identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
68D09C20397D5422834D382B1CD9F78F.text	68D09C20397D5422834D382B1CD9F78F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa)	<div><p>Plusiocampa (Plusiocampa) spp. indet.</p><p>Fig. 21</p><p>New material examined.</p><p>Bulgaria: Western Rhodopes Mts . • 1 ex., Asenovgrad District, village Dobrostan, cave Lednitsata, 05 Nov. 2008, leg. S. Deleva, I. Naydenov • 2 ex., Asenovgrad District, village Dobrostan, cave Ivanova voda, 1320 m alt., 04 Feb. 2006, leg. B. Petrov, N. Simov • 14 ex., Velingrad District, cave Lepenitza, clay, 25 Nov. 1993, leg. B. Petrov, P. Stoev • 3 ex., same locality, 21 May 2011, leg. B. Petrov, N. Toshkova • 1 ex., Smolyan District, village Maglishta, cave Dupkata, 813 m alt., rotten log, 16 Jul. 2006, leg. B. Petrov • 1 ex., Gotse Deltshev District, village Ribnovo, cave Skoka, 1100 m alt., 18 Nov. 2006, leg. B. Petrov • 2 ex., Slavyanka Mts., cave Snyagova dupka below Gotsev vrah, 2160 m alt., 26 Jul. 2004, leg. B. Petrov • 2 ex., Vitosha Mts., Sofia District, village Bosnek, cave Zhivata voda, 1000 m alt., 27 Nov. 2002, leg. B. Petrov, P. Beron, V. Beshkov; Predbalkan • 1 ex., Lovech District, village Mikre, cave Mandrata, 365 m alt., underground stream, 11 Mar. 2007, leg. B. Petrov, I. Borisov; 1 female, Lovech District, village Bezhanovo, cave Alchashkata peshtera, 17 Jul. 1985, leg. P. Beron, B. Garev; Danubian Plane • 2 females, Pleven District, village Bohot, cave Kirov vartop, 06 Apr. 1984, leg. P. Beron; Eastern Stara planina Mts . • 5 ex., Sliven District, near Sliven City, disused mine gallery in Karandila Nature Park, 04 Dec. 1991, leg. P. Stoev • 5 males, 1 juv., Kotel District, near town Kotel, cave Prikazna, 31 Oct. 2002, leg. P. Beron.</p><p>Remarks.</p><p>Unfortunately, the examined material was in very poor condition and did not allow identification to the species level. Particularly noteworthy are the specimens from the Snyagova Dupka Cave in the Slavyanka Mountains, as they may represent another, yet undescribed, new species.</p></div>	https://treatment.plazi.org/id/68D09C20397D5422834D382B1CD9F78F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
F1C6D018EC1D50A6B00C1FEFB0497C69.text	F1C6D018EC1D50A6B00C1FEFB0497C69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa) arbanisiensis Bareth & Conde 2001	<div><p>Plusiocampa (Plusiocampa) arbanisiensis Bareth &amp; Condé, 2001</p><p>Figs 1, 21</p><p>Plusiocampa (Plusiocampa) arbanisiensis Bareth &amp; Condé, 2001: 23, figs 7 a – g, 8 a, b. Type locality: Lyaskovskata pestera, village Arbanasi, Bulgaria.</p><p>Plusiocampa arbanisiensis: Beron, 2015: 92.</p><p>New material examined.</p><p>Bulgaria • 1 male, Predbalkan: Veliko Tarnovo District, Village Arbanasi, Arbanashkata (Lyaskovskata) Cave, 1 Mar. 2011, leg. B. Petrov • Eastern Stara Planina Mts: 3 females, Kotel District, Zlosten, Cave Lednitsata, 726 m alt., stones, clay, 11 Mar. 2011, leg. B. Petrov, S. Deleva, I. Alexandrova • Shumensko Plateau: 1 ex., Shumen District, Cave Biserna ( Zandana), 395 m alt., guano, rotten logs, 29 Jan. 2010, leg. A. Hubancheva, I. Borissov • Strandzha Mts.; 1 ex., Burgas District, Village Stoilovo, Cave Golyama vapa, 255 m alt., under stones, guano-clay, 06 Aug. 2006, leg. B. Petrov, T. Stoyanov .</p><p>Diagnosis.</p><p>(Bareth and Condé 2001; Sendra et al. 2020 b): Epicuticle smooth at high magnifications. Antennae with 23–26 antennomeres; five simple olfactory chemoreceptors in cupuliform organ; distal whorl of 8–11 gouge sensilla that are 25 µm long; slightly protruding frontal process with non-tuberculated to slightly tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 (2 + 1) la, 2 + 2 lp macrosetae; long, barbed notal macrosetae; abundant barbed notal clothing setae. Two ventral tibial macrosetae; subequal claws (ratio 1.05–1.1) with large but narrow lateral crests, posterior claw with a small backward overhang (Fig. 1 a). Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergites IV – V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 (2 + 1) la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7 (Fig. 1 b), urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae (Fig. 1 c); stylus setae with a few distal barbs (Fig. 1 d). Male with glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female appendages similar to those of male (Fig. 1 b).</p><p>Habitat and distribution.</p><p>Slightly cave-adapted species that has hitherto been found in caves from two geographically distant karst regions in the Balkan Peninsula – the Predbalkan in Bulgaria (Lyaskovskata Cave, the type locality) and the Karst Plateau in Slovenia (two caves near the town of Sežana). Here, we re-establish the species from its type locality (Arbanashkata Cave, also known as Lyaskovskata Cave), as well as from three additional caves in Bulgaria located in distinct geographical regions: the Eastern Stara Planina Mountains, the Shumen Plateau, and the Strandzha Mountains. As a result, the known range of the species is extended by approximately 200 km to the southeast and 100 km to the northeast of the type locality (Fig. 21).</p></div>	https://treatment.plazi.org/id/F1C6D018EC1D50A6B00C1FEFB0497C69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
ED7BBBD2F224565FBBA84A348B09B647.text	ED7BBBD2F224565FBBA84A348B09B647.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa) balcanica Sendra & Stoev & Selfa 2025	<div><p>Plusiocampa (Plusiocampa) balcanica Sendra &amp; Stoev sp. nov.</p><p>Figs 11, 12, 21</p><p>Etymology.</p><p>The specific epithet derives from the type locality, which is situated in Stara Planina (= Balkan) Mts. The specific epithet is a noun in the genitive case.</p><p>Material examined.</p><p>Holotype • male deposited in the Natural History Museum of Sofia, Non-insect Invertebrates – Id. 10835 NMNHS. Bulgaria: Central Stara planina Mts.: Karlovo District, village Hristo Danovo, cave Yulen Ere (Vodnata peshtera), 950 m alt., 27 Sep. 1997, leg. P. Beron &amp; T. Ivanova ; Paratypes • 3 females from the same locality, date, and collectors deposited in the Natural History Museum of Sofia, Non-insect Invertebrates Id. 10836-10838 NMNHS • 1 female from the same locality, date, and collectors deposited in MCNB - MZB 2025-7269; and 1 male from the same locality, date, and collectors deposited in MUVHN - ZE 4330 .</p><p>Description.</p><p>Body. Body length 5.5 mm and 6 - mm (holotype) males, 6.5 mm to 7 mm females. Epicuticle smooth under optical microscope; slender body with smooth or with one or two tiny distal barbs on clothing setae.</p><p>Head. Antennae have 37, 40, 40, and 41 antennomeres in four completed and intact antennae; antennae ~ 1.1-0.7 times longer than the body’s length, with medial antennomeres almost three times longer than wide and apical antennomere more than two times longer than wide. Cupuliform organ, occupying 1 / 9 of the total length in the last antennomere, has about 6–7 complex olfactory chemoreceptors. Distal and central antennomeres have two whorls of macrosetae with a few distal barbs and scattered smooth thin setae, in addition to a single distal whorl of 10‒14 long gouge sensilla (35-45 µm long) and up to two very short coniform sensilla. Proximal antennomeres have typical trichobothria, plus a small coniform sensillum on the third antennomere in ventral position. The frontal process has three slightly tuberculate smooth macrosetae; the three-insertion line macrosetae with a few distal barbs, with length ratios of a / i / p / x are 41 / 47 / 35 / 55, respectively, in the holotype. Labial palps are suboval with small latero-external sensillum, three guard setae, up to six setae on the anterior border, and up to 95 neuroglandular setae in the larger paratype.</p><p>Thorax. Thoracic macrosetae distribution: pronotum with 1 + 1 ma, 2 + 2 la 3-4, 1 + 1 lp 1, 3 macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la 1-3, 2 + 2 lp 2-3 and 1 + 1 ma, 1 + 1 la 1, 2 + 2 lp 2-3 long macrosetae. All macrosetae long with thin barbs along the basal two-thirds of their length; thin marginal setae barbed on distal portion double the length of clothing setae. Legs elongated, and the metathoracic legs reach the eighth abdominal segment. Tibia longer than the femur or tarsus; femur / tibia / tarsus plus claws: 0.10 / 0.11 / 0.99 mm in paratype 7 mm in lengths. Femora I – III with one long dorsal macrosetae, and three ventral macrosetae on tibia III. Calcars have long barbs almost all over. Two ventral rows of slightly barbed on medial portion setae on tarsus; end part of the tarsus with two dorsal smooth long setae plus two ventral long barbed setae. Claws are very unequal in size with posterior claw longer than the anterior claw (1.6‒1.8: ratio posterior / anterior). Claws have large lateral crests well developed in posterior claw with a large backward overhang. Pretarsus has similar long smooth setiform lateral processes on posterior claw but shorter on the smaller anterior claw (Fig. 11 a, b).</p><p>Abdomen. Distribution of abdominal macrosetae on tergites: 1 + 1 post macrosetae on I – III (0 + 0 on I – II urotergites in male paratype), 1 + 1 la, 5 + 5 post on IV – V urotergites, 2 + 2 la, 5 + 5 post on VI – VII; 6 + 6 post on VIII, and 8 + 8 post macrosetae on IX abdominal segment. All tergal abdominal macrosetae long and well-differentiated with thin barbs along most of its length. Urosternite I with 7 + 7 (7 + 1 + 7) well-barbed macrosetae (Fig. 15 a, b); urosternites II to VII with 5 + 5 macrosetae; urosternite VIII with 2 + 2 macrosetae; urosternal macrosetae with long barbs along distal half to two third. Stylus with an apical almost smooth setae with one large and one small basal barb, subapical seta with a few short barbs and ventromedial seta with some long barbs on half distal part (Fig. 12 c). All cerci broken, the most complete is 4.8 mm long with a basal article follow for eight primary articles; each primary article bear up to four untidy whorls of macrosetae with short thin barbs on half distal part in addition to several smooth long setae and the characteristic apical whorl of very thin short setae smooth or with one or two tiny barbs.</p><p>Secondary sex characters. Female urosternite I with large subcylindrical appendages, bearing each up to 30 glandular a 1 setae in a distal field (Fig. 12 a). Male with larger subcylindrical appendages than the female, bearing each up to 45 glandular setae a 1, in addition posterior portion of the urosternite with a few glandular g 1 setae, up to 14 on holotype (Fig. 12 b) with bears a whorl of 17 short setae around the terminal hole of the genital papilla.</p><p>Taxonomic position.</p><p>According to Sendra et al. (2020 b), the taxonomy of the species of Plusiocampa needs thorough revision and clarification. However, here we propose a new species: Plusiocampa (Plusiocampa) balcanica sp. nov., which is distinguished from the congeners ( Plusiocampa s. str.) by having two reliable taxonomic characters: very uneven pretarsal claws with a 1.6‒1.8: posterior / anterior ratio, and the lack of medial posterior macrosetae on the mesonotum and metanotum. There is yet another species that possesses this combination of characters: Plusiocampa (Plusiocampa) ternovensis Sendra &amp; Borko, 2020, found in three caves on the edge of a karst plateau north of the town of Ajdovščina, in Slovenia (Sendra et al. 2020 b).</p><p>However, P. (P.) balcanica sp. nov. can be distinguished from P. (P.) ternovensis by the following characters: 2 + 2 la macrosetae on the pronotum (vs. 4 + 4 in P. (P.) ternovensis; three ventral tibial macrosetae (vs. only one); 5 + 5 post macrosetae on urosternites V – VII (vs. 4 + 4); and larger, thicker appendages in both males and females (vs. much thinner ones in P. (P.) ternovensis .</p><p>Habitat and distribution.</p><p>This cave-adapted species is currently known only from a single cave, Yulen Ere (Vodnata Peshtera), near the village of Hristo Danovo in the Central Stara Planina Mts (= Balkan) in Bulgaria (Fig. 21). According to Beron (2015), the cave is situated at an altitude of 970 m and has a length of 227 m, with a denivelation of + 17 m. In addition to Plusiocampa (P.) balcanica sp. nov., the following species have been recorded in the cave: Araneae: Meta menardi (Latreille, 1804), Antrohyphantes balcanica (Drensky, 1931); Pseudoscorpiones: Chthonius sp.; Diptera: Triphleba antricola (Schmitz, 1918); Chiroptera: Rhinolophus ferrumequinum (Schreber, 1774), Rhinolophus hipposideros (Bechstein, 1800) (Beron 2015) . Two other caves, Zlatnata Peshtera and Mazata, are known from the same area. It is, therefore, likely that the new species also occurs in these caves.</p></div>	https://treatment.plazi.org/id/ED7BBBD2F224565FBBA84A348B09B647	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
C3F643A93FA5523ABBB024D873FED0E9.text	C3F643A93FA5523ABBB024D873FED0E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa) beroni Bareth & Conde 2001	<div><p>Plusiocampa (Plusiocampa) beroni Bareth &amp; Condé, 2001</p><p>Figs 2, 3, 4, 21</p><p>Plusiocampa (Plusiocampa) beroni Bareth &amp; Condé, 2001: 16, figs 3 b, 4 a – e. Type locality: Cave Magura, village Rabisha, Vidin District, Bulgaria.</p><p>Plusiocampa ( Stygiocampa [sic!]) aff. beroni: Beron et al., 2011 610.</p><p>Plusiocampa ( Stygiocampa [sic!]) beroni: Beron, 2015: 92.</p><p>Plusiocampa ( Stygiocampa [sic!]) cf. beroni: Beron, 2015: 92.</p><p>New material examined.</p><p>Bulgaria: Western Rhodopes Mts . • 13 ex., Gotse Deltsev District, Village Ribnovo, Cave Manuilovata, 1182 m alt., 30 Apr. 1994, leg. B. Petrov • 2 ex., Gotse Deltchev District, Village Gospodintsi, Cave Salievata, 02 Nov. 1994, leg. B. Petrov • 4 ex., Pazardzhik District, Peshtera town, Kupena Reserve, Cave Novata Peshtera, 540 m alt., 17 Oct. 1992, leg. B. Petrov • 3 ex., same locality and collector, 07 Nov. 2004; 6 ex., same locality, 27 Jun. 1986, leg. P. Beron • 4 ex., Kupena Reserve, Cave Vodnata Peshtera, 27 Jun. 1986, 890 m alt., leg. P. Beron • 4 ex., same locality, 02 Jul. 2006, leg. P. Stoev, B. Petrov • 1 ex., Kupena Reserve, Village Ravnogor, Cave Pavla, 1322 m alt., 01 Jul. 2006, leg. B. Petrov and P. Stoev; Vlahina Mountain • 3 ex., Blagoevgrad Distr., Village Logodazh, Cave Boichova Dupka, 14 Sep. 1994, leg. B. Petrov; Predbalkan • 2 males, 3 females, Lovetch District, Village Dragana, Sinyoto Ezero Cave, 25 Aug. 1985, leg. B. Garev.</p><p>Diagnosis.</p><p>(Bareth and Condé 2001): Epicuticle smooth at high magnifications (Fig. 4 b). Antennae with 26–29 antennomeres; distal whorl of 24–26 gouge sensilla that are 30 µm long (Fig. 2 a – c); 6–8 complex olfactory chemoreceptors on cupuliform organ, each with a prominent central protrusion surrounding a single fold (Fig. 2 d); border of cupuliform organ with a whorl of presumably glandular setae long and short, long ones with a shallow distal part with an apical opening (Fig. 2 d, e); non-protruding frontal process with non-tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae (Fig. 3 a, b; metanotum with 1 + 1 ma, 1 + 1 (2 + 1) la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. Two ventral tibial macrosetae; unequal claws (ratio 1.10-1.30) with large lateral crests, posterior claw with a backward overhang (Fig. 3 c – f). Urotergites I – II with 1 + 1 post macrosetae; urotergites III with 2 + 2 post macrosetae (1 + 1 la, 1 + 1 post on one specimen from Novata Peştera, western Rhodope); urotergite IV with 1 + 1 la, 4 + 4–5 + 5 post macrosetae; urotergite V with 1 + 1–2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2–3 + 3 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae (Fig. 4); stylus setae with a few barbs (Fig. 4 b). Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae (adults from Gruffre Padez, Sofia with short, thick appendages with a large area of glandular a 1 setae); female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicle wheels with 2 turns of 50 µm long and 5 µm diameter filament.</p><p>Habitat and distribution.</p><p>This species was originally described from Magura Cave near the village of Rabisha in northwestern Bulgaria (Bareth and Condé 2001). In the same publication, the authors also reported it from another cave in the region – Varkan near the village of Druzhba. Two other records – though uncertain – have been reported from Padezh Cave near the village of Breze and Novata Cave near the town of Peshtera.</p><p>Here, we confirm the species’ presence in caves in the Peshtera area and report its occurrence in other caves in the Rhodopes along the Mesta Valley, as well as in one cave in the Vlahina Mountain and one in the Predbalkan region. Thus, the species appears to be distributed across parts of western Bulgaria, although its range remains highly fragmented, with confirmed presence in only 10 caves to date (Fig. 21).</p><p>Despite being found exclusively in subterranean habitats, the species does not exhibit strong adaptations to the cave environment.</p></div>	https://treatment.plazi.org/id/C3F643A93FA5523ABBB024D873FED0E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
809483A18F3C5278B4FECB402B2EA0D6.text	809483A18F3C5278B4FECB402B2EA0D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa) bulgarica Silvestri 1931	<div><p>Plusiocampa (Plusiocampa) bulgarica Silvestri, 1931</p><p>Figs 5, 6, 7, 8, 9, 21</p><p>Plusiocampa (Plusiocampa) bulgarica Silvestri, 1931: 103, figs vi – viii. Type locality: Cave Charamijskata, village Trigrad, Bulgaria.</p><p>Plusiocampa bulgarica: Bareth, 1974: 657.</p><p>Plusiocampa ( Stygiocampa [sic!]) bulgarica: Beron et al., 2011: 610.</p><p>Plusiocampa ( Stygiocampa [sic!]) bulgarica: Beron, 2015: 92.</p><p>New material examined.</p><p>Bulgaria: Western Rhodopes Mts . • 2 ex., Asenovgrad District, Village Mostovo, Cave Gargina Dupka, 900 m alt., 14 Mar. 1992, leg. B. Petrov • 2 ex., same locality, 19 Apr. 1993, leg. P. Stoev • 2 males, same locality, guano, rocks, 26 Feb. 2009, leg. S. Deleva, A. Pavlova • 1 female, same locality, pitfall traps, 10 Apr. 2010 – 18 Apr. 2011, leg. S. Deleva • 1 ex., Asenovgrad District, Village Belitsa, Shepran Dupka Cave, 850 m alt., 08 Oct. 2005, leg. B. Petrov, T. Ivanova, I. Borisov • 17 ex., Asenovgrad District, Village Orehovo, Cheloveshkata Peshtera cave, 1110 m alt., 14–15 Nov. 1992, leg. P. Stoev • 3 ex., same locality, 07 Jun. 2007, leg. B. Petrov, P. Stoev • 2 ex., Laki District, Agovski Kamak area, Village Dryanovo, Cave Biserna, 1140 m alt., 09 Oct. 2005, leg. B. Petrov, I. Borisov, T. Ivanova • 1 female, 1 juvenile, Laki District, Village Dryanovo, Lisek Cave, 15 Nov. 2009, leg. S. Deleva • 3 females, 1 male, 1 juv., Laki District, Staro Selo close to Village Dryanovo, Cave Barzibogovata, 1110 m alt., 20 Oct. 2005, leg. I. Borisov • 10 ex., Dospat District, Village Borino, Hut Orphei, Cave Izvora, 20 Mar. 1993, leg. T. Ivanova • 1 ex., Chepelare District, Village Zabardo, Cave Tamnata Dupka, 1250 m alt., clay, 20 Mar. 2004, leg. B. Petrov, T. Stoyanov • 7 ex., same locality, 22 May 2010, leg. B. Petrov, P. Beron, T. Georgieva, S. Stoycheva • 3 ex., town Chepelare, Cave Samurskata, 28 Nov. 1994, leg. B. Petrov • 4 ex., Devin District, Village Trigrad, Cave Suhodolska, 1400 m alt., 13 Jul. 1997, leg. B. Petrov • 4 ex., Devin District, Village Yagodina, Cave Gorna Karanska, 1100 m alt., 04 Aug. 1997, leg. B. Petrov • 66 ex., same locality, trap at the end of the cave, 12 Mar. 1998 leg. B. Petrov • 2 ex., Devin District, Village Yagodina, Sanchova Dupka Cave, 990 m alt., clay-guano, 17 Sep. 2005, leg. B. Petrov • 1 ex., Devin District, Village Kesten, Cave Forgovo 1, 1300 m alt., 01 Oct. 2000, leg. B. Petrov • 2 ex., Devin District, Village Breze, Cave Byaloto Kamene, 1230 m alt., 04 Mar. 2007, leg. B. Petrov • 2 ex., Smolyan District, Village Mogilitsa, Cave Uhlovitsa, 1000 m alt., 06 Aug. 1999, leg. B. Petrov, V. Beshkov • 2 ex., Smolyan District, Village Progled, Cave Sbirkovata, 1350 m alt., 07 Aug. 1999, leg. B. Petrov, V. Beshkov • 1 male, same locality, 21 May 2017, leg. D. Georgiev • 10 ex., Smolyan District, Village Borikovo, Cave Borikovskata, 1120 m alt., clay, rotten wood, 19 Mar. 2004, leg. B. Petrov • 4 ex., Smolyan District, Village Borikovo, Borikovskata Peshtera Cave, 01 Jun. 2003, leg. T. Ivanova • 2 ex., Smolyan District, Village Gela, Cave Lednitsata, 1620 m alt., 19 Sep. 2005, leg. B. Petrov • 1 ex., Smolyan District, Village Dolno Vlahovo, Cave Potoka, 1150 m alt., 16 Jul. 2006, leg. B. Petrov • 2 ex., Smolyan District, Village Mogilitsa, Cave Byala Voda, 1045 m alt., 13 Jul. 2006, leg. B. Petrov • 1 ex., Smolyan District, Village Polkovnik Serafimovo, cave near Pametnika, 1080 m alt., 14 Jul. 2006, leg. B. Petrov • 4 ex., Smolyan District, Pamporovo, Mecha Dupka Cave, 1560 m alt., under stones in clay, 19 Oct. 2006, leg. B. Petrov, T. Stoyanov • 3 ex., Rudozem District, Village Boevo, Cave Boevskata, 1065 m alt., 14 Jul. 2006, B. Petrov leg.; Eastern Rhodopes Mts . • 1 ex., Dzhebel District, Village Stremtsi, Gyaurhambar Cave, 520 m alt., 08 Jun. 2007, leg. P. Stoev, B. Petrov; Pirin Mts . • 3 ex., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.3235&amp;materialsCitation.latitude=41.72768" title="Search Plazi for locations around (long 23.3235/lat 41.72768)">Cave Aleko</a>, 1692–1760 m alt., 41.72768 ° N, 23.32350 ° E, 26 Jun. 2002, leg. B. Petrov • 1 ex., same locality, 20. 11. 2011, leg. N. Toshkova • 1 ex., Banski Suhodol, Cave BS 25, BFSp <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.40306&amp;materialsCitation.latitude=41.79817" title="Search Plazi for locations around (long 23.40306/lat 41.79817)">No.</a> 2557, 41.79817 ° N, 23.40306 ° E, 2310 m alt., 02 Sep. 2013, leg. S. Goranov, S. Deltshev leg. • 5 ex., Sandanski District, above Village Ilindentsi, Cave Sharaliiska Peshtera, 1650 m alt., 21 Aug. 2002, leg. B. Petrov, P. Beron; Predbalkan • 1 ex., Pleven District, Village Dragana, Cave Skoka, under stones, clay-guano, 29 Jan. 1998, leg. B. Petrov, T. Ivanova, T. Troanski .</p><p>Diagnosis.</p><p>(Silvestri 1931; Bareth and Condé 2001; Sendra et al. 2020): Elongated body (Figs 6 a, 7 a). Epicuticle smooth at high magnifications (Fig. 7 c). Antennae with 24–30 antennomeres; distal whorl of 30–36 gouge sensilla that are 45–60 µm long with an apical thin extension (Fig. 5 c – f); 10 complex olfactory chemoreceptors in cupuliform organ each with two concentric floods around a central protrusion (Fig. 7 a); border of cupuliform organ with a whorl of presumably glandular setae long and short, long ones with a shallow distal part with an apical opening (Fig. 5 a, b); protruding frontal process with tubercular setae in most of the population (Fig. 6 b), rarer non-tuberculated population. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp, 1 + 1 mp macrosetae; metanotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp, 1 + 1 mp macrosetae; long, barbed notal macrosetae; a few long, distally barbed notal clothing setae (Figs 6 c, d, 7 b). Two or three ventral tibial macrosetae; unequal claws (ratio 1.30-1.5) with large lateral crests, posterior claw with a large backward overhang (Fig. 8 a – d). Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 0 + 0–1 + 1 la, 1 + 1–2 + 2 post macrosetae; urotergite IV with 1 + 1 la, 5 + 5 post macrosetae; urotergites V with 2 + 2 la, 5 + 5 post macrosetae; urotergites VI – VII with 3 + 3 la, 5 + 5 post macrosetae; urotergites VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae (Fig. 7 d, e). Urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae; stylus setae with a few barbs (Fig. 8 e); typical exertile vesicles (Fig. 8 e, f). Male with a large area of glandular g 1 setae and enlarged appendages with glandular a 1 setae (Fig. 9 a); female with subcylindrical appendages with glandular a 1 setae (Fig. 9 b, c).</p><p>Habitat and distribution.</p><p>A cave-adapted species, which is found in a high number of caves in the studied area. This species appears to be common in the Rhodopes, Pirin, the central parts of the Balkan Mountains, and the Predbalkan. In the Rhodopes, P. bulgarica is distributed throughout the region, and in the present study, it was recorded for the first time in the eastern part, in the region of Dzhebel. However, so far it has not been recorded in the lower, warmer parts of East Rhodopes. In Pirin, the species inhabits caves above 2000 m above sea level, and also in the Western and Central Rhodopes many of the localities are above 1000 m, which clearly shows that the species is cold-resistant. There are only two records of the species in Predbalkan and one in Central Stara Planina Mts. It is worth mentioning that some populations, particularly those of caves Boevskata and Skoka, and the caves in Pirin Mts. have non-protruding setae on frontal process in contrast to all other tuberculated setae populations (Fig. 21).</p></div>	https://treatment.plazi.org/id/809483A18F3C5278B4FECB402B2EA0D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
D2683A2AD77450A0B834A214383BFD2E.text	D2683A2AD77450A0B834A214383BFD2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa) isterina Conde 1993	<div><p>Plusiocampa (Plusiocampa) isterina Condé, 1993</p><p>Figs 10, 21</p><p>Plusiocampa isterina Condé, 1993: 736, figs 1 a, 2, 3 a, 4, 6 a. Type locality: Movile cave, Mangalia, Romania.</p><p>New material examined.</p><p>Bulgaria: Shumensko Plateau: Shumen District • 3 males, Nahodka 13 Cave, 28 Sep. 1996, leg. P. Beron (male with enlarged appendages with a 1, st I with g 1) • 4 ex., Cave Zandana, 16 Jul. 1980, leg. P. Beron; Strandzha Mts . • 1 male, Burgas District, Village Stoilovo, Peshterata s dvata vhoda (cave), 24 Jun. 1980, P. Beron, S. Andreev leg.</p><p>Diagnosis.</p><p>(Condé 1993, 1996; Sendra et al. 2020 b): Elongated body. Epicuticle smooth under optical microscope. Antennae with 23-25 antennomeres; four simple olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with some slightly tubercular setae. Pronotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 3 + 3 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 1 + 1 (2 + 1) la, 2 + 2 lp macrosetae; long, thin, barbed notal macrosetae; short, smooth clothing setae. Two ventral tibial macrosetae; unequal claws (1.3) with large lateral crests, posterior claw with a backward overhang; pretarsal processes short, with a few sparse barbs. Urotergites I – II with 1 + 1 post macrosetae; urotergite III with 2 + 2 post macrosetae; urotergites IV – V with 1 + 1 la, 5 + 5 post macrosetae; urotergites VI – VII with 2 + 2 la, 5 + 5 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae; first urosternite of male with a large field of up to 302 glandular g 1 setae on the posterior portion and supporting subcylindrical large appendages bearing a field of up to 43 glandular a 1 setae (Fig. 10); first urosternite in female with subcylindrical female appendages bearing glandular a 1 setae.</p><p>Habitat and distribution.</p><p>Non-troglomorphic species. Until now, the species was known only from Peştera Movile in Southern Dobruja, Romania. Here, we report it for the first time from Bulgaria, based on specimens found in two caves on the Shumen Plateau and one in the Strandzha Mountains. Additionally, Sendra et al. (2020) reported seven specimens of Plusiocampa (Plusiocampa) aff. isterina found in Ledenjača Cave, Bosnia and Herzegovina. These specimens were distinguished from the typical form by a higher number of antennomeres (25–28), the presence of six olfactory chemoreceptors in a relatively large cupuliform organ (occupying one-third of a short antennomere), and a more prominent frontal process bearing distinctive tuberculate setae.</p><p>It is interesting that two species of Diplura are recorded in the Zandana cave – P. arbanisiensis and P. isterina . Future research will show whether these are indeed different species or whether the material is misidentified (Fig. 21).</p></div>	https://treatment.plazi.org/id/D2683A2AD77450A0B834A214383BFD2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
AB36C86476DB502C9E275AB27F72161B.text	AB36C86476DB502C9E275AB27F72161B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Plusiocampa) lindbergi Conde 1956	<div><p>Plusiocampa (Plusiocampa) lindbergi Condé, 1956</p><p>Fig. 21</p><p>Plusiocampa (Plusiocampa) lindbergi Conde, 1956: 7, figs 1, 2. Type locality: Saint-Jean-le Prodrome cave, Serres District, Greece.</p><p>Material examined.</p><p>Greece: Menikio Mts. • 1 ex., Timiou Prodromou Monastery, 11 km NE from Serres, cave Piladele (Saint-Jean-le Prodrome), 490 m alt., in guano, 19 Mar. 2010, leg. B. Petrov; Thasos Island • 1 ex., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.62212&amp;materialsCitation.latitude=40.78384" title="Search Plazi for locations around (long 24.62212/lat 40.78384)">Skalo Rachoniou</a>, unnamed cave 40.78384 ° N, 24.62212 ° E, 07 Sep. 2010, leg. B. Petrov.</p><p>Diagnosis.</p><p>(Condé 1956; Sendra et al. 2020 b): Slightly troglomorphic species. Antennae with 34–37 antennomeres; five olfactory chemoreceptors in cupuliform organ of a possibly regenerated antenna; non-protruding frontal process with non tuberculate setae. Pronotum with 1 + 1 ma, 4 + 4 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; metanotum with 1 + 1 ma, 2 + 2 lp macrosetae; long, large, barbed notal macrosetae; smooth clothing setae. Two ventral tibial macrosetae; subequal claws (1.05–1.1) with middle-sized lateral crests, posterior claw with no backward overhang. Urotergites I – III with 1 + 1 post macrosetae; urotergite IV with 1 + 1 la, 2 + 2 post macrosetae; urotergites V – VII with 1 + 1 la, 4 + 4 post macrosetae; urotergite VIII with 6 + 6 post macrosetae; abdominal segment IX with 8 + 8 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae. Male with narrow area of glandular g 1 setae, enlarged subcylindrical appendages with up to 60 glandular a 1 setae; female with slender, subcylindrical appendages.</p><p>Habitat and distribution.</p><p>Subterranean species, known from Saint-Jean-le-Prodrome Cave (Spilaio Pelade) and from Mesovoid Shallow Substratum (MSS) near Xánthi, in the Macedonian region of Greece. Here, we report the species also from a cave on Thasos Island, which is not surprising given its close proximity to the mainland (Fig. 21).</p></div>	https://treatment.plazi.org/id/AB36C86476DB502C9E275AB27F72161B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
1A35A5D68E855B3B856EB87CAA96440B.text	1A35A5D68E855B3B856EB87CAA96440B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Stygiocampa) bureschi Silvestri 1931	<div><p>Plusiocampa (Stygiocampa) bureschi Silvestri, 1931</p><p>Figs 13, 14, 15, 16, 17, 21</p><p>Plusiocampa bureschi Silvestri, 1931: 100, figs iii – v. Type locality: Temnata dupka, Lakatnik Railway Station, Bulgaria.</p><p>Plusiocampa (Stygiocampa) rauseri Rusek, 1965: 94, figs 18–24.</p><p>Plusiocampa (Stygiocampa) bureschi: Beron, 2015: 92.</p><p>New material examined.</p><p>Bulgaria: Vrachanska Planina Mts • 2 ex., Svoge Ditstrict, Lakatnik Railway Station, cave Razhishkata peshtera (type locality of P. rauseri), 28 Feb. 1993, leg. B. Petrov • 15 ex., same locality, 22 Feb. 1995, leg. B. Petrov • 3 ex., Svoge Ditstrict, Lakatnik Railway Station, cave Kozarskata, 21 Mar. 1993, leg. P. Stoev • 3 ex., same locality, 03 Nov. 2010, leg. B. Petrov, N. Toshkova • 1 ex., Svoge Ditstrict, Lakatnik Railway Station, cave Svinskata dupka, 450 m, alt., 19 Apr. 2006, leg. B. Petrov • 1 ex., same locality, 03 Nov. 2010, leg. B. Petrov, N. Toshkova • 3 ex., Mezdra District, Cherepish Railway Station, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.59615&amp;materialsCitation.latitude=43.09811" title="Search Plazi for locations around (long 23.59615/lat 43.09811)">cave Novata</a> pestera, 43.09811 ° N, 23.59615 ° E, 562 m alt., 01 Jun. 2012, leg. N. Toshkova • 3 ex., Mezdra District, Cherepish Railway Station, cave Serapinovata peshtera, 15 Apr. 1993, leg. P. Stoev • 1 ex., Mezdra District, Cherepish Railway Station, cave Ezeroto, 03 Feb. 2008, leg. Y. Makulev, Zh. Zhechev • 4 ex., Sofia District, village Iskrets, cave Tsarkvishte, 14 Nov. 1993, leg. B. Petrov • 1 ex., Sofia District, village Breze, cave Yamata, 1000 m alt., 20 Jun. 1993, leg. B. Petrov • 1 female, Sofia District, village Tserovo, cave Vodnata peshtera, 09 May. 1982, leg. P. Beron • 2 ex., Vratsa District, village Chelopek, cave Dedova dupka, alt. 885 m alt., 23 Sep. 2017, leg. K. Lakovski, O. Genova • 1 male, 1 female, same locality, and collectors, 03 Nov. 2017 .</p><p>Diagnosis.</p><p>(Silvestri 1931; Condé 1956, 1959; Condé and Bareth 1996; Bareth and Condé 2001): Epicuticle with a surface in strips in anastomosis unions visible at high magnifications, not visible in optical microscope (Fig. 14 a). Antennae with 39–54 antennomeres elongated with up to 32 thin gouge sensilla on medial antennomers, covered by up to 5 whorls of macrosetae (Fig. 13 c, d); base of the trichobothria elevated overpassing the surface (Fig. 13 b); seven to nine complex olfactory chemoreceptors in cupuliform organ, each with a prominent central protrusion surrounding a single fold (Fig. 13 a). Labium subtriangular with the typical latero-anterior bacilliform sensillum in addition to a field of about 120 thin gustative glandular sensilla (Fig. 14 b). Pronotum with 1 + 1 ma, 2 + 2 la, 2 + 2 lp macrosetae; mesonotum with 1 + 1 ma, 1 + 1 lp macrosetae; metanotum without macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. No ventral tibial macrosetae; slightly unequal claws (1.1–1.3: ratio posterior / anterior) with large and narrow lateral crests, posterior claw with a small backward overhang (Fig. 15 a – e); calcars all covert with numerous long barbs (Fig. 15 f). Urotergite VII with 2 + 2 post macrosetae; urotergites VIII with 4 + 4 post macrosetae; abdominal segment IX with 6 + 6–7 + 7 post macrosetae; urosternite I with 7 + 7, urosternites II – VII with 5 + 5, urosternite VIII with 2 + 2 macrosetae (Fig. 16 c – f); stylus setae all covered with numerous long barbs (Fig. 17 a, b); typical exertile vesicles (Fig. 17 c, d). Male without glandular g 1 setae, long subcylindrical slightly enlarged appendages with glandular a 1 setae; female with very long subcylindrical appendages with glandular a 1 setae (Fig. 16 a, b). Spermatozoid fascicles 50 µm in diameter, 25 µm wide; wheels with 3 ½ turns of 300–350 µm and 5.6 µm diameter filament.</p><p>Habitat and distribution.</p><p>The species seems highly adapted to the cave environment, and until now, it has been known from 11 caves in the area surrounding Tserovo and Lakatnik railway stations (Beron 2015). With the present study, its range has expanded by approximately 14 km (in a straight line) to the west-southwest, where it has been recorded in caves near the villages of Breze and Iskrets, and by about 18 km to the north-northeast, where it has been found in three caves in the area of Cherepish railway station and the village of Chelopек. All recorded locations are within the karst region of the Vratsa Mountain, one of the most cave- and cave-organism-rich areas in the country (Fig. 21).</p></div>	https://treatment.plazi.org/id/1A35A5D68E855B3B856EB87CAA96440B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
CD2219913EEC5EB38D707C0C17C461E7.text	CD2219913EEC5EB38D707C0C17C461E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plusiocampa (Stygiocampa) petrovi Sendra & Stoev & Selfa 2025	<div><p>Plusiocampa (Stygiocampa) petrovi Sendra &amp; Stoev sp. nov.</p><p>Figs 18, 19, 20, 21</p><p>Etymology.</p><p>We name this species after the late colleague and friend Boyan Petrov, a zoologist at the National Museum of Natural History in Sofia, who collected most of the material studied in this publication. The specific epithet is a noun in the genitive case.</p><p>Material examined.</p><p>Holotype • female deposited in the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.58883&amp;materialsCitation.latitude=43.16261" title="Search Plazi for locations around (long 23.58883/lat 43.16261)">Natural History Museum</a> of Sofia, Non-insect Invertebrates – Id. 10839 NMNHS. Bulgaria: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.58883&amp;materialsCitation.latitude=43.16261" title="Search Plazi for locations around (long 23.58883/lat 43.16261)">Vrachanska</a> Planina Mts.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.58883&amp;materialsCitation.latitude=43.16261" title="Search Plazi for locations around (long 23.58883/lat 43.16261)">Vrachanski Balkan Natural Park</a>, Vratsa District, Village Pavolche, Cave Pukoya, 750 m alt., 43.16261 ° N, 23.58883 ° E, clay, 03 Jul. 2010, leg. S. Goranov leg.</p><p>Description.</p><p>Body. Body length 5.8 mm female holotype. Epicuticle smooth under optical microscope (Fig. 21 a); slender body with barbed clothing setae along distal two-thirds of each seta.</p><p>Head. Antennae with 41 antennomeres in one completed and intact antenna; antennae, 7 mm long, are approximately ~ 1.2 times longer than the body’s length, with medial antennomeres almost four times longer than wide and apical antennomere three times longer than wide. Cupuliform organ occupying 1 / 9 of total length in the last antennomere has about eight complex olfactory chemoreceptors. Distal and central antennomeres have four to five untidy whorls of smooth macrosetae and scattered smooth setae, in addition to a single distal whorl of 18‒20 long gouge sensilla (35-40 µm long) and up to two very short coniform sensilla 10-12 µm long. Proximal antennomeres with typical trichobothria, plus a small coniform sensillum on the third antennomere in ventral position. Frontal process has non protrusion with clothing setae and a couple of smooth long macrosetae, slightly longer than the three-insertion line macrosetae, with length ratios of a / i / p / x are 21 / 25 / 19 / 19, respectively, in the holotype (Fig. 18 a). Labial palps are suboval with small latero-external sensillum, two guard setae, up to six setae on anterior border, and up to 95 neuroglandular setae in the holotype.</p><p>Thorax. Thoracic macrosetae distribution (Fig. 19 a): pronotum with 1 + 1 ma, 1 + 1 la 3, 1 + 1 lp 1 macrosetae; mesonotum with 1 + 1 ma macrosetae; metanotum without macrosetae. All macrosetae relatively thin, long, and with thin barbs along the basal half to two-thirds of their length; thin marginal setae are also thin, barbed, and double the length of the clothing setae. Legs elongated, and the metathoracic legs overpass 1 / 3 of their length at the end of the abdomen. Tibia longer than the femur or tarsus; coxa / trochanter / femur / tibia / tarsus plus claws: 0.18 / 0.1 / 1.05 / 1.15 / 0.5 mm. Femora I – III with no dorsal macrosetae, and no ventral macrosetae on tibia I-III. Calcars have small thin barbs all over, pubescence type. Two ventral rows of barbed setae on the medial portion of the tarsus; end part of the tarsus with two dorsal smooth long setae plus two ventral long barbed setae. Claws unequal in size, that is, the posterior claw longer than the anterior claw (1.3‒1.4: ratio posterior / anterior). Claws with large lateral crests well developed in posterior claw with a large backward overhang. Pretarsus with similar, long, smooth and setiform lateral processes (Fig. 18 b).</p><p>Abdomen. Distribution of abdominal macrosetae on tergites: 1 + 1 post 5 short macrosetae on VI, 2 + 2 post 4, 5 on VII, 5 + 5 post on VIII; and 8 + 9 post on IX abdominal segment (fig. 22 b). All tergal abdominal macrosetae long and well-differentiated with thin barbs along their length. Urosternite I with 8 + 8 well-barbed macrosetae (Fig. 20); urosternites II to VII with 5 + 5 macrosetae; urosternite VIII with 2 + 2 macrosetae; long and well-barbed urosternal macrosetae. Stylus with an apical, subapical, and ventromedial seta completely covered with thin barbs all around except at the ending part; apical seta shorter than the others. One apparently intact cercus respectively 1.1 times the body length; divided basal article followed by seven primary articles. Length of the cerci increases from proximal to distal articles (basal: five secondary articles 0.36 mm; 0.19 mm; 0.25 mm; 0.28 mm; 0.30 mm; and seven primary articles 0.33 mm; 0.38 mm; 0.40 mm; 0.40 mm; 0.55 mm; 0.90 mm; 1 mm; 0.90 mm). Each article with a variable number of untidy whorls of macrosetae with short, thin barbs on three / fourth of the distal part and some smooth setae. All primary articles with a distal whorl of longer covered with barbs macrosetae and a more distal whorl of short thin barbed setae.</p><p>Secondary sex characters. Female urosternite I with large coniform appendages, bearing each up to 30 glandular a 1 setae in a distal field (Fig. 20). Male unknown.</p><p>Taxonomic position.</p><p>With the addition of Plusiocampa (Stygiocampa) petrovi sp. nov., the number of species in the subgenus Stygiocampa Silvestri, 1934, increases to nine. All are characterized by the absence of notal and urotergal macrosetae, while several species exhibit additional urosternal macrosetae: Plusiocampa (Stygiocampa) nivea (Joseph, 1882), Plusiocampa (Stygiocampa) remyi Condé, 1947, Plusiocampa (Stygiocampa) dalmatica Condé, 1959, Plusiocampa (Stygiocampa) christiani Condé &amp; Bareth, 1996, and Plusiocampa (Stygiocampa) barethae Sendra &amp; Rađa, 2021 (Joseph 1882; Condé 1947, 1959; Condé and Bareth 1996; Sendra et al. 2020 b, 2021 b). Among the Stygiocampa species, P. (S.) bureschi, P. (S.) christiani, and P. (S.) petrovi sp. nov. share a pronounced reduction in the number of both notal and urotergal macrosetae. Nevertheless, these species exhibit clear differences, such as the number of macrosetae on the pronotum (1 + 1 ma, 2 + 2-3 + 3 la, 1 + 1 lp in P. (S.) christiani; 1 + 1 ma, 2 + 2 la, 2 + 2 lp in P. (S.) bureschi and 1 + 1 ma, 1 + 1 la, 1 + 1 lp in P. (S.) petrovi sp. n.) and the number of post macrosetae on the seventh urotergite (0-1 + 1 post in P. (S.) christiani; 2 + 2 post in P. (S.) bureschi and P. (S.) petrovi sp. nov.). On the other hand, P. (S.) petrovi sp. nov. has more unequal claws (with a 1.3–1.4: ratio posterior / anterior) than its geographically closest congener, P. (S.) bureschi (with 1.1: ratio posterior / anterior).</p><p>Habitat and distribution.</p><p>Pukoya Cave is located in the Krushovitsa area near the village of Pavolche, Vratsa region, within the Vrachanski Balkan Nature Park. The cave is 48 meters long and 178 meters deep. It begins with a short, horizontal gallery, followed by two consecutive shafts; the second of these is approximately 15 meters deep. After that, the cave develops vertically. The middle sections are narrow, requiring ropes to reach the bottom, where water is present (Fig. 21).</p><p>In Beron’s (2015) comprehensive monograph on Bulgaria’s cave fauna, no organisms were reported from this cave. Plusiocampa (Stygiocampa) petrovi sp. nov. is the first species recorded in it. Notably, this new species was found in an area where another representative of the subgenus Stygiocampa, P. bureschi, is also present. The latter was recorded here from caves near the village of Chelopек (about 3 km in a straight line from Pavolche) and Cherepish railway station (approximately 6 km in a straight line).</p></div>	https://treatment.plazi.org/id/CD2219913EEC5EB38D707C0C17C461E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sendra, Alberto;Stoev, Pavel;Selfa, Jesús	Sendra, Alberto, Stoev, Pavel, Selfa, Jesús (2025): Cave-adapted Plusiocampinae (Diplura, Campodeidae) from the eastern part of the Balkan Peninsula. Deutsche Entomologische Zeitschrift 72 (2): 317-340, DOI: 10.3897/dez.72.162647
