 1 ♂, 1 ♀ (MHBU HBU (E) 339910 –339911):  Miga pass, Gongbogyamda County, Xizang / 29°84.105'N, 92°33.422'E / Alt. 4750 m / 2019-VII-26 / Xiu-Min Li &amp; Zhao Pan leg.</p><p>Description.</p><p>Male (Fig. 7, 8 A – C). Body length 10.5–11.3 mm, width 4.9–5.2 mm; shiny, black or brownish; antennae, palpi, and tarsi brown.</p><p>Head (Fig. 7 A, B). Anterior margin of clypeus slightly sinuate. Head widest at eye level. Lateral margin of head with pair of projections between antennal base and oculus, brownish red. Genal margin arcuately converging before eyes. Eyes barely protruding beyond contour of head. Vertex flat or slightly convex, with uniform punctures. Antennae (Fig. 7 D) slender and long, reaching pronotal base when posteriorly extended, antennomere III very long, 2.4 times as long as antennomeres II, antennomeres VIII – X oval, XI spindle-shaped. Length (width) ratio of antennomeres II – XI as follows: 11.8 (10.0): 28.1 (10.0): 14.5 (10.0): 14.5 (10.0): 14.5 (10.0): 14.5 (10.0): 13.3 (12.0): 13.3 (13.0): 13.3 (14.0): 19.5 (14.0).</p><p>Prothorax. Pronotum (Fig. 7 C) transverse, 1.34 times as wide as long, widest in middle, 1.71 times as wide as head. Ratio of width on anterior margin to its maximum width and posterior margin 0.52: 1.00: 0.92. Lateral margins of pronotum arcuately narrowing to anterior margin, bordered along entire length; posterior margin straight; anterior margin slightly emarginate; anterior angles widely, obtusely angled; posterior angles almost rectangular. Surface of pronotum slightly convex between lateral margins, covered with fine, dense punctation. Hypomera covered shallow, longitudinal wrinkles and granules. Prosternum before procoxae gently sloping. Prosternal process gently sloping behind procoxae, forming obtuse projection.</p><p>Pterothorax. Elytra oblong-oval and convex, 1.38–1.40 times as long as wide, 1.35–1.37 times as wide as pronotum, widest in apical third. Dorsal surface of elytra passing into outer (deflexed) surface without traces of humeral carina. Outer margin of epipleura visible in dorsal view at basal third and apex. Surface of elytra with dense, rather smooth punctation and wrinkles almost vanishing on apical declivity.</p><p>Legs (Fig. 7 E – J). Femora and tibiae moderately thickened. Ratio of length (width) pro-, meso-, and metatibiae: 56.2 (9.1): 55.8 (9.5): 94.1 (11.0). Protibiae straight with shorter spurs, inner surface of protibiae slightly widened at basal third; mesotibiae slightly arcuately curved; metatibiae arcuately curved, narrow. Ventral surface of protarsomeres I – III with hairy brush; ventral surface of mesotarsomeres I – II with hairy brush. Ratio of length (width) of metatarsomeres I – IV: 29.9 (10.0): 19.5 (10.0): 17.1 (10.0): 29.4 (7.9).</p><p>Abdomen. Abdominal ventrites rather sparsely covered with minute, pale, recumbent setae.</p><p>Aedeagus (Fig. 7 K – O). Length of aedeagus 2.34 mm, width 0.46 mm; length of parameres 0.89 mm, width 0.35 mm. Slightly curved to ventral side apically in lateral view. Parameres strongly elongate, widest at base, regularly narrowing towards apex; outer margins slightly curved to ventral side apically in lateral view. Spiculum gastrale as in Fig. 7 K. Posterior margin of abdominal sternite VIII sinuate (Fig. 7 O).</p><p>Female (Figs 8 A, B, 9 D – F). Body larger and wider than male, length 12.1–12.7 mm, width 6.0– 6.4 mm. Antennae shorter than male, not posteriorly reaching base of pronotum when posteriorly extended. Pronotum 1.5 times as wide as long, widest in middle, lateral margins subparallel from base to middle and narrowing toward anterior angles curved, sides of pronotum slightly convex; 1.73 times as wide as head; with very dense punctation. Elytra oval, more convex than male, 1.37 times as long as wide. Protibial spurs with small, pointed at apex. Distal gonocoxite (Fig. 8 A) rounded apically, densely covered with setae; spiculum ventrale as in Fig. 8 B.</p><p>Diagnosis.</p><p>This new species is morphologically similar to  P. himalayana, but can be distinguished from it by the following male character states: pronotum 1.29 times as wide as long, lateral margins of pronotum regularly arcuate (pronotum 1.34 times as wide as long, lateral margins arcuately narrowed at basal 2 / 3 in  P. himalayana); mesotarsomeres I – II with hair brushes, mesotarsomere III with small hairy tuft (ventral surface of mesotarsomeres I – III with hair brushes in  P. himalayana); parameres strongly elongate, widest at base, regularly narrowing towards apex, more acute from basal half to apex (parameres moderately elongate, regularly narrowing towards apex, more obtuse from basal half to apex in  P. himalayana).</p><p>Etymology.</p><p>This species is named after the type locality, Miga Pass.</p><p>Distribution.</p><p>Gongbogyamda County, Xizang, China.</p></div>	https://treatment.plazi.org/id/894F98F750CC5FA9A6235D3CC48509FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
AC264C895AA85A4AA470CA292460D167.text	AC264C895AA85A4AA470CA292460D167.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina nepalica Kaszab 1977	<div><p>Pseudognaptorina nepalica Kaszab, 1977</p><p>Type material examined.</p><p>Paratypes: 1 ♀ (HNHM SMF C 14534): “ Nepal. Expeditionen Jochen Martens, ” “  Gompabei Tarakot / 3300–3400 m / 1970-V-11–16 ”, “  Tagonoides (Pseudognaptorina) nepalica Kasz., det. N. Skopin, 1976 ” ;   1 ♂ (HNHM SMF C 14539), “ Dolpo, Tal der oberen Barbung Khola,  zwischen Terang und Tukot / 4000 m / 1970-VI-19 ” ;   1 ♂ (HNHM SMF C 14537), “ Dolpo,  Weg von Kangar nach Shimen / 4300–4500 m / 1973-VI-18 ” ;   1 ♂ (HNHM SMF C 14532), “ Dolpo, Tal der oberen Barbung Khola,  Charka / 4300–4400 m / 20–25.VI.1973 ”  .</p><p>Distribution.</p><p>Nepal.</p></div>	https://treatment.plazi.org/id/AC264C895AA85A4AA470CA292460D167	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
562B827691345636B165FE3E69513446.text	562B827691345636B165FE3E69513446.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina oblonga X.-M. Li 2025	<div><p>Pseudognaptorina oblonga X.-M. Li sp. nov.</p><p>Type materials.</p><p>Holotype: China • ♂ (MHBU HBU (E) 339912): Lhari County, Xizang / 2013-VII-22 / Xing-Long Bai &amp; Jun-Sheng Shan leg. Paratypes: China • 2 ♂♂, 4 ♀♀ (MHBU HBU (E) 339913 –339918): same data as holotype;   China • 4 ♂♂, 5 ♀♀ (MHBU HBU (E) 339919 –339927): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.40511&amp;materialsCitation.latitude=30.6184" title="Search Plazi for locations around (long 93.40511/lat 30.6184)">Arza Township</a>, Lhari County, Xizang / 30°37.104'N, 93°24.307'E / Alt. 4300 m / 2019-VIII-9 / Guo-Dong Ren, Ya-Lin Li &amp; Xing-Long Bai leg. ;  China • 1 ♂ (MHBU HBU (E) 339928): Lhari County, Xizang / 30°45.225'N, 93°13.162'E / Alt. 4762 m / 2023-VII-17 / Xiu-Min Li &amp; Tong-Yang Guo leg.</p><p>Description.</p><p>Male (Figs 10, 11 A – C). Body length 10.8–11.3 mm, width 5.0– 5.1 mm; shiny, black or brownish; antennae, palpi, and tarsi brown.</p><p>Head (Fig. 10 A, B). Anterior margin of clypeus slightly sinuate. Head widest at eye level. Lateral margin of head with pair of projections between antennal base and oculus, brownish red. Genal margin arcuately converging before eyes. Eyes barely protruding beyond contour of head. Vertex flat or slightly convex, with uniform punctures. Antennae (Fig. 10 D) slender and long, reaching beyond pronotal base when posteriorly extended, antennomere III very long, 3.1 times as long as antennomere II, antennomeres VIII – X oval, XI spindle-shaped. Length (width) ratio of antennomeres II – XI as follows: 10.0 (10.0): 31.3 (10.0): 15.9 (10.0): 15.1 (10.0): 16.2 (10.0): 16.2 (10.3): 15.3 (12.5): 15.3 (12.5): 14.8 (12.5): 19.8 (13.8).</p><p>Prothorax. Pronotum (Fig. 10 C) transverse, 1.40 times as wide as long, widest in middle, 1.67 times as wide as head. Ratio of width on anterior margin to its maximum width and posterior margin 0.57: 1.00: 0.96. Lateral margins of pronotum arcuately narrowing to anterior margin, bordered along entire length; posterior margin straight; anterior margin slightly emarginate; anterior angles widely, obtusely angled; posterior angles almost rectangular. Surface of pronotum very narrowly flattened along lateral margins from base nearly to anterior angles, covered with dense punctation. Hypomera covered shallow longitudinal wrinkles and granules. Prosternum before procoxae gently sloping. Prosternal process gently sloping behind procoxae, forming obtuse projection.</p><p>Pterothorax. Elytra oblong-oval and convex, 1.45–1.49 times as long as wide, 1.44–1.46 times as wide as pronotum, widest at apical third. Dorsal surface of elytra passing into outer (deflexed) surface without traces of humeral carina. Outer margin of epipleura visible in dorsal view at basal third and apex. Surface of elytra with dense, rather smooth punctation and wrinkle almost vanishing on apical declivity.</p><p>Legs (Fig. 10 E – J). Femora and tibiae moderately thickened. Ratio of length (width) of pro-, meso-, and metatibiae: 46.8 (6.5): 64.6 (7.5): 94.1 (8.0). Protibiae straight with shorter spur, inner surface of protibiae slightly widen in basal 1 / 3; mesotibiae slightly arcuately curved; metatibiae arcuately curved, narrow. Ventral surface of protarsomeres I – III with hairy brush; ventral surface of mesotarsomere I – III with hairy brush. Ratio of length (width) of metatarsomeres I – IV segments: 27.5 (9.9): 25.0 (9.1): 17.5 (8.6): 37.5 (9.3).</p><p>Abdomen. Abdominal ventrites rather sparsely covered with minute, pale, recumbent setae.</p><p>Aedeagus (Fig. 10 K – O). Length of aedeagus 2.77 mm, width 0.53 mm; length of parameres 0.88 mm, width 0.32 mm. Slightly curved to ventral side apically in lateral view. Parameres strongly elongate, widest at base, regularly narrowing towards apex; outer margins slightly curved to ventral side apically in lateral view. Spiculum gastrale as in Fig. 10 K. Posterior margin of abdominal sternite VIII sinuate (Fig. 10 O).</p><p>Female (Fig. 11 A, B, 12 D – F). Body larger and wider than male, length 12.1–12.5 mm, width 5.9–6.1 mm. Outer margin of head above base of antennae with widely, obtusely angled emargination, less sharp than in male. Antennae shorter than in male, not posteriorly reaching base of pronotum when posteriorly extended. Pronotum 1.45 times as wide as long, widest in middle, lateral margins subparallel from base to middle and narrowing toward anterior angles arcuately, sides of pronotum slightly convex; 1.72 times as wide as head; with very dense punctation. Elytra oval, more convex than male, 1.32 times as long as wide. Protibial spurs small, pointed at apex. Distal gonocoxite (Fig. 11 A) rounded apically, densely covered with setae; spiculum ventrale as in Fig. 11 B.</p><p>Diagnosis.</p><p>This new species is morphologically similar to  P. banbarica, but can be distinguished from it by the following male character states: ventral surface of mesotarsomeres I – III with hairy brush (ventral surface of mesotarsomeres I – II with hair brushes, mesotarsomere III with small hairy tuft in  P. banbarica); surface of elytra with fine punctures and irregular wrinkles (surface of elytra with dense, rather smooth punctation and wrinkle in  P. banbarica).</p><p>Etymology.</p><p>This species is named from the Latin adjective  “ oblonga ”, in reference to its elongate aedeagus.</p><p>Distribution.</p><p>Lhari County, Xizang, China.</p></div>	https://treatment.plazi.org/id/562B827691345636B165FE3E69513446	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
B87728E7F8045F438EBCBD2E62D3547E.text	B87728E7F8045F438EBCBD2E62D3547E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina obtusa Shi, Ren & Merkl 2005	<div><p>Pseudognaptorina obtusa Shi, Ren &amp; Merkl, 2005</p><p>Type material examined.</p><p>Holotype: China • ♂ (MHBU XZ 04061237): Markam County, Xizang / 29°36'N, 98°24'E / 3800–4000 m / 2004-VI-12 / Ai-Min Shi &amp; Yi-Bin Ba leg. Paratypes: China • 2 ♀♀ (MHBU XZ 04061238 –04061239): same data as holotype .</p><p>Distribution.</p><p>Xizang, China.</p></div>	https://treatment.plazi.org/id/B87728E7F8045F438EBCBD2E62D3547E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
816F668FF9C959E48B8DF7345A4780E0.text	816F668FF9C959E48B8DF7345A4780E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina rectangularis X.-M. Li 2025	<div><p>Pseudognaptorina rectangularis X.-M. Li sp. nov.</p><p>Type materials.</p><p>Holotype: China • ♂ (MHBU HBU (E) 339929): Hongyuan County, Sichuan / 32°01.96'N, 102°01.99'E / Alt. 3451 m / 2021-VII-20 / Xiu-Min Li leg.</p><p>Description.</p><p>Male (Figs 13, 14 A – C). Body length 12.6 mm, width 6.2 mm; shiny, black; antennae, palpi, and tarsi brown.</p><p>Head (Fig. 13 A, B). Anterior margin of clypeus slightly sinuate. Head widest at eye level. Lateral margin of head with pair of projections between antennal base and oculus, brownish red. Genal margin arcuately converging before eyes. Eyes barely protruding beyond contour of head. Vertex flat or slightly convex, with uniform punctures. Antennae (Fig. 13 D) slender, long, and reaching pronotal base when posteriorly extended; antennomere III very long, 2.9 times as long as antennomere II; antennomeres VIII – X oval; XI spindle-shaped. Length (width) ratio of antennomeres II – XI as follows: 10.9 (9.4): 27.1 (10.7): 14.8 (10.0): 16.2 (10.0): 15.5 (10.0): 18.9 (10.0): 15.1 (11.5): 13.9 (13.3): 13.9 (13.3): 15.2 (13.9).</p><p>Prothorax. Pronotum (Fig. 13 C) transverse, 1.52 times as wide as long, widest in middle, 1.84 times as wide as head. Ratio of width on anterior margin to its maximum width and posterior margin 0.55: 1.00: 0.97. Lateral margins of pronotum arcuately narrowing to anterior margin, bordered along entire length; posterior margin straight; anterior margin slightly emarginate; anterior angles widely, obtusely angled; posterior angles almost rectangular. Surface of pronotum very narrowly flattened along lateral margins from base nearly to anterior angles, covered with dense punctation. Hypomera covered shallow longitudinal wrinkles and granules; inner part covered with longitudinal rugae. Prosternum gently sloping before procoxae. Prosternal process gently sloping behind procoxae, forming obtuse projection.</p><p>Pterothorax. Elytra oblong-oval and convex, 1.31 times as long as wide, 1.43 times as wide as pronotum, widest at apical third. Dorsal surface of elytra passing into outer (deflexed) surface without traces of humeral carina. Outer margin of epipleura visible in dorsal view at basal third and apex. Surface of elytra with dense, rather smooth punctation and wrinkles almost vanishing on apical declivity.</p><p>Legs (Fig. 13 E – J). Femora and tibiae moderately thickened. Ratio of length (width) of pro-, meso-, and metatibiae: 57.2 (9.2): 59.1 (9.4): 77.9 (10.3). Protibiae straight with shorter spur, inner surface of protibiae nearly flat; mesotibiae slightly curved; metatibiae slightly curved. Ventral surface of protarsomeres I – III with hairy brush; ventral surface of mesotarsomeres I – II with hairy brush. Ratio of length (width) of metatarsomeres I – IV: 34.5 (9.7): 22.5 (10.0): 22.8 (10.0): 31.9 (7.7).</p><p>Aedeagus (Fig. 13 K – O). Length of aedeagus 2.18 mm, width 0.46 mm; length of parameres 0.79 mm, width 0.31 mm. Slightly curved to ventral side apically in lateral view. Parameres strongly elongate, widest at base, regularly narrowing towards apex; outer margins slightly curved to ventral side apically in lateral view. Spiculum gastrale as in Fig. 13 K. Posterior margin of abdominal sternite VIII sinuate (Fig. 13 O).</p><p>Diagnosis.</p><p>This new species is morphologically similar to  P. flata, but can be distinguished from it by the following male character states: pronotum transverse, 1.52 times as wide as long, surface flatted, posterior angles nearly rectangular (pronotum transverse, 1.36 times as wide as long, surface explanate and slightly concave, posterior angles obtusely rounded in  P. flata); ventral surface of mesotarsomeres I – II with hair brushes, mesotarsomere III with small hairy tuft (ventral surface of mesotarsomeres I – III with hair brushes in  P. flata).</p><p>Etymology.</p><p>This species is named from the Latin adjective  “ rectangularis ”, in reference to its sub-rectangular prothorax.</p><p>Distribution.</p><p>Hongyuan County, Sichuan, China.</p></div>	https://treatment.plazi.org/id/816F668FF9C959E48B8DF7345A4780E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
3838EE234D575E72AD226A6671BB5D25.text	3838EE234D575E72AD226A6671BB5D25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudognaptorina reni X.-M. Li 2025	<div><p>Pseudognaptorina reni X.-M. Li sp. nov.</p><p>Type materials.</p><p>Holotype: China • ♂ (MHBU HBU (E) 339930): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.154335&amp;materialsCitation.latitude=32.799667" title="Search Plazi for locations around (long 95.154335/lat 32.799667)">Vajra Mountain pass</a>, Zadowa County, Qinghai / 32°47.98'N, 95°09.26'E / Alt. 4718 m / 2019-VII-26 / Guo-Dong Ren &amp; Yi-Ping Niu leg.  Paratypes: China • 10 ♂♂, 20 ♀♀ (MHBU HBU (E) 339931 –339960): same data as holotype;   China • 3 ♂♂, 10 ♀♀ (MHBU HBU (E) 339961 –339973): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.72183&amp;materialsCitation.latitude=32.525" title="Search Plazi for locations around (long 94.72183/lat 32.525)">Konge township</a>, Bachen County, Xizang / 32°31.50'N, 94°43.31'E / Alt. 4556 m / 2022-VII-4 / Guo-Dong Ren &amp; Yi-Ping Niu leg.</p><p>Description.</p><p>Male (Figs 15, 16 A – C). Body length 13.6–14.1 mm, width 6.30–6.58 mm; black, slightly shiny, oval-oblong.</p><p>Head (Fig. 15 A, B). Anterior margin of clypeus slightly sinuate. Head widest at eye level. Lateral margin of head with pair of projections between antennal base and oculus, brownish red. Genal margin arcuately converging before eyes. Eyes barely protruding beyond contour of head. Vertex flat or slightly convex, with uniform punctures. Antennae (Fig. 15 D) slender and long, reaching pronotal base when posteriorly extended, antennomere III very long, 2.7 times as long as antennomere II, antennomeres VIII – X oval, XI spindle-shaped. Length (width) ratio of antennomeres II – XI as follows: 12.5 (10.0): 33.4 (10.0): 16.6 (10.0): 15.7 (10.0): 14.8 (10.0): 16.4 (10.0): 14.1 (12.9): 14.1 (14.1): 14.1 (14.7): 18.8 (15.5).</p><p>Prothorax. Pronotum (Fig. 15 C) transverse, 1.50 times as wide as long, widest in middle, 1.72 times as wide as head. Ratio of width on anterior margin to its maximum width and posterior margin 0.54: 1.00: 0.95. Lateral margins of pronotum arcuately narrowing to anterior margin, bordered along entire length; posterior margin straight; anterior margin slightly emarginate; anterior angles widely, obtusely angled; posterior angles almost rectangular. Surface of pronotum slightly convex between lateral margins, very narrowly flattened along lateral margins from base nearly to anterior angles, covered with fine, dense punctation. Hypomera covered shallow, longitudinal wrinkles and granules. Prosternum before procoxae gently sloping. Prosternal process gently sloping behind procoxae, forming obtuse projection.</p><p>Pterothorax. Elytra oblong-oval and convex, 1.43–1.48 times as long as wide, 1.39–1.41 times as wide as pronotum, widest at apical third. Dorsal surface of elytra passing into outer (deflexed) surface without traces of humeral carina. Outer margin of epipleura visible in dorsal view at basal third and apex. Surface of elytra with dense, rather smooth punctation and wrinkles almost vanishing on apical declivity.</p><p>Legs (Fig. 15 E – J). Femora and tibiae moderately thickened. Ratio of length (width) of pro-, meso-, and metatibiae: 56.0 (8.6): 49.6 (7.6): 68.0 (8.6). Protibiae straight with shorter spur, inner surface of protibiae slightly widened at basal third; mesotibiae slightly arcuately curved; metatibiae curved, narrow. Ventral surface of protarsomeres I – III with hairy brush; ventral surface of mesotarsomeres I – II with hairy brush. Ratio of length (width) of metatarsomeres I – IV: 38.7 (11.5): 21.1 (10.0): 19.6 (10.0): 31.1 (10.0).</p><p>Abdomen. Abdominal ventrites rather sparsely covered with minute, pale, recumbent setae.</p><p>Aedeagus (Fig. 15 K – O). Length of aedeagus 2.31 mm, width 0.51 mm; length of parameres 0.76 mm, width 0.33 mm. Slightly curved to ventral side apically in lateral view. Parameres strongly elongate, widest at base, regularly narrowing towards apex; outer margins slightly curved to ventral side apically in lateral view. Spiculum gastrale as in Fig. 15 K. Posterior margin of abdominal sternite VIII sinuate (Fig. 15 O).</p><p>Female (Figs 16 A, B, 17 D – F). Body larger and wider than male, length 14.1–15.0 mm, width 7.0– 7.7 mm. Antennae shorter than in male, reaching base of pronotum when posteriorly extended. Pronotum 1.56 times as wide as long, widest in middle, lateral margins subparallel from base to middle and arcuately narrowing toward anterior angles, sides of pronotum slightly convex; 1.83 times as wide as head, with very dense punctation. Elytra oval, more convex than in male, 1.37 times as long as wide. Protibial spurs small, pointed at apex. Distal gonocoxite (Fig. 16 A) rounded apically, densely covered with setae; spiculum ventrale as in Fig. 16 B.</p><p>Diagnosis.</p><p>This new species is morphologically similar to  P. exsertogena, but can be distinguished from it by the following male character states: pronotum 1.50 times as wide as long (pronotum 1.28 times as wide as long in  P. exsertogena); surface of elytra with fine punctures and without regular wrinkles (surface of elytra with fine punctures and irregular wrinkles in  P. exsertogena).</p><p>Etymology.</p><p>This species is named after Prof. Guo-Dong Ren, in recognition to his contributions in collecting specimens of  Pseudognaptorina .</p><p>Distribution.</p><p>Zadowa and Bachen Counties, Xizang, China.</p></div>	https://treatment.plazi.org/id/3838EE234D575E72AD226A6671BB5D25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Xiu-Min;Zhang, Bao-Yue;Li, Ji-Gang;Pan, Zhao	Li, Xiu-Min, Zhang, Bao-Yue, Li, Ji-Gang, Pan, Zhao (2025): Systematic review of the genus Pseudognaptorina Kaszab, 1977 (Coleoptera, Tenebrionidae, Blaptinae, Blaptini) from the Qinghai-Xizang Plateau, with description of six new species. ZooKeys 1234: 19-46, DOI: 10.3897/zookeys.1234.137739
