taxonID	type	description	language	source
1F4087B7FFDDFFC917E2FDE3FC77FB46.taxon	diagnosis	Diagnosis. Species of the genus (Figs 1, 2) are sexually dimorphic with females externally similar to Tufteulisna gen. n., Bitecta and Macotasa Moore, 1878. Males of Plumareola are characterised by the abdomen bearing bristles of long androconial scales on each segment, the rounded forewing apex and tornus, and the modified forewing cell bearing a cluster of androconial scales and having a comma-shaped transverse fold. The male genital capsule of Plumareola (Fig. 92) is characterised by the massive posteriorly directed lateral lobes of the juxta, which is herein considered as an autapomorphic feature of the genus. Additionally, Plumareola differs from other similar genera by the short valva (in proportion to the tegumen-vinculum complex) with a proximally narrow sacculus. The vesica of Plumareola is short and narrow and bears a broad cluster of spines. In the female genitalia of Plumareola (Fig. 145), the corpus bursae is extremely elongate and narrow, which is also unique among the Teulisna sensu lato. Note. The Malayan P. nigrisquama (Bucsek, 2012), comb. n. was originally described in Teulisna and is closely related to the type species of the genus Plumareola described from Java Island. The genus is under revision by Volynkin & Černý (in prep.).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDDFFC817E2FAEBFD6DFEF0.taxon	materials_examined	Type species: Teulisna pendleburyi Holloway, 1982, by present designation.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDDFFC817E2FAEBFD6DFEF0.taxon	diagnosis	Diagnosis. The new genus (Figs 3, 4) is closely related to Plumareola with males sharing such external diagnostic features as the abdomen bearing bristles of androconial hair-like scales and the forewing cell having a comma-shaped transverse fold. However, unlike in Plumareola, males of Tufteulisna gen. n. have shorter abdominal androconial hair-like scales and more angled forewing apex and tornus. The male genital capsule of Tufteulisna gen. n. (Fig. 93) is similar to Plumareola with its robust uncus, heavily sclerotised scaphium, and proximally narrow sacculus, but is distinct in the juxta lacking the processes, and the strongly enlarged bilobate lamella centralis with a massive and heavily sclerotised distal lobe forming a cucullus-like distal section of the valva, the feature considered herein as autapomorphic. The vesica of Tufteulisna gen. n. is longer than in Plumareola and lacks the spines. In the female genitalia, Tufteulisna gen. n. (Fig. 146) differs from Plumareola in the markedly shorter, pyriform and membranous corpus bursae, which has a sclerotised posterior end in the latter genus.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDDFFC817E2FAEBFD6DFEF0.taxon	etymology	Etymology. The genus name is an aggregate of the English ‘ tuft’ and the genus-group name Teulisna, and refers to the male abdomen of members of the new genus bearing tufts of hair-shaped androconial scales. Gender is feminine. Taxonomic content. The genus comprises two valid species: Tufteulisna pendleburyi (Holloway, 1982), comb. n. and Tufteulisna xanthura (Rothschild, 1920), comb. n., and is under revision by Volynkin & Černý (in prep.).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDCFFC817E2FEEEFABDFA4D.taxon	diagnosis	Diagnosis. The male forewing of the genus has a strongly narrowed cell and bears two clusters of androconial scales: an elongate one stretching between the veins R 1 and R 2, and a compact one on the vein Cu proximally (Fig. 5). The male genital capsule of Bitecta (Figs 94, 95) is reminiscent of Teulisna but distinguished by the combination of the following features. (1) The valva is proximally narrow and has nearly parallel margins. (2) The basis valvae is broader. (3) The lamella centralis is strongly elongate and narrow, stretching from the distal end of the editum to the inner side of the distal saccular process, with which it is strongly articulated. (4) The dorsal branch of the distal saccular process is posteriorly dilated and setose while its anterior section is narrow fold-like and stalked with the lamella centralis in the middle of the valva. (5) The vesica lacks the sclerotised elements. The features (3) and (4) are considered herein as characteristic of the genus. The vinculum structure of Bitecta is similar to Teulisna. The type species of the genus lacks the plostri with caselli and the processus intrinsicus but these structures are present in reduced condition in other, otherwise morphologically very similar species in the genus (see Holloway 2001), which suggests that the presence or lack of the intravincular sclerotised structures or coremata in itself cannot be considered as a generic or subgeneric character. The intravincular saccate corema is present, which is similar to most of the subgenera of Teulisna except for Poikilothosia subgen. n. The female genitalia of Bitecta (Fig. 147) differ from Teulisna in the narrow and membranous corpus and appendix bursae whereas in the latter genus the posterior section of the corpus bursae and the proximal section of the appendix bursae are gelatinous and the former also often bears areas of sclerotisation. Compared to Plumareola and Tufteulisna gen. n., the female genitalia of Bitecta have a longer ductus bursae with a sclerotised plate, and a markedly larger appendix bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDCFFCF17E2F8BAFDE6FCED.taxon	materials_examined	Type species: Bitecta fasciata Gaede, 1925, by present designation.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDCFFCF17E2F8BAFDE6FCED.taxon	diagnosis	Diagnosis. Species of the new genus (Fig. 7, 8) are externally reminiscent of Bitecta but differ in the narrower forewing having only one cluster of androconial scales (between the veins R 1 and R 2). In the male genital capsule (Fig. 96), Sundabitecta gen. n. differs from Bitecta in the combination of the following features. (1) The vinculum is shorter, narrower, has thicker arms and lacks the medial saccate corema, and the intravincular sclerotisations as well. (2) The valva is short, strongly dilated proximally and tapered distally. (3) The basis valvae is markedly narrower. (4) The lamella centralis is shorter, bears a crest-like pseudoampulla, and is displaced distally being situated within the inner surface of the distal saccular process and forming a complex with it. (5) The juxta is more heavily sclerotised, plate-like and has dorso-lateral processes, whereas in Bitecta, the juxta is similar to Teulisna, frame-like with a membranous ventro-medial section, and is extended dorsad with a caulis. Unlike in Bitecta, the phallus of Sundabitecta gen. n. has a developed carina protruding distad, and the vesica is broader and bears cornuti. In the female genitalia (Fig. 148), Sundabitecta gen. n. differs clearly from Bitecta in the broad and heavily gelatinous ductus bursae (it is membranous with a sclerotised plate in Bitecta), the globular corpus bursae with a gelatinous posterior section (it is elongate and membranous in Bitecta), and the very short and conical appendix bursae situated dorso-posteriorly and directed posteriad whereas it is utricular, situated postero-laterally and recurved in Bitecta.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDCFFCF17E2F8BAFDE6FCED.taxon	etymology	Etymology. The genus name is an aggregate of the geographical name Sunda and the genus-group name Bitecta and refers to the distribution of the new genus in Sundaland. Gender is feminine. Taxonomic content. The genus currently comprises one valid species: Sundabitecta fasciata (Gaede, 1925), comb. n. (= Teulisna harmani Holloway, 2001, syn. n.), and is under revision by Volynkin & Černý (in prep.).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDBFFCF17E2FC05FE89F93B.taxon	materials_examined	Type species: Teulisna maculata Fang, 2000, by present designation.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDBFFCF17E2FC05FE89F93B.taxon	diagnosis	Diagnosis. The only species of the new genus (Fig. 9, 10) is externally reminiscent of members of the genus Mithuna Moore, 1878 (illustrated by Holloway 2001) and has a broad forewing with a strongly convex costal margin. In male, the subcostal area between the veins R 1 and R 2 bears a cluster of androconial scales, which is similar to Sundabitecta gen. n. The female has forewing markings similar to male but the ground colour is darker, greyish-brown whereas it is ochreous brown in the male. The male genital capsule structure of Fangeulisna gen. n. (Fig. 97) is reminiscent of Bitecta but differs clearly in the long costa occupying the whole dorsal margin of the valva (vs. proximal one-third in Bitecta), and the lamella centralis stretching along and dorsally fused with the costa (it is articulated with the distal saccular process in Bitecta), and forming with it a swollen saccate and setose-spinulose ‘ cucullus’, which is unique among the Teulisna sensu lato. The distal saccular process of Fangeulisna gen. n. is narrow and has a reduced dorsal branch. The juxta of the new genus is frame-shaped with a short caulis, which is similar to Bitecta and Teulisna. The vinculum of Fangeulisna gen. n. is short and lacks the coremata and sclerotisations, which is similar to Sundabitecta gen. n. The female genitalia of Fangeulisna gen. n. (Fig. 149) are reminiscent of Bitecta but differ in the short ductus bursae lacking the sclerotised plate, the posteriorly sclerotised corpus bursae, the anterior section of the corpus bursae bearing a signum, and the reduced and conical appendix bursae, which is well-developed and utricular in Bitecta.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDBFFCF17E2FC05FE89F93B.taxon	etymology	Etymology. The generic name is an aggregate of the surname Fang and the genus-group name Teulisna. The new genus is dedicated to Dr Fang Chenglai, author of the Arctiidae volume of the fundamental book series “ Fauna Sinica ” (Fang 2000). Gender is feminine. Taxonomic content. The new genus is currently monotypic: Fangeulisna maculata (Fang, 2000), comb. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDBFFCE17E2F8B7FE88FAC4.taxon	diagnosis	Diagnosis. Species of the genus (Figs 11 – 91) are diverse externally, usually have distally angular forewing and display a moderate or limited sexual dimorphism. In most groups, males have a forewing with an elongate cluster of androconial scales below the vein Cu (similar to the Thysanoptyx generic complex). The male genitalia of the genus (Figs 98 – 144) are characterised by the combination of the following features. (1) The scaphium is thin, gelatinous or weakly sclerotised. (2) The vinculum is trapezoidal with thin but heavily sclerotised arms. The intravincular area has a medial saccate corema and a complex of sclerotisations: the processus intrinsicus, medius, introrsum, and the paired plostri bearing gelatinous and easily detachable bodies termed the caselli (Birket-Smith 1965; Volynkin 2024). Some of the aforementioned intravincular structures are absent in certain groups. (3) The juxta is frame-like with a membranous ventro-medial section and is extended into caulis dorsally. (4) The basis valvae is well-developed. (5) The costa is very short and occupies only the basal section of the dorsal margin of the valva, in most groups it lays only on the outer wall of the valva. (6) The editum is short, thin and gelatinous or weakly sclerotised. The tendon is present. (7) The sacculus is proximally broad, forming a fold, the dorsal margin of which is connected with the apex of the juxta by the membranous commissure. (8) The distal saccular process has a dorsal plate-like branch lying in the plane of the outer wall of the valva. (9) The lamella centralis is present, distally articulated with the dorsal branch of the distal saccular process, and in a number of groups bears a process. In the recent paper devoted to the genitalia morphology of Lithosiini (Volynkin 2024), it was suggested to use the descriptive term the crest of lamella centralis to refer to the protrusions of this sclerite. However, as in most groups of Teulisna it is not crest-shaped but thorn-shaped or clavate, the new term pseudoampulla is introduced herein to refer to the process of the lamella centralis. The phallus and vesica structures are diverse within the genus. The female genitalia of Teulisna (Figs 150 – 181) are characterised by the combination of the following features. (1) The ductus bursae is relatively short, dorso-ventrally flattened, gelatinous or with sclerotised plates. (2) The corpus bursae is saccate and has a gelatinous posterior section in many species bearing irregular areas of sclerotisation, wrinkles or clusters of spinulose scobination. In certain groups the corpus bursae also has diverticula and / or protrusions. (3) The appendix bursae is short conical, can be situated in various parts of the corpus bursae depending on the group; reduced in certain species. Notes. (1) The genus is subdivided into four subgenera in the present paper. The names Teulisna, Tegulata Walker 1862 and Corcura Walker 1862 were introduced in the same publication (Walker 1862), and Birket-Smith (1965), acting as the First Reviser (see Article 24.2 of ICZN (1999 )), determined the priority of Teulisna over Tegulata and Corcura. (2) The following species are herein excluded from Teulisna and transferred to the genus Mithuna: Mithuna tongdzuythanhi (Dubatolov & Bucsek, 2016), comb. n., Mithuna flava (Dubatolov & Bucsek, 2016), comb. n., Mithuna varia (Bucsek, 2020), comb. n., Mithuna mithunoides (Černý, 2009), comb. n., Mithuna tenebrosus (N. Singh & Kirti, 2016), comb. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFDAFFCE17E2FADDFBA7F93C.taxon	diagnosis	Diagnosis. The forewing ground colour is ochreous or pale brown in most species, with a large blackish medial spot (Figs 11, 12; also illustrated by Holloway (2001 )). The male genitalia of the nominate subgenus (Fig. 98; also illustrated by Holloway (2001 )) are characterised by the combination of the following features. (1) The caulis is broad plate-like. (2) The intravincular sclerotisations and caselli are fully developed. (3) The basis valvae is framed by the jugum basalis. (4) The lamella centralis is displaced proximally and bilobate with the dorsal (proximal) lobe situated at the dorsal corner of the base of the valva replacing the costa, and bears pseudoampulla, which may be erroneously considered as a costal process due to its placement. (5) The phallus is tubular and bears carinal process (es) in many species. The female genitalia of the nominate subgenus (Fig. 150) display no fundamental differences from other subgenera. Note. The subgenus is under revision by Volynkin & Černý (in prep.).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD9FFCD17E2FD96FEC9FBF9.taxon	description	Notes. (1) The type species of the subgenus has reduced intravincular sclerotisations with only the processus intrinsicus being present. However, other three species in the subgenus have fully developed intravincular structures. (2) The subgenus comprises four species endemic to the Indian subcontinent, which were reviewed by Volynkin (2023 a).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD9FFCD17E2FD96FEC9FBF9.taxon	diagnosis	Diagnosis. Species of the subgenus Cossa (Figs 13, 14; also illustrated by Volynkin (2023 a )) differ externally from Teulisna s. str. in the shortly pectinate male antenna, and the more elongate, narrow and distally less angular forewing, which is reminiscent of certain genera in the Thysanoptyx generic complex. The male genitalia of the subgenus Cossa (Fig. 99; also illustrated by Volynkin (2023 a )) are morphologically close to the nominate one but differ in the structure of the lamella centralis, which is narrow string-like whereas it is bilobate in Teulisna s. str. The female genitalia of the subgenus (Fig. 153; also illustrated by Volynkin (2023 a )) display no fundamental differences from other subgenera.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD9FFCC17E2FABCFC53FDDC.taxon	diagnosis	Diagnosis. Species of the genus (Figs 15 – 18) have forewing shape similar to the nominate subspecies with brown ground colour and one or two black spots at the costal margin proximally and / or medially. The male genitalia of Teulisna (Tegulata) (Figs 100, 101) differ from other subgenera in the lack of the distinct border between the basis valvae and the base of the sacculus whereas in other subgenera of Teulisna, the basis valvae is framed by the jugum basalis and is flexibly articulated with the sacculus. Besides this feature, the male genitalia ground plan of Teulisna (Tegulata) is similar to Teulisna s. str. but the former differs in the narrower lamella centralis, which bears a weakly sclerotised and lobular pseudoampulla and almost reaches the dorsal margin of the valva (similar to Cossa; whereas in the of Teulisna s. str. is bilobate and bears a sclerotised thorn-shaped or clavate pseudoampulla), the thin, string-shaped caulis (vs. broad plate-like one in both Teulisna s. str. and Cossa), and the broader sacculus with a larger dorsal branch of its distal process. The female genitalia of the subgenus (Figs 151, 152) display no fundamental differences from other subgenera.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD8FFCC17E2FB78FC2AF8FF.taxon	materials_examined	Type species: Teulisna ganesha Volynkin, 2023, by present designation.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD8FFCC17E2FB78FC2AF8FF.taxon	diagnosis	Diagnosis. Species of the subgenus (Figs 19 – 91) are externally reminiscent of Teulisna (Tegulata) but are clearly different in their male genitalia structures (Figs 102 – 144), which are characterised by the combination of the following features. (1) The vinculum lacks the processus intrinsicus, the medius and the medial saccate corema. The plostri with caselli are present in most species but the T. bipectinis and T. differens sp. n. species-groups. (2) The tendon is elongate and fused with its counterpart into a gelatinous string whereas other subgenera of Teulisna have a very short tendon. (3) The lamella centralis is short plate-like, with a thorn-shaped or triangular pseudoampulla in most species, situated in the middle of the valva. (4) The dorsal branch of the distal saccular process is relatively short. The female genitalia of the subgenus (Figs 154 – 181) display no fundamental differences from other subgenera.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD8FFCC17E2FB78FC2AF8FF.taxon	etymology	Etymology. The subgenus name is an aggregate of the Greek ‘ ποικίλος’ meaning ‘ diverse’ and the genus-group name Lithosia Fabricius, 1798, and refers to the surprising hidden species diversity of the new subgenus. Gender is feminine. Note. The subgenus is reviewed below in the present paper.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD7FFC217E2F8BAFD83FB5C.taxon	description	(Figs 19, 20, 102, 154) Teulisna spinulosa Volynkin, 2023, Ecologica Montenegrina, 68: 128, figs 1 – 3, 9, 12 (Type locality: “ Vietnam, Prov. Lao Cai, 1900 – 2000 m, Fan-si-pan Mts, 14 km NW of Sa Pa, 22 ° 20.9 ' N 103 ° 46.06 ' E ”).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD7FFC217E2F8BAFD83FB5C.taxon	materials_examined	Type material examined. Holotype (Figs 19, 102): male, “ Vietnam, Prov. Lao Cai, 1900 – 2000 m | Fan-si-pan Mts, 14 km NW [of] Sa Pa | 103 ° 46.06 ' E, 22 ° 20.9 ' N, 26 – 29. I. 1999 | leg. L. Peregovits, G. & L. Ronkay | coll. Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 425 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 5 males, same data as in holotype; 145 males, 2 females, [Lao Cai Prov.] 1600 m, Mt. Fan-si-pan (north), Cha-pa, primary forest, 22 ° 17 ' N 103 ° 44 ' E, 25 – 30. III. 1995, V. Sinyaev & A. Schintlmeister leg., gen. slide No.: ZSM Arct. 2021 - 427 ♀ (prepared by Volynkin); 3 males, 1 female, [Tuyen Quang Prov.] Tam Dao (sec. [ondary] forest), 60 km NW Hanoi, 950 m, 21 ° 34 ' N 105 ° 20 ' E, IV. 1995, V. Sinyaev leg., gen. slide Nos.: ZSM Arct. 2021 - 426 ♂, ZSM Arct. 2021 - 428 ♀ (prepared by Volynkin) (all in MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD7FFC217E2F8BAFD83FB5C.taxon	diagnosis	Diagnosis. The forewing length is 13.0 – 13.5 mm in males and 15.0 – 16.0 mm in females. Teulisna spinulosa is vaguely reminiscent of the Himalayan T. ganesha but distinguished by the shorter forewing with a more elongate apex, and the costal spot of the forewing situated more distally. The male genitalia of T. spinulosa are fundamentally different from T. ganesha and are most similar to the partly sympatric and externally quite dissimilar T. bipectinis, unlike which T. spinulosa has a thin and distally tapered uncus (it is swollen and posteriorly dilated in the congener), a shorter vinculum, a longer valva with a flattened, broadly triangular and smooth distal saccular process (it is club-shaped and serrulate in T. bipectinis), and a membranous anellus (it bears a broad dense ring-like cluster of minute triangular thorns in the congener). The phallus of T. spinulosa is shorter than in T. bipectinis, has a shorter and less upcurved coecum, and a longer and thinner carinal process. The vesica of T. spinulosa is broad saccate with two broad clusters of ornamentation distally: a dorsal one consisting of short but robust spinules, and a ventral cluster of short spinulose scobination, whereas the vesica of T. bipectinis is narrow tubular with a denser cluster of spinules basally and subbasally. The female genitalia of T. spinulosa can be easily distinguished from T. bipectinis by the shorter ductus bursae, and the broader corpus bursae having a dilated and densely spinulose gelatinous posterior section, and a short, narrow and gelatinous anterior section whereas in T. bipectinis, the posterior section is tubular and lacking the spinules while the anterior section is broader, teardrop-shaped, and membranous with a narrow cluster of scobination along its left wall.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD7FFC217E2F8BAFD83FB5C.taxon	distribution	Distribution. The species is currently known from Northern Vietnam (Lao Cai and Tuyen Quang Provinces) (Volynkin 2023 b).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD6FFC117E2FB5AFAF0FEF0.taxon	description	(Figs 21, 22, 103, 155)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD6FFC117E2FB5AFAF0FEF0.taxon	materials_examined	Material examined. CHINA: series of both sexes, Hunan, Nanling Mts, 1500 m, Shikengkong Mt., 24 ° 54 ' N 112 ° 57 ' E, 25. X. – 07. XI. 2003, V. Sinyaev leg., gen. slide No.: ZSM Arct. 2021 - 114 ♂ (prepared by Volynkin) (MWM / ZSM); VIETNAM: 1 male, 1 female, [Lao Cai Prov.] Mt. Fan-si-pan (north), Cha-pa, cloud forest, 22 ° 15 ' N 103 ° 45 ' E, 2 – 4. III. 1995, 2400 m, Dr. R. Brechlin leg., gen. slide Nos: ZSM Arct. 2019 - 524 ♂, 2019 - 525 ♀ (prepared by Volynkin) (MWM / ZSM); 26 males, 9 females, [Tuyen Quang Prov.] Tam Dao (sec. forest), 60 km NW Hanoi, 1200 m, 21 ° 34 ' N 105 ° 20 ' E, 1 – 5. V. 1993, Sinyaev & Simonov leg., gen. slide No.: ZSM Arct. 2021 - 115 ♂ (prepared by Volynkin) (MWM / ZSM); 14 males, 3 females, [Thua Thien Hue Prov.] Bach-Ma Nat. Park, 1200 m, 16 ° 10 ' N 107 ° 54 ' E, 26. VII. – 6. VIII. 1996, Sinyaev & Afonin leg., ex coll. Schintlmeister, gen. slide No.: ZSM Arct. 2021 - 116 ♂ (prepared by Volynkin) (MWM / ZSM); THAILAND: 1 male, Chiang Mai Prov., Doi Inthanon NP, 1416 m, 18 ° 30 ' 59 '' N 98 ° 28 ' 13 '' E, 6 – 7. vi. 2005, K. Černý leg., gen. slide No.: AV 8446 ♂ (CKC).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD6FFC117E2FB5AFAF0FEF0.taxon	diagnosis	Diagnosis. The forewing length is 10.0 – 12.0 mm in males and 10.5 – 13.0 mm in females. Teulisna bipectinis is externally vaguely reminiscent of T. spinulosa but differs in its smaller size, rounded forewing apex, greenish-grey colouration of both wings, the smaller costal spot in male and the lack of it in female, and the cluster of shiny bronze androconial scales on the male forewing (it is ochreous in the congener). The detailed comparison of the genitalia of both sexes of T. bipectinis and T. spinulosa is provided above in the diagnosis of the latter species.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD6FFC117E2FB5AFAF0FEF0.taxon	distribution	Distribution. The species is known from China (Yunnan, Hunan, Guangxi, Guangdong, and Hainan) (Fang 2000; Dubatolov et al. 2012), Vietnam (Lao Cai, Tuyen Quang, and Thua Thien Hue Provinces) (Volynkin 2023 b), and Northern Thailand (Chiang Mai Province) (present study).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD5FFC117E2FEEEFC19FD81.taxon	diagnosis	Diagnosis. Members of the species-group are externally similar to the T. differens and T. tricolor species-groups with their forewing having a medial costal protrusion. However, the male genitalia of the T. protuberans species-group are characterised by the broad saccate caselli, which are thin bunch-shaped or absent in other species-groups in the subgenus Poikilothosia subgen. n. Additionally, in the T. protuberans species-group, the phallus is markedly broader than in other congeners, and the distal saccular process is shorter (except for the T. differens species-group).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD5FFC117E2FD19FDBCF89E.taxon	description	(Figs 23 – 25, 104, 105, 156)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD5FFC117E2FD19FDBCF89E.taxon	materials_examined	Type material. Holotype (Figs 23, 104): male, “ Nepal, Annapurna Himal | Geirigan village, 1340 m | 83 ° 45 ' E, 28 ° 20 ' N, 25. VI. 1996 | leg. Gy. M. László & G. Ronkay | coll. Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 340 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. NEPAL: 2 females, same data as in holotype, gen. slide No.: ZSM Arct. 2021 - 451 ♀ (MWM / ZSM); 1 male, 1 female, Annapurna Himal, 1200 m, 1 km N of Syange, 28 ° 24 ' N 84 ° 25 ' E, 07. vi. 1996, Hreblay & Szaboky leg. (MWM / ZSM); 1 male, Royal Chitwan National Park, Bandarjohala Island, Jungle Island Resort, 27 ° 40 ' N 84 ° 28 ' E, 28 – 30. x. 1995, L. Lõkös & L. Peregovits leg., gen. slide No.: ZSM Arct. 2021 - 452 ♂ (MWM / ZSM); INDIA: 3 males, 2 females, NW Himalaya, 1500 m, Uttar [Himachal] Pradesh, Kumaon Distr., Bhimtal, Nainital [Naini Valley], vi. 1974, F. Smetacek leg., Lf [light trap], gen. slide No.: ZSM Arct. 2021 - 341 ♂ (MWM / ZSM); 1 male, Sikkim, 2800 [m], May [18] 95, Pilcher [leg.] / Pilcher coll., Pres. 1917 [by] Mrs Pilcher, gen. slide No.: AV 7565 ♂ (OUMNH); 1 female, Sikkim, 5000 ft., Kurseong, v. 1922, Mrs Fletcher coll. / Brit. Mus. 1926 – 142 (NHMUK); MYANMAR: 4 males, 3 females, [Kachin State] 21 km E [of] Putao, Nan Sa Bon Village, 550 m, 1 – 5. v. 1998, Murzin & Sinyaev leg., gen. slide Nos: ZSM Arct. 2021 - 343 ♂, 2021 - 344 ♂, 2021 - 453 ♀ (MWM / ZSM); 1 male, [Kachin State] 25 km E [of] Putao, env. Nan Sa Bon Vill., 800 m, 6 – 9. v. 1998, Murzin & Sinyaev leg. (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD5FFC117E2FD19FDBCF89E.taxon	diagnosis	Diagnosis. The forewing length is 12.0 – 13.0 mm in males and 14.0 mm in females. Teulisna distributa sp. n. is externally indistinguishable from the partially sympatric T. pilcheri sp. n. and the reliable identification requires the examination of the genitalia structures. The male genital capsule of T. distributa sp. n. differs clearly from other similar congeners in the proximally narrower valva, the larger, thorn-shaped pseudoampulla, and the narrower and cylindrical distal saccular process, which is flattened in similar species. The phallus of T. distributa sp. n. is markedly narrower than in other members in the T. protuberans species-group, and the vesica is considerably shorter and globular whereas it has a tubular or saccate main chamber in similar species. In the female genitalia, T. distributa sp. n. is distinguished from other species in the species-group by the narrower ostium bursae, the shorter and weakly sclerotised ductus bursae, and the membranous corpus bursae densely covered with minute spinules posteriorly, whereas in the similar congeners, the corpus bursae has gelatinous areas and sclerotised plates.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD5FFC117E2FD19FDBCF89E.taxon	distribution	Distribution. The new species is known from Nepal, North and Northeast India and North Myanmar (Kachin State).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD5FFC117E2FD19FDBCF89E.taxon	etymology	Etymology. The specific epithet refers to the wide distribution of the new species. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC017E2FF41FD08FB21.taxon	description	(Figs 26 – 29, 106 – 109, 157)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC017E2FF41FD08FB21.taxon	materials_examined	Type material. Holotype (Figs 26, 106): male, “ N. Thailand | Chiang Rai, 1090 m, road 1150, | km 17 from Wiang Pa Pao | 19 ° 18 ' 45 '' N, 99 ° 23 ' 24 '' E | 27. V. 2011 leg. K. Černý ” / “ Slide | AV 7871 ♂ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. THAILAND: 1 male, Chiang Rai Prov., 1090 m, road 1200, 19 ° 18 ' 39 '' N 99 ° 21 ' 47 '' E, 28. v. 2011, K. Černý leg., gen. slide No.: AV 7872 ♂ (CKC); 1 male, Chiang Mai Prov., Chae Son NP, 1496 m, 18 ° 51 ' 22 '' N 91 ° 22 ' 03 '' E, 9. vi. 2005, K. Černý leg., gen. slide No.: AV 7867 ♂ (CKC); 1 male, Mae Hong Son Prov., 12 km E [of] Khun Yuam, 1200 m, 18 ° 48 ' 10 '' N 98 ° 06 ' 04 '' E, Wald mit Ufervegetation [forest with riparian vegetation], 2. v. 2006, K. Černý leg., gen. slide No.: AV 7880 ♂ (CKC); 2 males, 1 female, 17 – 23. v. 1991, Mae Hong Son [Prov.], Ban Huai Po, 1600 – 2000 m, J. Horak leg., gen. slide Nos: AV 8459 ♂, AV 8460 ♀, AV 8463 ♂ (CKC); 1 male, Lampang [Prov.], Mae Chaem, 1453 m, 18 ° 56.119 ' N 99 ° 23.481 ' E, 9. v. 2008, K. Černý leg., gen. slide No.: AV 7868 ♂ (CKC); 1 male, Loei [Prov.], Phu Rua Nat. Park, 900 m, 20, 22. viii. 1987, Mamoru Owada [leg.], ex coll. Yasunori Kishida, gen. slide No.: YKSY 31 (prepared by S. - Y. Huang) (CHSY). Note. The male genitalia of the species display remarkable variability in the length and width of the distal saccular process, and length and the degree of the curvature of the vesica even within the same population (e. g., see figs 108, 109).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC017E2FF41FD08FB21.taxon	diagnosis	Diagnosis. The forewing length is 11.0 – 12.0 mm in males and 13.0 mm in the female. Teulisna tubularis sp. n. is externally very similar to T. obliquistria, T. subtilis sp. n. and T. variabilis sp. n. also occurring in Northern Thailand and the reliable identification requires the examination of the genitalia structures. Compared to the sympatric T. obliquistria, T. tubularis sp. n. has a slightly narrower forewing with a less prominent costal protrusion. The male genitalia of the new species differ from the similar congeners in the tubular and upcurved vesica. In the female genitalia of T. tubularis sp. n., the narrow corpus bursae with a heavily gelatinous posterior section bearing irregular sclerotisations, is diagnostic.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC017E2FF41FD08FB21.taxon	distribution	Distribution. The new species is currently known from Northern Thailand (Mae Hong Son, Chiang Rai, Chiang Mai, Lampang, and Loei Provinces). The record of ‘ Teulisna protuberans ’ from Laos by Bucsek (2020) probably belongs to T. tubularis sp. n. but requires confirmation.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC017E2FF41FD08FB21.taxon	etymology	Etymology. The specific epithet is derived from the tubular vesica of the new species. The name is an adjective in apposition to the generic name.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC717E2FAB8FBFFFD28.taxon	description	(Figs 29 – 31, 110, 111, 158)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC717E2FAB8FBFFFD28.taxon	materials_examined	Type material examined. Lectotype (designated herein) (Figs 29, 110): male, “ Momeit. | Burmah. | 2000 ft. | June 1890. | W. Doherty. ” / “ Tegulata | obliquistria. Hmpsn. | type ♂ ” / round label with red margin “ Type ” / “ Collectio | H. J. Elwes. ” / “ Rothschild | Bequest | B. M. 1939 – 1. ” / blue label “ Arctiidae | genitalia slide | No. 4864 ♂ ” / QR-code label with unique ID “ NHMUK 010401722 ” (NHMUK). Additional material examined. THAILAND: 1 male, 23 – 31. v. 1991, Mae Hong Son [Province], Ban Si Lang, 1200 m, J. Horák leg., gen. slide No.: AV 7869 ♂ (CKC); 3 males, 2 females, 17 – 23. v. 1991, Mae Hong Son [Prov.], Ban Huai Po, 1600 – 2000 m, J. Horak leg., gen. slide Nos: AV 8461 ♀, AV 8462 ♀, (CKC); 1 female, Mae Hong Son Prov., Ban Huai Kung, 1060 m, 18 ° 08 ' 09 '' N 98 ° 08 ' 31 '' E, Waldrand [forest edge], 3. v. 2006, K. Černý leg., gen. slide No.: AV 9298 ♀ (CKC). Note. The species was originally described from an unspecified number of specimens of both sexes from two places (“ Sikhim …; Momeit, Burma ”) (Hampson 1894). Subsequently, Hampson (1900) limited his treatment of the taxon to the specimens from “ Momeit, Burma ”. Although only male sex was mentioned by Hampson (1900), the number of the type specimens was not specified by the author therefore this cannot be considered as a valid lectotype designation as the existence of syntypes is possible. In order to stabilise the nomenclature, the male specimen labelled as “ Type ” is herein designated as the lectotype.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC717E2FAB8FBFFFD28.taxon	diagnosis	Diagnosis. The forewing length is 11.0 – 12.0 mm in males and 12.0 mm in females. Teulisna obliquistria is externally very similar to T. tubularis sp. n. and T. subtilis sp. n. also occurring in Northern Thailand and differs from those species only in the slightly more prominent costal protrusion of the forewing. The reliable identification requires the examination of the genitalia structures, which are most similar to T. inthanona sp. n., the comparison with which is provided below in its diagnosis. The male genitalia of T. obliquistria are also similar to T. tubularis sp. n., from which they differ in the shorter and narrower distal saccular process, the somewhat longer and proximally narrower phallus, the broader proximal section of the vesica with a reniform ventral diverticulum (vs. one or two short semiglobular diverticula laterally and dorsally in T. tubularis sp. n.), the presence of the tubular medial diverticulum with a terminal cornutus, and the shorter and straighter distal diverticulum. In the female genitalia, T. obliquistria is distinguished from T. tubularis sp. n. by the anteriorly broader ductus bursae, the shorter but broader corpus bursae with a markedly shorter posterior sclerotised area, the lack of the posterior ventral protrusion of the corpus bursae, and the considerably smaller appendix bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD4FFC717E2FAB8FBFFFD28.taxon	distribution	Distribution. The species is currently known from Northern Myanmar (Shan State) (Hampson 1894) and Northern Thailand (Mae Hong Son Province) (present study).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC717E2FCC6FCA0FA0D.taxon	description	(Figs 32, 112)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC717E2FCC6FCA0FA0D.taxon	materials_examined	Type material. Holotype (Figs 32, 112): male, “ N. Thailand | Chiang Mai-prov. | Doi Inthanon-NP, 1416 m | 18 ° 30 ' 59 '' N, 98 ° 28 ' 13 '' E | 6. – 7. VI. 2005 leg. K. | Cerný [recte: Černý] ” / “ Slide | AV 7878 ♂ | A. Volynkin ” (MWM / ZSM, ex CKC).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC717E2FCC6FCA0FA0D.taxon	diagnosis	Diagnosis. The forewing length is 12.0 mm in the male holotype. Teulisna inthanona sp. n. is morphologically most similar to T. obliquistria, from which the male of the new species differs in the somewhat broader forewing. Compared to T. obliquistria, the male genital capsule of T. inthanona sp. n. has a markedly longer and thicker pseudoampulla and a longer and slightly broader distal saccular process. The phallus of the new species is somewhat broader than in the similar congener. In the vesica, T. inthanona sp. n. differs from T. obliquistria in the markedly broader proximal section with a larger, saccate ventral diverticulum (it is reniform in T. obliquistria), the broader medial diverticulum, and the considerably longer and broader distal diverticulum with a larger terminal cornutus. The female is unknown.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC717E2FCC6FCA0FA0D.taxon	distribution	Distribution. The new species is currently known only from its type locality in Northern Thailand (Chiang Mai Province).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC717E2FCC6FCA0FA0D.taxon	etymology	Etymology. The specific epithet is derived from the Mount Doi Inthanon, where the new species is found. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC617E2F9A5FA9AFC70.taxon	description	(Figs 33 – 35, 113, 159, 160)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC617E2F9A5FA9AFC70.taxon	materials_examined	Type material examined. Lectotype (designated herein) (Figs 33, 159): female, “ Darjeeling. ” / “ Tegulata | protuberans ♀. | (type). Moore ” / “ coll. Atkinson | Moore ” / pink label “ Origin. ” / “ Coll. | Staudinger ” / “ 764. ” / “ 258. ” / “ Ilema | protuberans Moore | sq. Hmpsn. ” (MfN). Additional material examined. INDIA: 1 male, Sikkim, Mt. Kanchenjunga (SE), 27 ° 30 ' N 88 ° 20 ' E, 2600 m, 9 – 10. viii. 1995, E. Afonin & V. Sinyaev leg., gen. slide Nos.: ZSM Arct. 2021 - 342 ♂ (MWM / ZSM); 2 males, 1 female, same locality and collectors as previous but 2000 m, 22 – 31. vii. 1995, gen. slide Nos: ZSM Arct. 2021 - 461 ♂, 2021 - 462 ♀ (MWM / ZSM); 1 male, Sikkim, 5000 ', Aug. [18] 95, Pilcher [leg.] / Pilcher coll., Pres. 1917 [by] Mrs Pilcher, gen. slide No.: AV 7564 ♂ (OUMNH); 1 male, Gopaldhara, Mirik, Sikkim, H. Stevens [leg.] / Rothschild Bequest, B. M. 1939 - 1 (NHMUK); 1 male, [Sikkim] Kurseong, A. Hilsons [leg.], unique ID: NHMUK 010292598 (NHMUK). Note. The species was described from an unspecified number of female specimens deposited in the collection of Otto Staudinger (currently housed in MfN, Berlin) (Moore 1878), which makes the existence of syntypes possible. In order to stabilise the nomenclature, the specimen having Moore’s handwritten type label along with the Staudinger’s pink “ Origin. ” type label is herein designated as the lectotype.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC617E2F9A5FA9AFC70.taxon	diagnosis	Diagnosis. The forewing length is 12.5 – 13.5 mm in males and 13.5 – 15.0 mm in females. The male of T. protuberans differs from other similar congeners in the more contrast forewing pattern with intense rusty and slate grey suffusion in the medial and postmedial areas, respectively. The female of T. protuberans is very similar to another Himalayan species, T. distributa and differs only in the dark grey suffusion on the forewing. The male genitalia of T. protuberans are most reminiscent of T. inthanona sp. n. but differ in the shorter pseudoampulla directed more inwards, the distally narrower distal saccular process, and the markedly broader, longer and medially twisted proximal section of the vesica having a considerably larger ventral (proximal) diverticulum, the shorter and broader, conical medial diverticulum lacking the cornutus, and the shorter and broader distal diverticulum with a smaller, horn-shaped terminal cornutus, which is thorn-shaped with a broader base in T. inthanona sp. n. In the female genitalia, T. protuberans can be easily distinguished from other congeners by the long, narrow, membranous and finely scobinate corpus bursae having two irregular gelatinous and rugose postero-lateral protrusions at the junction with the ductus bursae, and a large heavily gelatinous latero-anterior protrusion.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD3FFC617E2F9A5FA9AFC70.taxon	distribution	Distribution. The species is known from Northeast India (Sikkim and north of West Bengal).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD2FFC617E2FC6EFB5BF89E.taxon	description	(Figs 36, 37, 114, 161)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD2FFC617E2FC6EFB5BF89E.taxon	materials_examined	Type material. Holotype (Figs 36, 114): male, “ 29 - VII- 2017 | Xizang, Pailong [in Chinese] | 2030 m ” / “ leg. Si-yao Huang | Shu-qin Ji, Fu-hong Wei | & Shi-fang Mo ” / [gen. slide No.] “ XZTe 01 ” (SCAU). Paratypes. CONTINENTAL CHINA: 1 female, same data as in holotype, gen. slide No.: XZTe 02 (prepared by Huang) (SCAU); 1 male, data as previous but 30. vii. 2017 (SCAU).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD2FFC617E2FC6EFB5BF89E.taxon	diagnosis	Diagnosis. The forewing length is 10.5 mm in males and 12.0 mm in the female. Teulisna sayinyingpo sp. n. is externally vaguely reminiscent of T. khasiana sp. n. with its short costal protrusion of the forewing, but can be easily distinguished by the smaller size, the less elongate forewing with brown ground colour (vs. reddish-brown in T. khasiana sp. n.), and the intense fuscous suffusion on the hindwing. The male genitalia of T. sayinyingpo sp. n. are most similar to the Thai T. subtilis sp. n. but differ in the markedly narrower dorsal section of the valva, the proximally narrower and apically rounded distal saccular process (it is apically pointed in the congener), the shorter phallus with a longer coecum, the broader, semiglobular medial diverticulum bearing a band-shaped cluster of short spines, and the narrower distal diverticulum bearing a shorter and considerably thinner, spike-shaped terminal cornutus, which is robust and spearhead-shaped in T. subtilis sp. n. As the female of T. subtilis sp. n. is unknown, the female genitalia of T. sayinyingpo sp. n. were compared to T. protuberans instead, from which the new species differs in the 7 th abdominal sternite having subostial lobes, the markedly shorter and narrower corpus bursae, which is medially curved sideways, bears heavier spinulose scobination and lacks the irregular postero-lateral gelatinous rugose protrusions.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD2FFC617E2FC6EFB5BF89E.taxon	distribution	Distribution. The new species is currently known only from its type locality in the south-east of the Tibet Autonomous Region (Xizang) of China.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD2FFC617E2FC6EFB5BF89E.taxon	etymology	Etymology. Sa Yi Nyingpo is the Tibetan name for K ṣitigarbha, one of the four principal bodhisattvas in East Asian Buddhism. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FBCCFD17F802.taxon	description	(Figs 40 – 42, 116, 162)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FBCCFD17F802.taxon	materials_examined	Type material. Holotype (Figs 40, 116): male, [NE India, Meghalaya, Khasi Hills] “ Khasis, | Nat. Coll. ” / “ Doncaster | private coll. | Purch. 1927 ” / “ Slide | AV 7562 ♂ | A. Volynkin ” (OUMNH). Paratypes. INDIA: 1 male, 3 females, same data as in holotype, gen. slide Nos: AV 7563 ♀, AV 7568 ♀ (OUMNH); 1 male, 1 female, [Meghalaya, Khasi Hills] Khasis, April 1894, Nat. Coll., gen, slide No.: AV 7567 ♂ (OUMNH); 1 male, Khasia Hills, Assam / Doncaster private coll., Purch. 1927 (OUMNH); 1 male, 1 female, [Meghalaya, Khasi Hills] Khasia Hills, Assam, Nissary (NHMUK); 1 male, [Meghalaya, Khasi Hills] Cherrapunji, Sept. 1893, Assam / Rothschild Bequest, B. M. 1939 - 1, unique ID: NHMUK 010916182 (NHMUK).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FBCCFD17F802.taxon	diagnosis	Diagnosis. The forewing length is 13.5 – 14.0 mm in males and 15.5 – 16.0 mm in females. Teulisna khasiana sp. n. is externally different from other similar congeners in the strongly elongate forewing with a short costal protrusion and reddish-brown ground colour. In the male genitalia, T. khasiana sp. n. is distinguished from similar species in the lack of the pseudoampulla and the bilobate vesica with a medial-lateral lobe bearing a vesica ejaculatorius, three short subdiverticula of various shapes, and an elongate and narrow tubular subdiverticulum bearing a spearhead-shaped terminal cornutus. The distal lobe is represented by the broad and tubular distal diverticulum bearing a robust spike-shaped terminal cornutus. The female genitalia of T. khasiana sp. n. are most similar to T. obliquistria but differ clearly in the markedly narrower ductus bursae, the anteriorly tapered corpus bursae with a postero-ventral sclerotised protrusion situated on the left side (whereas in T. obliquistria, the corpus bursae is saccate and its postero-ventral sclerotised protrusion is situated on the right side), and the larger appendix bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FBCCFD17F802.taxon	distribution	Distribution. The new species is currently known from Northeast India (Meghalaya).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FBCCFD17F802.taxon	etymology	Etymology. The specific epithet is derived from the Khasi Hills, where the new species is found. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FF41FE1FFBD5.taxon	description	(Figs 38, 39, 115)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FF41FE1FFBD5.taxon	materials_examined	Type material. Holotype (Figs 38, 115): male, “ Lao-NE [NE Laos], Hua Phan [Huaphanh] prov., | 20 ° 12.328 ' N 104 ° 00.621 ' E, | Phu Phan Mt., | 17. v. – 3. vi. 2007, ~ 1750 m, | Vít. Kubáň leg. ” / “ Slide | AV 7881 ♂ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. LAOS: 1 male, same data as in holotype, gen. slide No.: AV 7882 ♂ (CKC); THAILAND: 2 males, Changwat [Province] Chiang Mai, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 13. ix. 1999, A. Szabó & Z. Czere leg., gen. slide No.: ZSM Arct. 2021 - 349 ♂ (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FF41FE1FFBD5.taxon	diagnosis	Diagnosis. The forewing length is 12.0 mm in males. Teulisna subtilis sp. n. is superficially very similar to T. tubularis sp. n. and the identification requires the examination of the genitalia structures. The male genitalia of T. subtilis sp. n. are most similar to the externally rather dissimilar T. sayinyingpo sp. n. and the detailed comparison is provided above in the diagnosis of the latter species. Compared to T. tubularis sp. n., the male genitalia of T. subtilis sp. n. have a markedly shorter pseudoampulla, a narrower phallus, a longer, tubular and upcurved basal section of the vesica (it is distally directed in the similar congener), the short and clavate medial diverticulum lacking the terminal cornutus and with a vesica ejaculatorius originating laterally, and a narrower and shorter distal diverticulum bearing a spearhead-shaped terminal cornutus, which is thorn-shaped in the similar congener. The female is unknown.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FF41FE1FFBD5.taxon	distribution	Distribution. The new species is currently known from Northern Laos (Huaphanh Province) and Northern Thailand (Chiang Mai Province).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD1FFC517E2FF41FE1FFBD5.taxon	etymology	Etymology. The specific epithet is a Latin adjective meaning ‘ subtle’ and refers to the narrow vesica of the new species.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFC417E2FF41FAE0FB21.taxon	description	(Figs 43 – 45, 117, 118, 163)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFC417E2FF41FAE0FB21.taxon	materials_examined	Type material. Holotype (Figs 43, 117): male, “ N. Vietnam, 1400 m | [Hoa Binh Prov.] Mai-chau, 20.50 ' N [20 ° 50 ' N] | 104.50 ' E [104 ° 50 ' E], 40 km SE Moc | chau, Urwald [jungle], 7. – 15. IV | 1995, leg. Sinjaev [recte: V. Sinyaev], ex. | coll. Schintlmeister | Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 475 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 2 males, 1 female, same data as in holotype, gen. slide Nos: ZSM Arct. 2021 - 346 ♂, 2021 - 347 ♂, 2021 - 474 ♀ (MWM / ZSM); 1 female, [Lao Cai Prov.] 1600 m, Mt. Fan-si-pan (North), Cha-pa, Primärurwald [primary jungle], 22 ° 17 ' N 103 ° 44 '' E, 20 – 30. iv. 1995, V. Sinyaev & Einheim. Sammler [local collector] leg., gen. slide No.: ZSM Arct. 2021 - 473 ♀ (MWM / ZSM); THAILAND: 3 males, Chiang Mai Prov., Chae Son NP, 1496 m, 18 ° 51 ' 22 '' N 91 ° 22 ' 03 '' E, 9. vi. 2005, K. Černý leg., gen. slide Nos: AV 7864 ♂, AV 7866 ♂, AV 9294 ♂ (CKC); 2 males, Chiang Rai [Prov.], 1090 m, road 1150, km 17 from Wiang Pa Pao, 19 ° 18 ' 45 '' N 99 ° 23 ' 24 '' E, 27. v. 2011, K. Černý leg., gen. slide Nos: AV 7870 ♂, AV 9295 ♂ (CKC).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFC417E2FF41FAE0FB21.taxon	diagnosis	Diagnosis. The forewing length is 11.0 – 13.5 mm in males and 15.5 – 16.0 mm in females. Teulisna taidona sp. n. is externally reminiscent of T. bucseki sp. n. and T. variabilis sp. n. and differs from the former species in the somewhat smaller size and the more intense fuscous suffusion on both wings. Compared to T. variabilis sp. n., T. taidona sp. n. has a somewhat more prominent costal protrusion of the forewing. The male genitalia of T. taidona sp. n. are most similar to T. bucseki sp. n. but distinguished by the shorter and narrower distal saccular process, the proximally narrower phallus, the larger subbasal diverticulum, the somewhat larger dorsal diverticulum lacking the cornutus, the somewhat longer medial section of the vesica, and the presence of only one distal diverticulum bearing a considerably longer, horn-shaped terminal cornutus whereas T. bucseki sp. n. has two distal diverticula bearing shorter cornuti, the larger of which is claw-shaped while the smaller one is horn-shaped. The female genitalia of T. taidona sp. n. differ from T. bucseki sp. n. in the anteriorly narrower corpus bursae lacking the lateral sclerotised pocket, and the markedly larger appendix bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFC417E2FF41FAE0FB21.taxon	distribution	Distribution. The new species is known from Northern Vietnam (Hoa Binh and Lao Cai Provinces) and Northern Thailand (Chiang Mai and Chiang Rai Provinces).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFC417E2FF41FAE0FB21.taxon	etymology	Etymology. The specific epithet is derived from Tai Dón (White Tai) people, who live in Vietnam, Laos and Yunnan Province of China. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFDB17E2FAB8FE1AFE79.taxon	description	(Figs 46 – 48, 119, 120, 164)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFDB17E2FAB8FE1AFE79.taxon	materials_examined	Type material. Holotype (Figs 46, 119): male, “ N-Vietnam [Lao Cai Prov.] 1600 m | Mt. Fan-si-pan (Nord) | Cha-pa, Primärurwald [primary jungle] | 22.17 '' N 103.44 '' E [22 ° 17 ' N 103 ° 44 ' E] | 20. – 30. IV. 1995 | leg. V. Sinjaev [recte: Sinyaev] & | Einheim. Sammler [local collector] | Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 464 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 1 male, 3 females, same data as in holotype, gen. slide Nos: ZSM Arct. 2021 - 468 ♂, 2021 - 471 ♀, 2021 - 472 ♀ (MWM / ZSM); 1 male, [Lao Cai Prov.] 16 [00] – 1800 m, Mt. Fan-si-pan (West), Cha-pa, Sekund. - Wald [secondary forest], 22 ° 20 ' N 103 ° 40 ' E, xii. 1994, Sinyaev & Einh. Slr. [local collector] leg., gen. slide No.: ZSM Arct. 2021 - 476 ♂ (MWM / ZSM); 1 male, 1 female, [Lao Cai Prov.] Fansipan near Chapa, 16 [00] – 1800 m, 22 ° 20 ' N 103 ° 40 ' E, v. 1995, secondary forest, native collectors [leg.] (MfN).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFDB17E2FAB8FE1AFE79.taxon	diagnosis	Diagnosis. The forewing length is 13.0 – 14.0 mm in males and 15.0 – 16.0 mm in females. Teulisna bucseki sp. n. is morphologically most similar to the sympatric T. fimbriata, from which it differs externally in the more prominent costal protrusion of the forewing. The male genital capsules of the two species are very similar. The phallus of T. bucseki sp. n. is somewhat more distally dilated than in T. fimbriata. The main differences between the species are found in their vesica structures. Compared to T. fimbriata, T. bucseki sp. n. has a large saccate and heavily scobinate subbasal diverticulum (it is very short and finely scobinate in the congener), and a smaller proximal diverticulum bearing a smaller cornutus. The female genitalia of the two species are very similar but in T. bucseki sp. n., the lateral right sclerotised protrusion is broader than in T. fimbriata and has a smaller sclerotised pocket, and the appendix bursae is markedly larger.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFDB17E2FAB8FE1AFE79.taxon	distribution	Distribution. The new species is known from Northern Vietnam (Lao Cai Province).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFD0FFDB17E2FAB8FE1AFE79.taxon	etymology	Etymology. The new species is named after Karol Bucsek, renowned expert in the Asian Arctiinae, who greatly contributed to the knowledge of the Lithosiini of Southeast Asia. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FE70FB26FB5A.taxon	description	(Figs 49 – 51, 121, 122, 165)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FE70FB26FB5A.taxon	materials_examined	Type material. Holotype (by monotypy) (Fig. 49): male, “ Chang Yang | A. E. Pratt Coll. | July 1888 ” / “ Tegulata | fimbriata | sp. n. | Type ” / round label with red margin “ Type ” / “ Leech Coll. | 1900 – 64. ” / QR-code label with unique ID “ NHMUK 010401737 ” (NHMUK). Additional material examined. VIETNAM: 3 males, 1525 m, [Lao Cai Prov.] Mt. Fan-si-pan (North), Cha-pa, Primärurwald [primary jungle] (22 ° 17 ' N 103 ° 44 ' E), 28. x. – 3. xi. 1994, Sinyaev & Einh. Samml. [local collector] leg., gen. slide Nos: ZSM Arct. 2021 - 466 ♂, 201 - 470 ♂ (MWM / ZSM); 1 male, same data as previous but 1600 m, 20 – 30. vi. 1995 (MWM / ZSM); 2 males, 1400 m, [Hoa Binh Prov.] Mai-chau, Urwald [jungle], 40 km SE [of] Moc-chau, 20 ° 50 ' N 104 ° 50 ' E, 07 – 15. iv. 1995, Sinyaev & Einh. Samml. [local collector] leg., gen. slide No.: ZSM Arct. 2021 - 478 ♂ (MWM / ZSM); 1 female, [Lao Cai Prov.] Phansipan [Fansipan], Sapa, 1000 m, v. 1990, Kuban leg., gen. slide No.: AV 9301 ♀ (CKC).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FE70FB26FB5A.taxon	diagnosis	Diagnosis. The forewing length is 13.0 – 14.0 mm in males and 15.0 mm in the female examined. Teulisna fimbriata is similar to T. bucseki sp. n. and the detailed comparison is provided above in the diagnosis of the latter.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FE70FB26FB5A.taxon	distribution	Distribution. The species is known from China (Hubei Province) (Leech 1890), Northern Laos (Bucsek 2020) and Northern Vietnam (Lao Cai and Hoa Binh Provinces) (present study).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FB57FCCAF824.taxon	description	(Figs 52 – 54, 123, 166)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FB57FCCAF824.taxon	materials_examined	Type material. Holotype (Figs 52, 123): male, “ China, Sichuan-prov. | Mt. Omeishan | 11. – 19. Juli [July] 2006 ” / “ Museum Witt ” / “ Slide | CW 00096 ♂ ” [prepared by A. Volynkin] (MWM / ZSM). Paratypes. CONTINENTAL CHINA: 1 male, 1 female, same data as in holotype, gen. slide Nos: CW 00094 ♀, CW 00095 ♂ [prepared by Volynkin] (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FB57FCCAF824.taxon	diagnosis	Diagnosis. The forewing length is 14.0 mm in males and 15.0 mm in the female. Teulisna omeiana sp. n. is externally reminiscent of T. fimbriata but distinguished by the less prominent costal protrusion of the forewing and the paler, more greyish forewing ground colour with a less contrast pattern. In the male genitalia, the new species is similar to T. fimbriata and T. bucseki sp. n. but distinguished by the broader distal saccular process, the narrower phallus, the lack of the subbasal scobinate diverticulum, the proximal diverticulum bearing a conical cornutus (it is short thorn-shaped with a broad plate-like base in the congeners), the presence of the second, utricular proximal diverticulum, the longer and scobinate medial section of the vesica, and the shorter and narrower distal diverticula bearing longer but thinner cornuti. The female genitalia of T. omeiana sp. n. differ from T. fimbriata and T. bucseki sp. n. in the broader corpus bursae with a more heavily sclerotised lateral protrusion bearing a markedly larger appendix bursae but lacking the sclerotised pocket, which is characteristic of T. fimbriata and T. bucseki sp. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FB57FCCAF824.taxon	distribution	Distribution. The new species is currently known only from its type locality in Sichuan Province, Southwest China.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCFFFDB17E2FB57FCCAF824.taxon	etymology	Etymology. The specific epithet is derived from the Mount Omei, where the new species is found. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFDA17E2FF41FE0EF954.taxon	description	(Figs 55 – 60, 124 – 127, 167)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFDA17E2FF41FE0EF954.taxon	materials_examined	Type material. Holotype (Figs 55, 167): female, “ Sikkim. | 2800 | May’ [18] 95. | Pilcher. ” / “ Pilcher coll. | Pres. 1917 | [by] Mrs Pilcher ” / “ Slide | AV 7566 ♀ | A. Volynkin ” (OUMNH). Paratypes. INDIA: 1 male, [West Bengal, Darjeeling] Darjiling, gen. slide No.: AV 7572 ♂ (OUMNH); 3 males, [Meghalaya, Khasi Hills] Khasis, Nat. Coll. / Doncaster private coll., Purch. 1927, gen. slide Nos: AV 7105 ♂, AV 7560 ♂, AV 7569 ♂ (OUMNH); 1 female, [Assam] Dullabcherra / Doncaster private coll., Purch. 1927, gen. slide No.: AV 7570 ♀ (OUMNH); NEPAL: 1 male, Annapurna Himal, 1200 m, 1 km N of Syange, 28 ° 24 ' N 84 ° 25 ' E, 07. vi. 1996, Hreblay & Szaboky leg., gen. slide No.: ZSM Arct. 2021 - 450 ♂ (prepared by A. Volynkin) (MWM / ZSM); 1 male, Annapurna Himal, valley of Kali Gandaki, 1300 m, near Tatopani, 28 ° 29 ' N 83 ° 39 ' E, 20. vi. 1996, Gy. M. László & G. Ronkay leg., gen. slide No.: ZSM Arct. 2021 - 350 ♂ (MWM / ZSM); 1 female, E. Nepal, Sagarmatha, Okhaldhunga, 1800 m, 16. viii. 1990, K. Ito leg., ex coll. Yasunori Kishida, gen. prep. No. YKSY 34 (CHSY); 1 female, same data, but 3. vi. 1991, K. Ito leg., ex coll. Yasunori Kishida (CHSY); 1 male, 1 female, Kathmandu, Godavari, 1600 m, 30. v. 1990, ex coll. Yasunori Kishida, gen. prep. No.: YKSY 32 (♂) (CHSY). Notes. (1) Although the females of T. protuberans and T. pilcheri sp. n. were matched with the males with a reasonable degree of confidence, the species belonging of the males can only be determined with certainty by molecular analysis or a breeding experiment, which were not available in the current study. In order to avoid a possible confusion, we herein designate a female clearly different in its genitalia from the female lectotype of T. protuberans as the holotype of the new species. (2) The populations from Himalaya and Khasi Hills display slight differences in their male genitalia, namely in the scobination of the medial diverticulum and the length and width of the distal section of the vesica. It is difficult to evaluate the taxonomic importance of these differences based on the limited material and without molecular data therefore those populations are considered herein as belonging to the same biological species.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFDA17E2FF41FE0EF954.taxon	diagnosis	Diagnosis. The forewing length is 13.0 – 14.0 mm in males and 15.0 mm in females. Teulisna pilcheri sp. n. differs from another widely distributed Himalayan species T. distributa sp. n. in the forewing with a more prominent costal protrusion and a larger black discal marking. The male genitalia of T. pilcheri sp. n. are vaguely reminiscent of T. fimbriata and T. bucseki sp. n. but differ clearly in the markedly smaller pseudoampulla, the narrower distal saccular process, the larger proximal diverticulum with a smaller cornutus, the recurved medial section of the vesica with a broad but very short medial diverticulum bearing a robust horn-shaped cornutus, and the strongly dilated distal section of the vesica with a single broad conical distal diverticulum bearing a robust spike-shaped terminal cornutus. In the female genitalia, T. pilcheri sp. n. is distinguished from T. fimbriata and T. bucseki sp. n. by the shorter anterior apophyses, the posteriorly broader ductus bursae, and the posterior section of the corpus bursae bearing a considerably larger left lateral protrusion but lacking the right sclerotised one with a sclerotised pocket, which is characteristic of the similar congeners. Additionally, the anterior section of the corpus bursae of T. pilcheri sp. n. is longer and narrower than in T. fimbriata and T. bucseki sp. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFDA17E2FF41FE0EF954.taxon	distribution	Distribution. The new species is known from Nepal and Northeast India (Sikkim, north of West Bengal, Meghalaya and Assam).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFDA17E2FF41FE0EF954.taxon	etymology	Etymology. The new species is named after J. G. Pilcher, collector of the holotype. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFD917E2F94DFB9EFBD6.taxon	description	(Figs 61 – 63, 128 – 130, 168)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFD917E2F94DFB9EFBD6.taxon	materials_examined	Type material. Holotype (Figs 61, 128): male, “ N. Thailand, Lampang | Chae Son NP, 1496 m | 18 ° 51,080 ' N, 99 ° 22,016 ' E | 10. V. 2008 leg. K. Černý ” / “ Slide | AV 7873 ♂ | A. Volynkin ” (MWM / ZSM, ex CKC). Paratypes. THAILAND: 2 females, same data as in holotype, gen. slide Nos: AV 7874 ♀, AV 9296 ♀ (CKC); 1 male, Chiang Mai Prov., Fang Distr., Doi Pha Hom Pok, 2050 m, 20 ° 07 ' 30 '' N 99 ° 08 ' 49 '' E, 5 – 6. v. 2006, K. Černý leg., gen. slide No.: AV 7875 ♂ (CKC); 1 female, same locality, 21 – 25. viii. 2006, T. Ihle leg., gen. slide No.: AV 7876 ♀ (CKC); 1 male, Chiang Mai Prov., Fang Distr., Doi Pha Hom Pok, 2110 m, 20 ° 06 ' 16 '' N 99 ° 07 ' 46 '' E, 23 – 24. v. 2011, K. Černý leg., gen. slide No.: AV 9297 ♂ (CKC); 1 male, Chiang Mai Prov., Fang Distr., Doi Ang Khang, 1425 m, 29 ° 54 ' 11 '' N 99 ° 02 ' 32 '' E, 25 – 26. iv. 2006, K. Černý leg., gen. slide No.: AV 7877 ♂ (CKC). Note. The dorsal diverticulum of the vesica of T. variabilis sp. n. has a dorsally directed subdiverticulum, which remarkably varies in its size and shape in different populations.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFD917E2F94DFB9EFBD6.taxon	diagnosis	Diagnosis. The forewing length is 12.5 – 13.5 mm in males and 15.0 mm in females. Teulisna variabilis sp. n. is externally similar to T. fimbriata, T. obliquistria sp. n. and T. tubularis sp. n., and the reliable identification requires the examination of the genitalia structures. The male genital capsule of T. variabilis sp. n. is reminiscent of T. bucseki sp. n. and T. fimbriata but can be distinguished by the longer pseudoampulla protruding beyond the dorsal margin of the valva. The vesica structure of T. variabilis sp. n. is very characteristic and differs from other congeners in the large and curved dorsal and medial diverticula making the vesica S-shaped, and the medial diverticulum bearing a long and robust, horn-shaped cornutus. In the female genitalia, T. variabilis sp. n. is most similar to T. omeiana sp. n. but differs in the posterior section of the corpus bursae bearing a large and irregular sclerotised plate, which is fused with the sclerotisation of the ductus bursae, the gelatinous and rugose right lateral protrusion of the corpus bursae (it is sclerotised in T. omeiana sp. n.), and the anterior section of the corpus bursae bearing a single elliptical signum whereas T. omeiana sp. n. has two small round signa. Additionally, the anterior apophysis of T. variabilis sp. n. is shorter and thicker than in T. omeiana sp. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFD917E2F94DFB9EFBD6.taxon	distribution	Distribution. The new species is found in Northern Thailand (Lampang and Chiang Mai Provinces).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCEFFD917E2F94DFB9EFBD6.taxon	etymology	Etymology. The specific epithet refers to the variable shape of the dorsal subdiverticulum in the new species’ vesica. The name is an adjective in apposition to the generic name.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCDFFD917E2FBD3FCC1F807.taxon	description	(Figs 64, 65, 131, 169)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCDFFD917E2FBD3FCC1F807.taxon	materials_examined	Type material. Holotype (Figs 64, 131): male, [NE India, Meghalaya, Khasi Hills] “ Khasis | Nov. 1894 | Nat. Coll ” / “ Slide | AV 7571 ♂ | A. Volynkin ” (OUMNH). Paratype. INDIA: female, [Meghalaya, Khasi Hills] Khasis, Nat. Coll. / Doncaster private coll., Purch. 1927, gen. slide No.: AV 7106 ♀ (OUMNH).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCDFFD917E2FBD3FCC1F807.taxon	diagnosis	Diagnosis. The forewing length is 13.5 mm in the male holotype and 14.5 mm in the female paratype. Teulisna robusta sp. n. is externally reminiscent of T. pilcheri sp. n. and differs in the smaller black discal marking in the male and the somewhat more prominent costal protrusion in the female. In the male genitalia, T. robusta sp. n. is most similar to T. variabilis sp. n. but distinguished by the longer and medially narrower uncus, the shorter pseudoampulla, and the narrower distal saccular process. The phalli of the two species are similar. In the vesica of T. robusta sp. n., the dorsal diverticulum is straight conical, membranous and bears a small spike-shaped terminal cornutus whereas in T. variabilis sp. n., it is curved, heavily granulose, lacks the cornutus but has a dorsal subdiverticulum. The distal (ventral) section of the vesica of T. robusta sp. n. is longer than in T. variabilis sp. n., has a markedly smaller conical medial diverticulum (it is massive and C-like curved in T. variabilis sp. n.), and a broader distal diverticulum bearing a straighter but considerably thicker terminal cornutus. The female genitalia of T. robusta sp. n. differ clearly from T. variabilis sp. n. in the posterior section of the corpus bursae lacking the irregular sclerotised plate, the more elongate and narrower anterior section of the corpus bursae bearing a long and broad area of sclerotisation but lacking the signum, and the more weakly gelatinous right lateral protrusion of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCDFFD917E2FBD3FCC1F807.taxon	distribution	Distribution. The new species is currently known only from Khasi Hills in Northeast India (Meghalaya).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCDFFD917E2FBD3FCC1F807.taxon	etymology	Etymology. The specific epithet refers to the robust cornutus in the new species’ vesica. The name is an adjetive in apposition to the generic name.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFD817E2FF41FE0AFA84.taxon	description	(Figs 66, 67, 132, 170)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFD817E2FF41FE0AFA84.taxon	materials_examined	Type material. Holotype (Figs 66, 132): male, “ N-Vietnam [Lao Cai Prov.] 1600 m | Mt. Fan-si-pan (Nord) | Cha-pa, Primärurwald [primary jungle] | (22.17 '' N 103.44 '' E [22 ° 17 ' N 103 ° 44 ' E]) | 20. – 30. IV. 1995 leg. | Sinjaev & einh. Samml [V. Sinyaev & local collector] | Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 469 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 2 males, 3 females, same data as in holotype, gen. slide Nos: ZSM Arct. 2021 - 463 ♀, 2021 - 465 ♂, 2021 - 467 ♂ (MWM / ZSM); 1 female, [Lao Cai Prov.] 16 [00] – 1800 m, Mt. Fan-si-pan (West), Cha-pa, Sek. Wald [secondary forest] (22 ° 20 ' N 103 ° 40 ' E), 20 – 30. x. 1994, Sinyaev & Einh. Saml. [local collector] leg. (MWM / ZSM); 1 male, [Lao Cai Prov.] Fansipan near Chapa, 16 [00] – 1800 m, 22 ° 20 ' N 103 ° 40 ' E, v. 1995, secondary forest, native collectors [leg.] (MfN); 1 male, [Tuyen Quang Prov.] Tam Dao (Sek. Wald [secondary forest]), 60 km NW [of] Hanoi, 950 m, 21 ° 34 ' N 105 ° 20 ' E, iv. 1995, V. Sinyaev leg. (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFD817E2FF41FE0AFA84.taxon	diagnosis	Diagnosis. The forewing length is 13.5 – 14.0 mm in males and 15.0 mm in females. Teulisna brevis sp. n. is morphologically most similar to T. swanni sp. n. and the detailed comparison is provided below in the diagnosis of the latter species. The new species is also externally reminiscent of the sympatric T. fimbriata and the identification requires the examination of the genitalia structures. The male genital capsule of T. brevis sp. n. is distinguished from T. fimbriata by the shorter uncus, the shorter pseudoampulla directed more distally, and the markedly narrower distal saccular process. The phallus of T. brevis sp. n. is considerably shorter and distally broader than in T. fimbriata. The vesica shape of T. brevis sp. n. is diagnostic and differs from T. fimbriata in the narrower proximal section lacking diverticula, the long and conical medial diverticulum directed dorsally, and the short distal diverticulum bearing a massive claw-shaped cornutus. The female genitalia of T. brevis sp. n. differ clearly from T. fimbriata in the shorter anterior apophysis, the long and triangular postvaginal plate (it is elliptical in the congener), the narrower pyriform corpus bursae with a gelatinous protrusion situated latero-ventrally on the right side (vs. two broad lateral protrusions laterally on both sides in T. fimbriata), the lack of the lateral sclerotised pocket, in the right lateral protrusion of the corpus bursae, and the very short appendix bursae situated latero-anteriorly whereas it is well-developed conical and situated latero-ventrally in T. fimbriata.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFD817E2FF41FE0AFA84.taxon	distribution	Distribution. The new species is currently known from Northern Vietnam (Lao Cai and Tuyen Quang Provinces).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFD817E2FF41FE0AFA84.taxon	etymology	Etymology. The specific epithet is a Latin adjectie meaning ‘ short’ and refers to the short phallus of the new species.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFDF17E2FA1DFDACFDDF.taxon	description	(Figs 68, 69, 133, 171)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFDF17E2FA1DFDACFDDF.taxon	materials_examined	Type material. Holotype (Figs 68, 133): male, [N Myanmar, Kachin State, Myitkyina District] “ Upper Burma: | Htawgaw. A. E. Swann. [leg.] | B. M. 1923 – 347. ” / QR-code label with unique ID “ NHMUK 010292599 ” (NHMUK). Additional material. INDIA: 1 female, [Meghalaya, Khasi Hills] Khasis, Nat. Coll. / Doncaster private coll., Purch. 1927, gen. slide No.: AV 7561 ♀ (OUMNH). Note. The single female specimen from Khasi Hills (NE India, Meghalaya) (Fig. 69) has the female genitalia (Fig. 171) similar to T. brevis sp. n., which is closely related to T. swanni sp. n. For this reason, the specimen is treated herein as a possible female of T. swanni sp. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFDF17E2FA1DFDACFDDF.taxon	diagnosis	Diagnosis. The forewing length is 14.0 mm in the male holotype and 15.0 mm in the possible female. Teulisna swanni sp. n. is externally indistinguishable from the allopatric T. brevis sp. n. and is distinguished only by the genitalia structures. The male genital capsules of the two species are very similar but T. swanni sp. n. has a somewhat thicker pseudoampulla than in T. brevis sp. n. The phallus of T. swanni sp. n. is proximally broader and markedly shorter than in T. brevis sp. n. The main differences between the species are found in the vesica structures: compared to T. brevis sp. n., T. swanni sp. n. has a utricular subbasal diverticulum, a longer main chamber of the vesica, a somewhat shorter medial diverticulum, and a larger terminal cornutus in the distal diverticulum. In the female genitalia, the possible female of T. swanni sp. n. differs from T. brevis sp. n. in the broader and considerably shorter postvaginal plate, the narrower ductus bursae, and the somewhat more elongate, posteriorly broader and anteriorly more finely scobinate corpus bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFDF17E2FA1DFDACFDDF.taxon	distribution	Distribution. The new species is known from Northern Myanmar (Kachin State) and possibly from Northeast India (Meghalaya).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCCFFDF17E2FA1DFDACFDDF.taxon	etymology	Etymology. The species is named after Captain Arthur E. Swann, who collected not only the holotype of the new species but numerous other Heterocera in northern Myanmar in 1921 – 1923. The name is a noun in the genitive case.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDF17E2FCA0FE0CF89E.taxon	description	(Figs 70, 71, 134, 172)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDF17E2FCA0FE0CF89E.taxon	materials_examined	Type material. Holotype (Figs 70, 134): male, “ N-Vietnam [Lao Cai Prov.] 1600 m | Mt. Fan-si-pan (Nord) | Cha-pa, Primärurwald [primary jungle] | 22.17 '' N 103.44 '' E [22 ° 17 ' N 103 ° 44 ' E] | 20. – 30. IV. 1995 | leg. V. Sinjaev [recte: Sinyaev] & | Einheim. Sammler [local collector] | Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 345 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. VIETNAM: 5 males, 3 females, same data as in holotype, gen. slide Nos: ZSM Arct. 2021 - 348 ♂, 2021 - 454 ♀ (MWM / ZSM); 1 male, 1 female, Tam Dao (sec. forest), 60 km NW Hanoi, 950 m, 21 ° 34 ' N 105 ° 20 ' E, iv. 1995, V. Sinyaev leg. (MWM / ZSM); LAOS: 1 female, Hua Phan [Houaphanh] Prov., 20 ° 12.328 ' N 104 ° 00.621 ' E, Phu Phan Mt., 17. v. – 3. vi. 2007, ~ 1750 m, Vít. Kubáň leg., gen. slide No.: AV 7883 ♀ (CKC).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDF17E2FCA0FE0CF89E.taxon	diagnosis	Diagnosis. The forewing length is 14.0 – 15.0 mm in males and 16.5 – 17.5 mm in females. Teulisna differens sp. n. has the largest costal protrusion of the forewing among Poikilothosia subgen. n. species and is most reminiscent of T. taidona sp. n., from which it can be distinguished by its larger size and more evenly suffused hindwing. The male genitalia ground plan of T. differens sp. n. is similar to the T. protuberans species-group but differs in the broad swollen uncus, the lack of the caselli and plostri, the longer and broader juxta, and the more distally displaced protrusion of the dorsal fold of the sacculus. The phallus of T. differens sp. n. is narrow tubular and lacking the carinal process, and the vesica is relatively narrow and bears two robust conical cornuti (similar to the T. kishidai species-group). The female genitalia ground plan of T. differens sp. n. is reminiscent of the T. tricolor species-group but differs in the less medially constricted corpus bursae having a lateral sclerotised protrusion, and the very short conical and membranous appendix bursae situated postero-ventrally and directed posteriorly whereas in the T. tricolor species-group, the appendix bursae is clavate with a proximally sclerotised and rugose proximal section while its distal dilated section is membranous, and it is situated laterally and directed latero-anteriorly.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDF17E2FCA0FE0CF89E.taxon	distribution	Distribution. The new species is known from Northern Vietnam (Lao Cai and Tuyen Quang Provinces) and Northern Laos (Houaphanh Province).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDF17E2FCA0FE0CF89E.taxon	etymology	Etymology. The specific epithet is a Latin participle meaning ‘ different’ and refers to the genitalia of the new species, which are markedly different from the externally similar members of the T. protuberans species-group.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDF17E2FDDAFC6CFD06.taxon	diagnosis	Diagnosis. The only species included into the species-group is externally similar to species of the T. protuberans and T. tricolor species-groups but certain characters in its genitalia of both sexes suggest its distinctness (see below in the diagnosis of the new species).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCBFFDE17E2F818FCBBFDFE.taxon	diagnosis	Diagnosis. Species of the species-group are externally similar to the T. protuberans and T. differens species-groups but their male genitalia differ in the phallus bearing a carinal process. Additionally, compared to the T. protuberans species-group, the male genitalia of the T. tricolor species-group have a shorter vinculum with thin bunch-shaped caselli (vs. thick saccate ones in the T. protuberans species-group), a longer distal saccular process, and a smaller phallus (in proportion to the genital capsule). The female genitalia of the species-group are characterised by the corpus bursae clearly subdivided into the posterior and anterior sections by the medial constriction: the posterior section of the corpus bursae is heavily gelatinous with areas of sclerotisation while the anterior section is globular and membranous. The appendix bursae structure is diagnostic: in the T. tricolor species-group it is clavate with a proximally sclerotised and rugose proximal and a membranous dilated distal sections, whereas in other species-groups the appendix bursae is conical and membranous or gelatinous. Additionally, the 7 th abdominal sternite of the species-group has short rounded subostial lobes (also found only in T. sayinyingpo sp n. of the T. protuberans species-group).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDE17E2FDF1FCBAFA07.taxon	description	(Figs 72, 73, 135, 174)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDE17E2FDF1FCBAFA07.taxon	materials_examined	Type material. Holotype (Figs 72, 135): male, “ Taiwan, Prov. Tai-Tung, | Hsiangyang, Police station | 2320 m, 29 – 30. IV. 1997. | leg. Gy. Fábián & S. T. Kovács | coll.: Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 459 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. TAIWAN ISLAND: 1 male, same data as in holotype, gen. slide No.: ZSM Arct. 2021 - 351 ♂ (MWM / ZSM); 1 male, Hualien Co., 10 km NE of Tienchin, 24 ° 04 ' N 121 ° 20 ' E, M. Hreblay & P. Steger leg., gen. slide No.: ZSM Arct. 2021 - 352 ♂ (MWM / ZSM); 1 female, Nantou Co., 3 km SW of Tsuifeng, 24 ° 06 ' N 121 ° 10 ' E, M. Hreblay & P. Steger leg., gen. slide No.: ZSM Arct. 2021 - 460 ♀ (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDE17E2FDF1FCBAFA07.taxon	diagnosis	Diagnosis. The forewing length is 14.0 – 14.5 mm in males and 17.5 mm in the female. Teulisna spatula sp. n. is morphologically similar to T. mingchyra sp. n. and the detailed comparison is provided below in the diagnosis of the latter species. Externally, the new species is also reminiscent of certain members of the T. protuberans species-group such as T. fimbriata but can be distinguished by the hindwing intensely suffused with grey. The male genital capsule of T. spatula sp. n. differs clearly from species of the T. protuberans species-group in the longer and more distally dilated valva with a narrower sacculus bearing a longer, upcurved and spatulose dilated distal process (it is short triangular in the T. protuberans species-group), and the shorter and narrower phallus bearing a small thorn-shaped dorsal carinal process directed dorsally. The female genitalia of the new species are characterised by the posterior section of the corpus bursae having a large elliptical ventral dilation.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDE17E2FDF1FCBAFA07.taxon	distribution	Distribution. The new species is endemic to Taiwan Island (Taitung, Hualien and Nantou Counties).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDE17E2FDF1FCBAFA07.taxon	etymology	Etymology. The specific epithet refers to the spatulate dilated distal saccular process of the new species. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDD17E2F9A2FCA0FD06.taxon	description	(Figs 74, 75, 136, 175)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDD17E2F9A2FCA0FD06.taxon	materials_examined	Type material. Holotype (Figs 74, 136): male, “ Taiwan, Prov. Taoyuan, | Ming Chyr Forest Recreation | Area, 1160 m, 17 – 18. IV. 1997. | leg. Gy. Fábián & S. T. Kovács | coll.: Museum Witt ” / “ Slide | ZSM Arct. | 2021 - 353 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. TAIWAN ISLAND: 1 male, 1 female, same data as in holotype, gen. slide No.: ZSM Arct. 2021 - 456 ♀ (MWM / ZSM); 1 male, Ilan Co., Fu Shan Bot. Garden, LTER site, 700 m, 24 ° 45 ' 13 '' N 121 ° 35 ' 39 '' E, 8 – 9. iv. 1997, L. Peregovits & A. Kun leg., gen. slide No.: ZSM Arct. 2021 - 455 ♂ (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDD17E2F9A2FCA0FD06.taxon	diagnosis	Diagnosis. The forewing length is 13.5 mm in males and 14.5 mm in the female. Teulisna mingchyra sp. n. is externally distinguished from the similar T. spatula sp. n. by the smaller size, the straighter proximal half of the forewing costal margin (it is somewhat concave in T. spatula sp. n.), the shorter medial costal protrusion of the forewing, the more unicolorous and ochreous forewing ground colour, the markedly smaller proximal costal spot of the forewing, and the paler hindwing only slightly suffused with grey at the apex whereas in T. spatula sp. n., it is evenly suffused with grey medially and distally. The male genital capsule of T. mingchyra sp. n. differs from T. spatula sp. n. in the thicker and more strongly downcurved uncus, the distally broader valva, and the distally narrower distal saccular process. The phallus of T. mingchyra sp. n. is slightly broader than in T. spatula sp. n. The vesicae of the two species are similar but in T. mingchyra sp. n., the medial lateral diverticulum is longer, the distal diverticulum is broader, and the terminal cornutus is thicker than the corresponding structures of T. spatula sp. n. Compared to T. spatula sp. n., the female genitalia of T. mingchyra sp. n. have a longer ductus bursae, a narrower posterior sclerotised section of the corpus bursae lacking the ventral elliptical dilation, and a longer appendix bursae with a broader distal section. Additionally, the subostial depression of the 7 th abdominal sternite of T. mingchyra sp. n. is broad V-shaped whereas it is narrower, somewhat deeper and U-shaped in T. spatula sp. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDD17E2F9A2FCA0FD06.taxon	distribution	Distribution. The new species is endemic to Taiwan Island (Taoyuan and Ilan Counties).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFCAFFDD17E2F9A2FCA0FD06.taxon	etymology	Etymology. The specific epithet is derived from the Ming Chyr Forest, where the new species is found. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC9FFDD17E2FCA0FAE9F919.taxon	description	(Figs 76, 77, 137, 176)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC9FFDD17E2FCA0FAE9F919.taxon	materials_examined	Type material. Holotype (Figs 76, 137): male, “ Taiwan, Prov. Taoyuan, | Ming Chyr Forest Recreation | Area, 1160 m, 17 – 18.03.1996. | leg.: Gy. Fábián & L. Németh. ” / “ Slide | ZSM Arct. | 2021 - 354 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. TAIWAN ISLAND: 2 males, 1 female, same loclaity and collectors as in holotype, 29. ii. 1996, gen. slide Nos: ZSM Arct. 2021 - 457 ♂, 2021 - 458 ♀ (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC9FFDD17E2FCA0FAE9F919.taxon	diagnosis	Diagnosis. The forewing length is 17.0 mm in males and 17.5 mm in the female. Teulisna plica sp. n. is externally similar of T. spatula sp. n. but distinguished by the broader forewing with a shorter costal protrusion, and the paler hindwing only slightly suffused with grey subapically. The male genitalia of T. plica sp. n. are reminiscent of the externally dissimilar T. tricolor but the genital capsule of the new species differs from the latter in the longer and thinner uncus, the narrower dorsal section of the valva, the fold-shaped lamella centralis (it is plate-like with a short proximal pseudoampulla in T. tricolor), and the distally spatulate distal saccular process, which is flattened but distally tapered in T. tricolor. The phallus of T. plica sp. n. is distally narrower than in T. tricolor and bears a smaller carinal process. The vesica configurations of the two species are similar but unlike in T. tricolor, the new species has a shorter main chamber, a dorsal medial diverticulum lacking the cornutus, a smaller terminal cornutus, and a shorter distal diverticulum. As the female of T. tricolor is unknown, the female genitalia of T. plica sp. n. were compared to T. mingchyra sp. n. instead, and T. plica sp. n. differs from the latter species in the somewhat shorter ductus bursae, the broader posterior sclerotised section of the corpus bursae having a broad conical lateral protrusion, and the appendix bursae having a broader posterior and a shorter and narrower distal sections. In addition, the subostial depression of the 7 th abdominal sternite of T. plica sp. n. is caliciform whereas it is V-shaped in T. mingchyra sp. n.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC9FFDD17E2FCA0FAE9F919.taxon	distribution	Distribution. The new species is endemic to Taiwan Island (Taoyuan County).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC9FFDD17E2FCA0FAE9F919.taxon	etymology	Etymology. The specific epithet is derived from the Latin ‘ plica’ meaning ‘ a fold’ and refers to the fold-like pseudoampulla of the new species. The name is a noun in the nominative singular.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FA1FFDD1F83D.taxon	description	(Figs 80, 81, 139, 173) Teulisna kishidai Huang, Volynkin & Müller, 2024, Zootaxa, 5506 (4): 578, figs 1, 2, 9, 10, 15 (Type locality: “ Myanmar (West), Chin State, Mt. Victoria (Na Ma Toung) National park, Mindat town guesthouse, 21 ° 22.315 ' N 93 ° 58.601 ' E, 1453 m ”).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FA1FFDD1F83D.taxon	materials_examined	Type material examined. Holotype (Figs 80, 139): male, male, “ Myanmar (West) | Chin State, Mt. Victoria | (Na Ma Toung) National park | Mindat town guesthouse | 21 ° 22.315 ' N, 93 ° 58.601 ' E | 1453 m, 21. V. 2012 | leg. S. Löffler & S. Naumann ” / handwritten label “ [Slide No.] GML 06 ” (ZSM). Paratypes: 2 males, 7 females, Myanmar, Chin State, Mt. Victoria (Na Ma Toung) NP, Chin Hills, Mindat City, Kall Nue, 21 ° 22.314 ' N 93 ° 58.611 ' E, 1457 m, 13. VIII. 2007, P. Spona, Th. Ihle & S. Löffler leg., gen. slide Nos.: GML 07 (male) and GML 08 (female) (CGM); 1 male, Myanmar, Chin State, Kanpetlet, 1700 m, 19. VII. 2014, T. Tanaka leg. (CHSY).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FA1FFDD1F83D.taxon	diagnosis	Diagnosis. The forewing length is 15.0 – 15.5 mm in both sexes. For the morphological characteristics, see above, under the diagnosis of the species-group.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FA1FFDD1F83D.taxon	distribution	Distribution. The species is currently known only from Mount Victoria in Chin State, western Myanmar (Huang et al. 2024).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FF41FDC2FC89.taxon	description	(Figs 78, 79, 138) Ilema tricolor Wileman, 1911, The Entomologist, 44: 109 (Type locality: [Taiwan, Nantou County, Mt. Lantashan] “ Rantaizan (7500 ft.) ”).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FF41FDC2FC89.taxon	materials_examined	Type material examined. Holotype (by monotypy) (Fig. 78): male, “ Rantaizan, | ♂ Formosa. | 7,300 ft. | 13. V. 1909. | A. E. Wileman. ” / “ 542 A. F ” / “ Ilema | tricolor | sp. n. | Type ♂ ” / round label with red margin “ Type ” / QR-code label with unique ID “ NHMUK 010401724 ” (NHMUK). Additional material examined. TAIWAN ISLAND: 1 male, Formosa, [Nantou County, Mt. Lantashan] Rantaizan, 12. v. 1909, A. E. Wileman [leg.], 1913 – 180, unique ID: NHMUK 010916177 (NHMUK).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FF41FDC2FC89.taxon	diagnosis	Diagnosis. The forewing length is 13.0 mm in males. Teulisna tricolor is externally distinct from similar congeners in its unusual contrast wing colouration with the forewing having a rusty brown proximal and dark fuscous distal sections, while the hindwing is pale creamy with a dark fuscous outer fascia. The male genitalia of T. tricolor are similar to T. plica sp. n. and the detailed comparison is provided above in the diagnosis of the latter species. The female is unknown.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FF41FDC2FC89.taxon	distribution	Distribution. The species is endemic to Taiwan Island and is currently known only from its type locality in Nantou County.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC8FFDC17E2FC21FEFBFA82.taxon	diagnosis	Diagnosis. Superficially, the only species of the species-group is reminiscent of the T. pallida species-group but is easily distinguishable by its sexually dimorphic hindwing, which is broad and orange with a medially angled margin in the male and more pointed, pale yellow with a smooth margin in the female. The male genitalia ground plan of the T. kishidai species-group is similar to the T. pallida species-group but differs in the shorter and narrower dorsal section of the valva, longer and broader lamella centralis, and the larger, unilobate blade-shaped distal saccular process, which is distally bilobate in all members of the T. pallida species-group. The vesica of the T. kishidai species-group is narrow with a short main chamber and tubular diverticula whereas in the T. pallida species-group, the main chamber of the vesica is broad and has conical, saccate or semiglobular diverticula. The female genitalia of the T. kishidai species-group are characterised by the strongly elongate and relatively narrow posterior section of the corpus bursae, and the reduced appendix bursae, which is large and gelatinous in the T. pallida species-group.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD317E2FF46FCE9FDDC.taxon	diagnosis	Diagnosis. Species of the T. pallida species-group are characterised by the limited sexual dimorphism and the lack of the costal protrusion of the forewing. In the male genitalia, the wide, laterally flattened and distally bilobate distal saccular process is diagnostic. Additionally, the dorsal section of the valva is large and the vesica is broad with several diverticula of various shapes and a single terminal cornutus, which is thin and strongly elongate. In the female genitalia, the posterior section of the corpus is broad and heavily gelatinous with areas of sclerotisation, and the appendix bursae is large, gelatinous and situated laterally. Note. The species-group was erected and reviewed by Volynkin (2023 c); another new species was subsequently described by Huang et al. (2024).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD317E2FDDAFCD0FA42.taxon	description	(Figs 82, 83, 140, 177)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD317E2FDDAFCD0FA42.taxon	materials_examined	Type material examined. Neotype (designated herein) (Figs 82, 140): male, “ Indien U [ttar] P [radesh], Kumaon | Nainital [Naini Valley] 2000 m | 13. VII. 1984 | leg. W. Thomas ” / “ Slide | ZSM Arct. | 2021 - 367 ♂ | A. Volynkin ” (MWM / ZSM). Additional material examined. INDIA: 15 males, 4 females, same data as neotype, gen. slide No.: ZSM Arct. 2021 - 371 ♀ (prepared by Volynkin) (MWM / ZSM); 2 males, 1 female, Dalhousie, 92 - 98, Harford Coll., 6 / 91 (NHMUK); 1 female, Kangra, 83 - 26 (NHMUK); NEPAL: 1 male, 2835 m, Kalinchok Mt., 6 km NNE Muldi (Murre), 27. vi. 1997, M. Hreblay & K. Csak leg., gen. slide No.: ZSM Arct. 2021 - 368 ♂ (prepared by Volynkin) (MWM / ZSM). Note. The species has been described on the basis of the unspecified number of males held in F. Moore collection (Moore 1878). Unfortunately, no syntypes have been located in the NHMUK where the F. Moore collection is currently housed, and the type material of the species is apparently lost. In order to stabilise the nomenclature, the male specimen matching the original description and originating from the same state of India, Himachal Pradesh is herein designated as neotype.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD317E2FDDAFCD0FA42.taxon	diagnosis	Diagnosis. The forewing length is 13.5 – 14.5 mm in males and 14.0 – 15.0 mm in females. The detailed comparison with the most similar T. ganesha is provided below in the diagnosis of the latter species.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD317E2FDDAFCD0FA42.taxon	distribution	Distribution. The species is currently known from North India (Himachal Pradesh) (Moore 1878; Hampson 1900) and Nepal (Volynkin 2023 c).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD217E2FA5FFE11FD67.taxon	description	(Figs 84, 85, 141, 178) Teulisna ganesha Volynkin, 2023, Ecologica Montenegrina, 63: 17, figs 3, 4, 10, 11, 15 (Type locality: “ Nepal, Ganesh Himal, 3 km NE of Sunpati, 2330 m ”).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD217E2FA5FFE11FD67.taxon	materials_examined	Type material examined. Holotype (Figs 84, 141): male, “ Nepal, Ganesh Himal | 3 km NE of Sunpati, | 2330 m, 13. VI. 1993, | leg. M. Hreblay, G. Csorba ” / “ Slide | ZSM Arct. | 2021 - 369 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. NEPAL: 5 males, the same data as holotype (MWM / ZSM); 1 male, 1 female, Solu Khumbu Himal, 10 km S of Lukla, Bupsa, 2300 m, 3. vii. 1993, M. Hreblay & G. Csorba leg., gen. slide Nos.: ZSM Arct. 2021 - 370 ♂ and 2021 - 374 ♀ (prepared by Volynkin) (MWM / ZSM); 1 female, 2835 m, Kalinchok Mt., 6 km NNE Muldi (Murre), 27. vi. 1997, M. Hreblay & K. Csak leg., gen. slide No.: ZSM Arct. 2021 - 373 ♀ (prepared by Volynkin) (MWM / ZSM). Additional material examined. INDIA: 1 female, Sikkim, 5. v. [18] 89, J. G. Pilcher [leg.] / Rothschild Bequest, B. M. 1939 - 1, unique ID: NHMUK 010292596 (NHMUK).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD217E2FA5FFE11FD67.taxon	diagnosis	Diagnosis. The forewing length is 13.5 – 14.0 mm in males and 14.5 – 15.5 mm in females. Teulisna ganesha is externally reminiscent of T. pallida but distinguished by the greyish-brown body (it is pale ochreous in the congener), the considerably more intense brown suffusion on the forewing, the smaller costal spots, and the indistinct postero-lateral spot on the forewing. The male genital capsule of T. ganesha differs from T. pallida in the considerably shorter pseudoampulla and the markedly longer and distally narrower ventral lobe of the distal saccular process. The vesica of T. ganesha differs can be readily distinguished from four other species in the species-group in the longer and curved proximal ventral diverticulum, and the markedly smaller and semiglobular proximal dorsal diverticulum. Additionally, compared to T. pallida, T. ganesha has a markedly shorter medial dorsal diverticulum, a developed medial ventral diverticulum (absent in T. pallida), and a shorter and considerably thinner cornutus on the distal dorsal diverticulum. In the female genitalia, T. ganesha differs from T. pallida in the broader ductus bursae, the markedly smaller appendix bursae lacking the dorsal sclerotised pocket, and the narrower anterior section of the corpus bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC7FFD217E2FA5FFE11FD67.taxon	distribution	Distribution. The species is currently known from Nepal (Volynkin 2023 c) and Northeast India (Sikkim) (present study).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD217E2FC82FE67FB00.taxon	description	(Figs 86, 87, 142, 179)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD217E2FC82FE67FB00.taxon	materials_examined	Type material examined. Holotype (Figs 86, 142): male, “ Indien [India] | Sikkim | Pemayangtse | 2000 m | 23. – 28. VII. 1990 | leg. Dr. W. Thomas ” / “ Slide | ZSM Arct. | 2021 - 375 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. INDIA: 1 female, the same data as holotype (MWM / ZSM); 120 specimens of both sexes, Sikkim, SE Kanchenjunga Mt., 27 ° 30 ' N 88 ° 20 ' E, 2000 m, 22 – 31. vii. 1995, Afonin & Sinyaev leg., ex coll. Schintlmeister, gen. slide No.: ZSM Arct. 2021 - 372 ♀ (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD217E2FC82FE67FB00.taxon	diagnosis	Diagnosis. The forewing length is 12.5 – 13.5 mm in males and 15.0 – 16.0 mm in females. Teulisna thomasi is externally reminiscent of T. hreblayi and the detailed comparison is provided below in the diagnosis of the latter species.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD217E2FC82FE67FB00.taxon	distribution	Distribution. The species is currently known from two localities in Sikkim (Northeast India) (Volynkin 2023 c).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD117E2FA9EFED1FD67.taxon	description	(Figs 90, 91, 143, 180)	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD117E2FA9EFED1FD67.taxon	materials_examined	Type material examined. Lectotype of Cossa quadrisignata (designated by Volynkin 2023 c) (Figs 90, 143): male, “ Darj. [eeling] ” / “ Cossa | quadrisignata. | (type) Moore ” / “ coll. Atkinson | Moore ” / “ Ilema | quadrisignata | Moore | s. g. Hmpsn. ” / “ * 308. ” / “ 762 ” / “ Coll. | Staudinger ” / pink label “ Origin. ”, DNA voucher No. MFNLEP 1045, gen. slide No.: MfN- 367 (prepared by Volynkin) (MfN). Lectotype of Cossa brunnea (designated by Volynkin 2023 c) (Fig. 91): female with abdomen of Brunia antica glued on, “ Darj. [eeling] ” / “ Cossa | brunnea. ♀. | (type) Moore ” / “ coll. Atkinson | Moore ” / “ Ilema | brunnea | Moore | s. g. Hmpsn. ” / “ 276. ” / “ 759 ” / “ Coll. | Staudinger ” / pink label “ Origin. ”, DNA voucher No. MFNLEP 1042, gen. slide No.: MfN- 364 (prepared by Volynkin) (MfN). Additional material examined. INDIA: 1 female, Sikkim, 7000 ft., Aug. 1896, Pilcher / 99 – 206., unique ID: NHMUK 010292595 (NHMUK).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD117E2FA9EFED1FD67.taxon	diagnosis	Diagnosis. The forewing length is 13.0 mm in the male lectotype and 13.5 mm in females. Teulisna quadrisignata is most morphologically similar to T. thomasi, from which it differs externally in the somewhat smaller size, the slightly narrower forewing with a markedly darker, brown ground colour (it is deep ochreous in the congener), the more intense dark suffusion in the medial area of the forewing, the darker subterminal area with a diffuse submarginal line, and the larger proximal costal spot in the male. Additionally, the female of T. quadrisignata has a remarkably darker hindwing than in T. thomasi. In the male genital capsule, T. quadrisignata differs from T. thomasi in the nearly straight lamella centralis (it is downcurved in T. thomasi), the narrower distal saccular process with a smaller ventral lobe, and the somewhat shorter vinculum. The phallus of T. quadrisignata is proximally broader than in T. thomasi. The vesica of T. quadrisignata is markedly narrower than in the similar congener and has a proximal dorsal diverticulum (it is absent in T. thomasi); the proximal ventral diverticulum is shorter, and the terminal cornutus is considerably longer and nearly straight whereas it is medially curved in T. thomasi. Additionally, compared to T. thomasi, the medial ventral diverticulum of T. quadrisignata is longer and broader. The female genitalia of T. quadrisignata differ from T. thomasi in the shorter and narrower ductus bursae, the more medially constricted corpus bursae with a more rugose posterior gelatinous section having a ventral pocket, and a smaller anterior globular membranous section, and the conical and more rugose appendix bursae, which is saccate in T. thomasi.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC6FFD117E2FA9EFED1FD67.taxon	distribution	Distribution. The species is currently known from Northeast India (northern West Bengal and Sikkim).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC5FFD717E2FC82FD23FBD9.taxon	description	(Figs 88, 89, 144, 181) Teulisna hreblayi Volynkin & Huang in Huang, Volynkin & Müller, 2024, Zootaxa, 5506 (4): 580, figs 3, 4, 11, 12, 16 (Type locality: “ East Nepal, Deorali Danda, 1 km N of Yamphudin, 2000 m ”).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC5FFD717E2FC82FD23FBD9.taxon	materials_examined	Type material examined. Holotype (Figs 88, 144): male, “ East-Nepal | Deorali Danda | 1 km N of Yamphudin | 2000 m, 19. VI. 1998 | leg. Márton Hreblay | & Balázs Benedek | Museum Witt ” / “ Slide | CW 00091 ” [prepared by A. Volynkin] (MWM / ZSM). Paratypes: 5 males, 7 females, same data as holotype, gen. slide No.: CW 00093 ♀ (prepared by Volynkin) (MWM / ZSM); 4 males, East Nepal, Kanchenjunga Himal, Khambachen, 4150 m, 28. VI. 1998, Márton Hreblay & Balázs Benedek leg., gen. slide. No.: CW 00092 ♂ (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC5FFD717E2FC82FD23FBD9.taxon	diagnosis	Diagnosis. Teulisna hreblayi is externally reminiscent of T. thomasi and T. quadrisignata but can be readily distinguished by the submarginal area of the forewing much less suffused with fuscous in both sexes. The male genital capsule of T. hreblayi differs from T. thomasi and T. quadrisignata in the broader dorsal section of the valva with a somewhat more convex dorsal margin, the lamella centralis bearing a robust spike-like process directed dorsally (it is thinner and directed more anteriorly in T. quadrisignata and vestigial in T. thomasi), and the markedly shorter ventral lobe of the distal saccular process. The phallus of T. hreblayi is considerably longer and broader (in proportion to the genital capsule) than in similar congeners. In the vesica structure, the new species is most similar to T. quadrisignata but distinguished by the longer and broader main chamber, the larger, sack-like and more heavily granulose 1 st proximal diverticulum directed anteriorly (it is rather utricular and directed ventrally in the congener), the 2 nd proximal diverticulum lacking graniculi, the longer, broader and more heavily granulose 1 st medial diverticulum, the lack of the 2 nd medial diverticulum, the longer and broader distal diverticulum bearing a markedly longer cornutus, and the larger apical diverticulum. The female genitalia of T. hreblayi are most similar to T. thomasi, from which the former species differs clearly in the shorter ductus bursae with convex lateral margins, the shorter and rugose posterior section of the corpus bursae with a gelatinous ventral diverticulum posteriorly (it is absent in the congener), the somewhat broader and globular anterior section of the corpus bursae displaced laterally on the left side (it is sack-like and situated anteriorly in T. thomasi), the longer signum, and the configuration of the appendix bursae, which in the new species is heavily sclerotised with a membranous apex, strongly elongate (longer than the corpus bursae), and has a dilated, semiglobular proximal section and a narrow, cylindrical and straight distal section whereas the appendix bursae of T. thomasi is sack-like and markedly shorter than the corpus bursae.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC5FFD717E2FC82FD23FBD9.taxon	distribution	Distribution. The new species is currently known from eastern Nepal (Huang et al. 2024). Acknowledgements The senior author expresses his sincere thanks to the following colleagues for their kind assistance during visits to the collections under their care: Axel Hausmann, Mei-Yu Chen and Ulf Buchsbaum (ZSM); James Hogan (OUMNH); Alberto Zilli and Geoff Martin (NHMUK); and Wolfram Mey and Theo Leger (MfN). The third author would like to express his sincere thanks to Yasunori Kishida (Tokyo, Japan) for providing specimens for the study. Our special thanks go to Rob de Vos (RMNH, Leiden, the Netherlands) for pictures of the syntypes of Plumareola lineola van Eecke provided. Images of specimens deposited in NHMUK are used with permission and are copyright of the Trustees of the Natural History Museum, London and made available under Creative Commons License 4.0, CC-BY (https: // creativecommons. org / licenses / by / 4.0 /).	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
1F4087B7FFC5FFD717E2FC82FD23FBD9.taxon	description	Bucsek, K. (2020) Contribution to the knowledge of Lithosiini (Erebidae, Arctiinae) of central and northern Laos, part 4. Entomofauna carpathica, 32 (2), 47 – 87. Černý, K. & Pinratana, A. (2009) Moths of Thailand. Vol. 6. Arctiidae. Brothers of Saint Gabriel in Thailand, Bangkok, 283 pp. Dubatolov, V. V. & Bucsek, K. (2016) New lichen-moth taxa (Lepidoptera, Arctiidae, Lithosiinae) from Vietnam. Euroasian entomological journal, 15 (3), 228 – 238. Dubatolov, V. V., Kishida, Y. & Wang, M. (2012) New records of lichen-moth from the Nanling Mts., Guangdong, South China, with descriptions of new genera and species (Lepidoptera, Arctiidae: Lithosiinae). Tinea, 22 (1), 25 – 52. Fang, C. L. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). 19. Science Press, Beijing, 590 pp. [in Chinese with English summary] Hampson, G. F. (1894) The Fauna of British India, including Ceylon and Burma Fauna Br. India (Moths). Vol. 2. Taylor and Francis, London, pp. v-xxii, 1 – 609. Hampson, G. F. (1900) Catalogue of the Arctiadae (Nolinae, Lithosiinae) in the collection of the British Museum. In: Catalogue of the Lepidoptera Phalaenae in the collection of the British Museum, 2. British Museum (Natural History), London, xx + 589 pp., pl. XVIII – XXXV. Holloway, J. D. (2001) The Moths of Borneo, part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279 – 486. Huang, S. - Y., Volynkin, A. V. & Müller, G. (2024) Two new species of the genus Teulisna Walker (Lepidoptera, Erebidae, Arctiinae, Lithosiini) from Myanmar and Nepal. Zootaxa, 5506 (4), 577 – 586. https: // doi. org / 10.11646 / zootaxa. 5506.4.7 ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London, 306 pp. Kirti, J. S. & Singh, N. (2016) Arctiid Moths of India. Vol. 2. Nature Books India, New Delhi, 214 pp. Kononenko, V. S. (2010) Noctuidae Sibiricae. Vol. 2. Micronoctuidae, Noctuidae: Rivulinae – Agaristinae (Lepidoptera). Entomological Press, Sorø, 475 pp. Lafontaine, J. D. & Mikkola, K. (1987) Las-och-nyckel systemen i de inre genitalierna av Noctuidae (Lepidoptera) som taksonomiska kaennetecken. [Lock-and-key systems in the inner genitalia of Noctuidae (Lepidoptera) as a taxonomic character.] Entomologiske Meddelelser, 55, 161 – 167. [In Swedish with English summary] Leech, J. H. (1890) New species of Lepidoptera from China. The Entomologist, 23, 26 – 50, 81 – 83, 109 – 114, 187 – 192 + pl. 1. Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1878, 3 – 37, pls. 1 – 3. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07927. x Singh, N., Kirti, J. S., Datta, H. S., Joshi, R. & Volynkin, A. V. (2019) A review of the genus Dolgoma Moore from India, with notes on the genus taxonomy and descriptions of a new genus, four new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa, 4683 (1), 33 – 54. https: // doi. org / 10.11646 / zootaxa. 4683.1.2 Volynkin, A. V. (2023 a) On the taxonomy of the genus Teulisna Walker from India with a description of a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa, 5351 (3), 380 – 388. https: // doi. org / 10.11646 / zootaxa. 5351.3.6 Volynkin, A. V. (2023 b) Teulisna spinulosa, a new species from Northern Vietnam (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 68, 127 – 133. Volynkin, A. V. (2023 c) Taxonomic review of the Teulisna pallida (Moore) species-group with descriptions of two new species from the Himalayas (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 63, 15 – 23. https: // dx. doi. org / 10.37828 / em. 2023.63.2 Volynkin, A. V. (2024 a) Biampullisna, a new genus for “ Teulisna ” uniplaga Hampson (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Ecologica Montenegrina, 78, 72 – 78. https: // dx. doi. org / 10.37828 / em. 2024.78.8 Volynkin, A. V. (2024 b) On the terminology of the genitalia structures of lichen moths (Lepidoptera: Erebidae: Arctiinae: Lithosiini) with some references to Noctuidae. Ecologica Montenegrina, 73, 176 – 207. https: // dx. doi. org / 10.37828 / em. 2024.73.18 Walker, F. (1862) Catalogue of the Heterocerous lepidopterous insects collected at Sarawak, in Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the proceedings of the Linnean Society. Zoology, 6, 82 – 145.	en	Volynkin, Anton V., Černý, Karel, Huang, Si-Yao (2025): On the generic assignment of the taxa placed in the Oriental footman-moth genus Teulisna with the description of three new genera, one subgenus and eighteen new species. Ecologica Montenegrina 88: 104-163, DOI: 10.37828/em.2025.88.10, URL: https://doi.org/10.37828/em.2025.88.10
